treatments-xml/data/03/9E/2C/039E2C42BD34FF9663F0FE26FDFB4CDA.xml
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<document id="98BB82AB7B3681AD2423A2B82EC79A90" ID-DOI="10.11646/zootaxa.5448.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11231707" ID-ZooBank="79A27DBA-56C0-4D86-BA36-35D4070CED9A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1716279273055" checkinUser="plazi" docAuthor="Joshi, Shantanu, Sarkar, Vivek, Sawant, Dattaprasad, Pawar, Ujwala &amp; Kunte, Krushnamegh" docDate="2024" docId="039E2C42BD34FF9663F0FE26FDFB4CDA" docLanguage="en" docName="zootaxa.5448.3.2.pdf" docOrigin="Zootaxa 5448 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5448.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta khasia Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="361" masterDocId="FFA7543ABD3CFF9B6367FFB5FFF24E17" masterDocTitle="Description of Protosticta khasia sp. nov. and Yunnanosticta siangi sp. nov., with new records of P. samtsensis Gurung &amp; Phan, 2023 from Northeast India (Odonata: Zygoptera: Platystictidae)" masterLastPageNumber="370" masterPageNumber="348" pageNumber="356" updateTime="1716379561080" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="4072F96DA16E2E53AF918B7F4104CDB8">Description of Protosticta khasia sp. nov. and Yunnanosticta siangi sp. nov., with new records of P. samtsensis Gurung &amp; Phan, 2023 from Northeast India (Odonata: Zygoptera: Platystictidae)</mods:title>
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<mods:namePart id="9F95EA82C6BF420BF12457C9FE2E08DE">Joshi, Shantanu</mods:namePart>
<mods:nameIdentifier id="B23D1DDE71F34EF931907704F65BDBBA" type="ORCID">0000-0002-4552-5746</mods:nameIdentifier>
<mods:affiliation id="79824585215B285B40B4A347F832381D">Department of Biological Sciences, University of Arkansas, Fayetteville, 72701, Arkansas, USA &amp; National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India</mods:affiliation>
<mods:nameIdentifier id="2721890483CD82F8E3024D5AE3E42044" type="email">sj058@uark.edu</mods:nameIdentifier>
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<mods:namePart id="5A633A91D850CD9BB237F91631524D66">Sarkar, Vivek</mods:namePart>
<mods:nameIdentifier id="5BA19BFB3ADB71C355105CFA11AA3030" type="ORCID">0000-0002-0091-0288</mods:nameIdentifier>
<mods:affiliation id="E317FBB958B9A5EDCAF3C550214AF09E">Wildlife Institute of India, Chandrabani, Dehradun, 248001, Uttarakhand, India</mods:affiliation>
<mods:nameIdentifier id="BA0C17F8F04E7DE8198B1C24D80E981A" type="email">viveksarkar87@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D7718B51B88EAB9C24708A0649C2750C">Sawant, Dattaprasad</mods:namePart>
<mods:nameIdentifier id="76A8BF52FD1039E28EFB2A970E17EDD9" type="ORCID">0000-0003-0533-9292</mods:nameIdentifier>
<mods:affiliation id="15CE44A7DC6073C01718A8A47A93E5D2">Government Medical College and Hospital, Sindhudurg, 416534, Maharashtra, India</mods:affiliation>
<mods:nameIdentifier id="73C4192B40F2BE9B7768237D393CC354" type="email">dattaprasad.101@gmail.com</mods:nameIdentifier>
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<mods:namePart id="62FC86A3983D7A8A2558C719AFE250E6">Pawar, Ujwala</mods:namePart>
<mods:nameIdentifier id="7242BB5894D976CE1FA82B5AED8B3E4D" type="ORCID">0000-0003-1281-2228</mods:nameIdentifier>
<mods:affiliation id="A990614D0F437651AED2C2F80054D240">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India</mods:affiliation>
<mods:nameIdentifier id="27ED0FDD2E2A88CEB0AA8C0062FE45CF" type="email">ujwalamp@ncbs.res.in</mods:nameIdentifier>
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<mods:namePart id="74983063C2FF84E05A8DFE04519282DB">Kunte, Krushnamegh</mods:namePart>
<mods:nameIdentifier id="311666994F08D265D3B3C6FA67789C13" type="ORCID">0000-0002-3860-6118</mods:nameIdentifier>
<mods:affiliation id="373CA704188BC862CD4DC696E2C3CEC8">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India</mods:affiliation>
<mods:nameIdentifier id="2B6810FF99632B93D7F370967E84D151" type="email">krushnamegh@ncbs.res.in</mods:nameIdentifier>
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<mods:date id="7D221ED5376F95D06CFC6E4D92AE79B7">2024</mods:date>
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<taxonomicName id="4C37E6D7BD34FF9363F0FE26FE834FBA" authority="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte, 2024" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[151,369,402,429]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="356" phylum="Arthropoda" rank="species" species="khasia" status="sp. nov.">
<emphasis id="B9434146BD34FF9363F0FE26FE834FBA" bold="true" box="[151,369,402,429]" italics="true" pageId="8" pageNumber="356">Protosticta khasia</emphasis>
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<taxonomicNameLabel id="A270FC3DBD34FF93621FFE26FE244FBA" box="[376,470,403,429]" pageId="8" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
Joshi &amp; Sarkar
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<paragraph id="8B889D54BD34FF9363F0FE02FEE04FC6" blockId="8.[151,668,402,465]" box="[151,274,439,465]" pageId="8" pageNumber="356">
(
<figureCitation id="130C81D1BD34FF9363F8FE02FEF84FC6" box="[159,266,439,465]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="9.[152,255,1943,1967]" captionStartId-1="10.[152,255,1943,1967]" captionStartId-2="11.[152,255,1904,1928]" captionStartId-3="12.[152,255,1645,1669]" captionTargetBox-0="[201,1386,181,1919]" captionTargetBox-1="[173,1414,181,1919]" captionTargetBox-2="[151,1435,1333,1881]" captionTargetBox-3="[154,1434,181,1621]" captionTargetId-0="figure-20@9.[201,1386,181,1919]" captionTargetId-1="figure-19@10.[173,1414,181,1919]" captionTargetId-2="figure-462@11.[151,1435,1333,1881]" captionTargetId-3="figure-19@12.[154,1434,181,1621]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 6. Protosticta khasia sp. nov. Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view." captionText-1="FIGURE 7. Protosticta khasia sp. nov. Paratype female (IBC-BX522): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) ventral view; (e) lateral view; (f) dorsal view." captionText-2="FIGURE 8. Caudal appendages of Protosticta spp. All scale bars 0.5 mm. Protosticta damacornu: (a) ventro-lateral view. Protosticta khasia sp. nov.; (b) ventral view; (c) dorsal view; (d) lateral view. Protosticta fraseri: (e) dorsal view; (f) lateral view. Protosticta himalaica (redrawn from Fraser 1933): (g) dorsal view; (h) lateral view. Protosticta samstensis: (i) dorsal view; (j) lateral view." captionText-3="FIGURE 9. Protosticta khasia sp. nov. (a) paratype female (IBC-BX522) in situ; (b) habitat at type locality." figureDoi-0="http://doi.org/10.5281/zenodo.11231724" figureDoi-1="http://doi.org/10.5281/zenodo.11231729" figureDoi-2="http://doi.org/10.5281/zenodo.11231733" figureDoi-3="http://doi.org/10.5281/zenodo.11231735" httpUri-0="https://zenodo.org/record/11231724/files/figure.png" httpUri-1="https://zenodo.org/record/11231729/files/figure.png" httpUri-2="https://zenodo.org/record/11231733/files/figure.png" httpUri-3="https://zenodo.org/record/11231735/files/figure.png" pageId="8" pageNumber="356">Figs. 69</figureCitation>
)
</paragraph>
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<emphasis id="B9434146BD34FF9363F0FE4BFE774C0F" bold="true" box="[151,389,510,536]" pageId="8" pageNumber="356">Material examined</emphasis>
</paragraph>
<paragraph id="8B889D54BD34FF9363F0FDF2FD0C4C92" blockId="8.[151,1437,582,789]" pageId="8" pageNumber="356">
<materialsCitation id="3B5F9709BD34FF9363F0FDF2FD0C4C92" collectingDate="2017-05-26" collectorName="Vivek Sarkar" country="India" county="East Khasi Hills district" elevation="1393" latitude="25.2784" location="Maraikaphon" longLatPrecision="7" longitude="91.7226" municipality="Cherrapunjee" pageId="8" pageNumber="356" specimenCount="1" specimenCount-male="1" stateProvince="Meghalaya" typeStatus="holotype">
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<typeStatus id="548C23F6BD34FF9363F0FDF2FEF44C76" box="[151,262,583,609]" pageId="8" pageNumber="356" type="holotype">Holotype</typeStatus>
.
<specimenCount id="9D3156DDBD34FF936275FDF3FECF4C73" box="[274,317,582,612]" count="1" pageId="8" pageNumber="356" type="male">1 ♂</specimenCount>
</emphasis>
(IBC-BX521),
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,
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,
<collectingCounty id="62E9E5D8BD34FF936050FDF2FBB44C76" box="[823,1094,583,609]" pageId="8" pageNumber="356">East Khasi Hills district</collectingCounty>
,
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,
<collectingCountry id="F320DDC4BD34FF9367BCFDF2FAE74C76" box="[1243,1301,583,609]" name="India" pageId="8" pageNumber="356">India</collectingCountry>
(
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<geoCoordinate id="EE03FB93BD34FF9363F0FDDEFEFE4C92" box="[151,268,619,645]" degrees="91.7226" direction="east" orientation="longitude" pageId="8" pageNumber="356" precision="5" value="91.7226">91.7226 E</geoCoordinate>
,
<quantity id="4CCF30B1BD34FF93627FFDDEFE9F4C92" box="[280,365,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.393" pageId="8" pageNumber="356" unit="m" value="1393.0">
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ASL),
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<collectingDate id="EFCD427CBD34FF9362D8FDDEFDDD4C92" box="[447,559,619,645]" pageId="8" pageNumber="356" value="2017-05-26">26.v.2017</collectingDate>
</date>
,
<collectorName id="26C2F882BD34FF93615DFDDEFD3F4C92" box="[570,717,619,645]" pageId="8" pageNumber="356">Vivek Sarkar</collectorName>
leg.
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</paragraph>
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<materialsCitation id="3B5F9709BD34FF9363A0FD3AFBBA4CBE" box="[199,1096,654,684]" pageId="8" pageNumber="356" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B9434146BD34FF9363A0FD3AFE9E4CBB" bold="true" box="[199,364,654,684]" pageId="8" pageNumber="356">
<typeStatus id="548C23F6BD34FF9363A0FD3AFEC44CBE" box="[199,310,655,681]" pageId="8" pageNumber="356" type="paratype">Paratype</typeStatus>
.
<specimenCount id="9D3156DDBD34FF936225FD3BFE9E4CBB" box="[322,364,654,684]" count="1" pageId="8" pageNumber="356" type="female">1 ♀</specimenCount>
</emphasis>
(IBC-BX522), same location, date, and collector as the holotype
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.
</paragraph>
</subSubSection>
<subSubSection id="C32DCEDFBD34FF9363A0FD06FBF74D02" pageId="8" pageNumber="356" type="etymology">
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<emphasis id="B9434146BD34FF9363A0FD06FEBC4CDA" bold="true" box="[199,334,691,717]" pageId="8" pageNumber="356">Etymology.</emphasis>
The species epithet khasia is derived from the noun “Khasi” which refers to members of a Scheduled Tribe from northeastern
<collectingCountry id="F320DDC4BD34FF936142FD62FDAD4CE6" box="[549,607,727,753]" name="India" pageId="8" pageNumber="356">India</collectingCountry>
and parts of
<collectingCountry id="F320DDC4BD34FF936195FD62FC8A4CE6" box="[754,888,727,753]" name="Bangladesh" pageId="8" pageNumber="356">Bangladesh</collectingCountry>
. We dedicate this species to the Khasi people for their active role in restoring and conserving habitats (e.g.,
<bibRefCitation id="EFA6E0A5BD34FF936043FD4EFC0B4D02" author="Poffenberger, M." box="[804,1017,762,789]" pageId="8" pageNumber="356" pagination="4474 - 4494" refId="ref9517" refString="Poffenberger, M. (2015) Restoring and Conserving Khasi Forests: A Community-Based REDD Strategy from Northeast India. Forests, 6, 4474 - 4494. https: // doi. org / 10.3390 / f 6124382" type="journal article" year="2015">Poffenberger 2015</bibRefCitation>
).
</paragraph>
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<emphasis id="B9434146BD34FF9363F0FCD3FE494D96" bold="true" box="[151,443,870,897]" pageId="8" pageNumber="356">
Description of
<typeStatus id="548C23F6BD34FF936228FCD2FE494D96" box="[335,443,871,897]" pageId="8" pageNumber="356" type="holotype">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B889D54BD34FF9363F0FC3EFF1E4DB2" blockId="8.[151,443,870,933]" box="[151,236,907,933]" pageId="8" pageNumber="356">
(
<figureCitation id="130C81D1BD34FF9363F8FC3EFF114DB2" box="[159,227,907,933]" captionStart="FIGURE 6" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-20@9.[201,1386,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Protosticta khasia sp. nov. Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231724" httpUri="https://zenodo.org/record/11231724/files/figure.png" pageId="8" pageNumber="356">Fig. 6</figureCitation>
)
</paragraph>
<paragraph id="8B889D54BD34FF9363F0FC66FCA84AB6" blockId="8.[151,1437,979,1941]" pageId="8" pageNumber="356">
<emphasis id="B9434146BD34FF9363F0FC66FF264DFA" box="[151,212,979,1005]" italics="true" pageId="8" pageNumber="356">Head</emphasis>
(
<figureCitation id="130C81D1BD34FF936383FC66FEA94DFA" box="[228,347,979,1005]" captionStart="FIGURE 6" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-20@9.[201,1386,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Protosticta khasia sp. nov. Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231724" httpUri="https://zenodo.org/record/11231724/files/figure.png" pageId="8" pageNumber="356">Figs. 6ab</figureCitation>
). Eyes dark olive green. Median lobe of labium pale brown, lateral lobes dark brown, pointed at the apices anteriorly, ventrally labium covered by brown setae, posterior-ventral surface with a pale blue spot; lower one-fourth of labrum black, rest turquoise blue, anteclypeus turquoise blue; mandibles black; postclypeus, antefrons and postfrons black; antefrons and postfrons with a bronze metallic sheen. Antennal base dark brown, apical half dirty white; pedicel lower half dirty yellow, upper half dark brown; flagellum broken. Vertex and occipital area black with metallic green sheen; median ocellus creamy-white.
</paragraph>
<paragraph id="8B889D54BD34FF9363A0FB1EFC744846" blockId="8.[151,1437,979,1941]" pageId="8" pageNumber="356">
<emphasis id="B9434146BD34FF9363A0FB1EFEEA4AD2" box="[199,280,1195,1221]" italics="true" pageId="8" pageNumber="356">Thorax</emphasis>
(
<figureCitation id="130C81D1BD34FF936241FB1EFE6B4AD2" box="[294,409,1195,1221]" captionStart="FIGURE 6" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-20@9.[201,1386,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Protosticta khasia sp. nov. Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231724" httpUri="https://zenodo.org/record/11231724/files/figure.png" pageId="8" pageNumber="356">Figs. 6a, c</figureCitation>
). Prothorax: propleuron black, anterior lobe yellow with a black band medially; median lobe black medially, lateral lobes dark brown; anterior half of posterior lobe black, posterior half dark brown medially, lateral lobes dark brown, extending laterally, slightly triangular at apex. Pterothorax black with metallic green sheen on the dorsum, ventral half extensively marked with yellow as follows; anterior carina and mesothoracic stigma of mesostigmal plate dark brown. Mesepimeron marked with pale blue stripe at mesopleural ridge, extending from posterior to anterior end reaching till two-thirds of its length tapering and pointed anteriorly; metepisternum marked yellow broadly from anterior end reaching about 3/4
<superScript id="7C42301CBD34FF936196FA37FD0C4B87" attach="left" box="[753,766,1410,1424]" fontSize="6" pageId="8" pageNumber="356">th</superScript>
of its length; metathoracic spiracle yellow, metepisternum brown below the metathoracic spiracle extending posteriorly; lower 1/3
<superScript id="7C42301CBD34FF9360DFFA13FC344BA3" attach="left" box="[952,966,1446,1460]" fontSize="6" pageId="8" pageNumber="356">rd</superScript>
of mesinfraepisternum yellow, rest black; mesocoxa, metacoxa, metinfraepisternum, metepimeron, yellow; venter of metathorax yellow tipped with black at the posterior end. Legs: yellow, minute black spot posteriorly at the base of prothoracic coxae, posterior surface of femur dark brown, tibia dark brown with a thin yellow stripe anteriorly, tarsus and claws dark brown. Legs covered with minute brown setae throughout and large black setae laterally.
</paragraph>
<paragraph id="8B889D54BD34FF9363A0F9EEFD9B488E" blockId="8.[151,1437,979,1941]" pageId="8" pageNumber="356">
<emphasis id="B9434146BD34FF9363A0F9EEFEE14863" box="[199,275,1627,1652]" italics="true" pageId="8" pageNumber="356">Wings.</emphasis>
Hyaline; Pt dark brown and rhomboid, as long as 1 cell below, slightly larger in FW. Ax
<quantity id="4CCF30B1BD34FF93679BF9EEFAD54862" box="[1276,1319,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="356" unit="in" value="2.0">2 in</quantity>
all wings; Px
<quantity id="4CCF30B1BD34FF9363DBF9CAFF07488E" box="[188,245,1663,1689]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="8" pageNumber="356" unit="in" value="16.0">16 in</quantity>
right &amp;
<quantity id="4CCF30B1BD34FF936233F9CAFE7F488E" box="[340,397,1663,1689]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="8" pageNumber="356" unit="in" value="17.0">17 in</quantity>
left FW,
<quantity id="4CCF30B1BD34FF936291F9CAFDDD488E" box="[502,559,1663,1689]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="8" pageNumber="356" unit="in" value="16.0">16 in</quantity>
HW.
</paragraph>
<paragraph id="8B889D54BD34FF9363A0F916FD6D493E" blockId="8.[151,1437,979,1941]" pageId="8" pageNumber="356">
<emphasis id="B9434146BD34FF9363A0F916FEC348AA" box="[199,305,1699,1725]" italics="true" pageId="8" pageNumber="356">Abdomen</emphasis>
(
<figureCitation id="130C81D1BD34FF936227F916FE6348AA" box="[320,401,1699,1725]" captionStart="FIGURE 6" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-20@9.[201,1386,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Protosticta khasia sp. nov. Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231724" httpUri="https://zenodo.org/record/11231724/files/figure.png" pageId="8" pageNumber="356">Fig. 6a</figureCitation>
). Black marked with yellow. Ventral half of S12 yellow except dark brown tergites, anterior border of S2 with a thin yellow annule dorsally, S37 with yellow annules at the anterior border, broader sequentially. S8 and S9 with basal bright markings laterally, tapering downwards and larger on S8, M shaped turquoise blue marking dorsally at the posterior border of S9.
</paragraph>
<paragraph id="8B889D54BD34FF9363A0F886FD344982" blockId="8.[151,1437,979,1941]" pageId="8" pageNumber="356">
<emphasis id="B9434146BD34FF9363A0F886FE90495A" box="[199,354,1843,1869]" italics="true" pageId="8" pageNumber="356">Genital ligula</emphasis>
(
<figureCitation id="130C81D1BD34FF936217F886FE19495A" box="[368,491,1843,1869]" captionStart="FIGURE 6" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-20@9.[201,1386,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Protosticta khasia sp. nov. Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231724" httpUri="https://zenodo.org/record/11231724/files/figure.png" pageId="8" pageNumber="356">
Figs.
<quantity id="4CCF30B1BD34FF9362C9F886FE38495A" box="[430,458,1843,1869]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="8" pageNumber="356" unit="g" value="6.0">6g</quantity>
h
</figureCitation>
). Ligula curved midway in lateral view, junction of apical segments with two rounded flaps laterally with a tuft of hair at apices; apical segment furnished with a hood bifurcating into two broad arms. The two arms thinner at apices, apex club shaped.
</paragraph>
<caption id="DF48CDDCBD35FF9263FFF822FDDC49E0" ID-DOI="http://doi.org/10.5281/zenodo.11231724" ID-Zenodo-Dep="11231724" httpUri="https://zenodo.org/record/11231724/files/figure.png" pageId="9" pageNumber="357" startId="9.[152,255,1943,1967]" targetBox="[201,1386,181,1919]" targetPageId="9" targetType="figure">
<paragraph id="8B889D54BD35FF9263FFF822FDDC49E0" blockId="9.[151,1436,1943,2039]" pageId="9" pageNumber="357">
<emphasis id="B9434146BD35FF9263FFF822FEEB49B8" bold="true" box="[152,281,1943,1967]" pageId="9" pageNumber="357">FIGURE 6.</emphasis>
<taxonomicName id="4C37E6D7BD35FF926247F82DFE2A49B8" authority="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte, 2024" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[288,472,1943,1967]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="357" phylum="Arthropoda" rank="species" species="khasia" status="sp. nov.">
<emphasis id="B9434146BD35FF926247F82DFE2A49B8" box="[288,472,1943,1967]" italics="true" pageId="9" pageNumber="357">Protosticta khasia</emphasis>
</taxonomicName>
<emphasis id="B9434146BD35FF9262B8F822FDC249B8" bold="true" box="[479,560,1943,1967]" pageId="9" pageNumber="357">
<taxonomicNameLabel id="A270FC3DBD35FF9262B8F822FDC249B8" box="[479,560,1943,1967]" pageId="9" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view.
</paragraph>
</caption>
<caption id="DF48CDDCBD36FF9163FFF822FAE149C4" ID-DOI="http://doi.org/10.5281/zenodo.11231729" ID-Zenodo-Dep="11231729" httpUri="https://zenodo.org/record/11231729/files/figure.png" pageId="10" pageNumber="358" startId="10.[152,255,1943,1967]" targetBox="[173,1414,181,1919]" targetPageId="10" targetType="figure">
<paragraph id="8B889D54BD36FF9163FFF822FAE149C4" blockId="10.[151,1436,1943,2003]" pageId="10" pageNumber="358">
<emphasis id="B9434146BD36FF9163FFF822FEEA49B8" bold="true" box="[152,280,1943,1967]" pageId="10" pageNumber="358">FIGURE 7.</emphasis>
<taxonomicName id="4C37E6D7BD36FF916279F82DFE2449B8" authority="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte, 2024" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[286,470,1943,1967]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="358" phylum="Arthropoda" rank="species" species="khasia" status="sp. nov.">
<emphasis id="B9434146BD36FF916279F82DFE2449B8" box="[286,470,1943,1967]" italics="true" pageId="10" pageNumber="358">Protosticta khasia</emphasis>
</taxonomicName>
<emphasis id="B9434146BD36FF9162BBF822FDDE49B8" bold="true" box="[476,556,1943,1967]" pageId="10" pageNumber="358">
<taxonomicNameLabel id="A270FC3DBD36FF9162BBF822FDDE49B8" box="[476,556,1943,1967]" pageId="10" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC-BX522): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) ventral view; (e) lateral view; (f) dorsal view.
</paragraph>
</caption>
<paragraph id="8B889D54BD37FF9063A0FF22FDC74F56" blockId="11.[151,1437,151,357]" pageId="11" pageNumber="359">
<emphasis id="B9434146BD37FF9063A0FF22FE554EA6" box="[199,423,151,177]" italics="true" pageId="11" pageNumber="359">Caudal appendages</emphasis>
(
<figureCitation id="130C81D1BD37FF9062D1FF22FDDF4EA6" box="[438,557,151,177]" captionStart="FIGURE 6" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[201,1386,181,1919]" captionTargetId="figure-20@9.[201,1386,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Protosticta khasia sp. nov. Holotype male (IBC-BX521): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) dorsal view; (e) lateral view; (f) ventral view; (gh) genital ligula: (g) ventral view; (h) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231724" httpUri="https://zenodo.org/record/11231724/files/figure.png" pageId="11" pageNumber="359">Figs. 6df</figureCitation>
). Cerci black at base, paler brown towards apices and internally. Cerci broader at base folded inwards, excavated midway, strongly bent downwards, ending in knob-shaped apices, outer margin with dense setae. Paraproct dark brown with small but conspicuous basal spine at the upper margin, apices bifurcated into two arms, the outer arm about twice as thick as the inner arm, curved in and upwards, blunt at apices. Inner arm thin and curved, with pointed apices.
</paragraph>
<paragraph id="8B889D54BD37FF9063A0FEFEFB544F72" blockId="11.[151,1437,151,357]" box="[199,1190,330,357]" pageId="11" pageNumber="359">Measurements. Abdomen + caudal appendages = 38.5, FW = 26.827.1, HW = 29.830.</paragraph>
<paragraph id="8B889D54BD37FF9063F0FE1CFE4D4FD4" blockId="11.[151,447,425,487]" box="[151,447,425,451]" pageId="11" pageNumber="359">
<heading id="D0C02A38BD37FF9063F0FE1CFE4D4FD4" bold="true" box="[151,447,425,451]" fontSize="11" level="1" pageId="11" pageNumber="359" reason="1">
<emphasis id="B9434146BD37FF9063F0FE1CFE4D4FD4" bold="true" box="[151,447,425,451]" pageId="11" pageNumber="359">
Description of
<typeStatus id="548C23F6BD37FF906228FE1CFE4D4FD4" box="[335,447,425,451]" pageId="11" pageNumber="359" type="paratype">paratype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B889D54BD37FF9063F0FE78FF194FF0" blockId="11.[151,447,425,487]" box="[151,235,461,487]" pageId="11" pageNumber="359">
(
<figureCitation id="130C81D1BD37FF9063F8FE78FF114FF0" box="[159,227,461,487]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-19@10.[173,1414,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Protosticta khasia sp. nov. Paratype female (IBC-BX522): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) ventral view; (e) lateral view; (f) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231729" httpUri="https://zenodo.org/record/11231729/files/figure.png" pageId="11" pageNumber="359">Fig. 7</figureCitation>
)
</paragraph>
<paragraph id="8B889D54BD37FF9063F0FDBAFBAB4C82" blockId="11.[151,1437,526,1309]" pageId="11" pageNumber="359">
<emphasis id="B9434146BD37FF9063F0FDBAFF264C3E" box="[151,212,527,553]" italics="true" pageId="11" pageNumber="359">Head</emphasis>
(
<figureCitation id="130C81D1BD37FF906383FDBAFEAD4C3E" box="[228,351,527,553]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-19@10.[173,1414,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Protosticta khasia sp. nov. Paratype female (IBC-BX522): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) ventral view; (e) lateral view; (f) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231729" httpUri="https://zenodo.org/record/11231729/files/figure.png" pageId="11" pageNumber="359">Figs. 7ab</figureCitation>
). Labium brown, lateral lobes pointed outward; upper 3/4
<superScript id="7C42301CBD37FF90608AFDBBFC084C0B" attach="left" box="[1005,1018,526,540]" fontSize="6" pageId="11" pageNumber="359">th</superScript>
of labrum turquoise blue, rest black; anteclypeus turquoise blue; anterior half of mandibles turquoise blue, rest of the face black with metallic sheen. Scapus black, apical 1/5
<superScript id="7C42301CBD37FF9062C5FDE3FE5D4C73" attach="left" box="[418,431,598,612]" fontSize="6" pageId="11" pageNumber="359">th</superScript>
yellow; lower half of pedicel yellow, rest black; flagellum broken; vertex black; eyes olive green, bluish green in situ; median ocelli brown, appear creamy white at some angles.
</paragraph>
<paragraph id="8B889D54BD37FF9063A0FD2AFC674DEA" blockId="11.[151,1437,526,1309]" pageId="11" pageNumber="359">
<emphasis id="B9434146BD37FF9063A0FD2AFEEA4CAE" box="[199,280,671,697]" italics="true" pageId="11" pageNumber="359">Thorax</emphasis>
(
<figureCitation id="130C81D1BD37FF906240FD2AFE6C4CAE" box="[295,414,671,697]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-19@10.[173,1414,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Protosticta khasia sp. nov. Paratype female (IBC-BX522): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) ventral view; (e) lateral view; (f) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231729" httpUri="https://zenodo.org/record/11231729/files/figure.png" pageId="11" pageNumber="359">Figs. 7a, c</figureCitation>
). Prothorax: propleuron dark brown; anterior lobe yellow marked with a black dorsal stripe medially similar to male, median lobe dark brown with variegated yellow markings on lateral lobes; posterior lobe brown, lateral margins yellowish-brown. Pterothorax: mesepisternum and mesepimeron black with metallic green sheen; dorsal carina with fine yellow marking along both sides on mesepisternum; mesepimeron with a faint blue marking at the humeral suture, broader posteriorly, tapering and reaching about 12 its length anteriorly; metepisternum broadly yellow anteriorly, brown posteriorly; metathoracic spiracle black; metepimeron and venter yellow with a blue tinge at the posterior end; lower 1/4
<superScript id="7C42301CBD37FF906001FCC3FC814D93" attach="left" box="[870,883,886,900]" fontSize="6" pageId="11" pageNumber="359">th</superScript>
of mesinfraepisternum yellow, rest dark brown; metinfraepisternum yellow except a brown band at the upper end covering about 1/5
<superScript id="7C42301CBD37FF90675DFC2FFBB54DBF" attach="left" box="[1082,1095,922,936]" fontSize="6" pageId="11" pageNumber="359">th</superScript>
its length; coxae yellow. Legs: femur yellow, posteriorly dark brown with small variegated black markings; tibia dark brown, yellow anteriorly; tarsus and claws dark brown; legs covered with black setae laterally.
</paragraph>
<paragraph id="8B889D54BD37FF9063A0FBB2FAF04A36" blockId="11.[151,1437,526,1309]" box="[199,1282,1031,1057]" pageId="11" pageNumber="359">
<emphasis id="B9434146BD37FF9063A0FBB2FEF94A37" box="[199,267,1031,1056]" italics="true" pageId="11" pageNumber="359">Wings</emphasis>
. Hyaline, veins dark brown. Pt blackish-brown, larger in FW. Ax 2; Px
<quantity id="4CCF30B1BD37FF906749FBB2FB954A36" box="[1070,1127,1031,1057]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="11" pageNumber="359" unit="in" value="17.0">17 in</quantity>
FW and HW.
</paragraph>
<paragraph id="8B889D54BD37FF9063A0FB9EFC3C4AA6" blockId="11.[151,1437,526,1309]" pageId="11" pageNumber="359">
<emphasis id="B9434146BD37FF9063A0FB9EFEC34A52" box="[199,305,1067,1093]" italics="true" pageId="11" pageNumber="359">Abdomen</emphasis>
(
<figureCitation id="130C81D1BD37FF906226FB9EFE664A52" box="[321,404,1067,1093]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-19@10.[173,1414,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Protosticta khasia sp. nov. Paratype female (IBC-BX522): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) ventral view; (e) lateral view; (f) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231729" httpUri="https://zenodo.org/record/11231729/files/figure.png" pageId="11" pageNumber="359">Fig. 7a</figureCitation>
). Dark brown, marked with yellow as follows: ventral half of S12 yellow, on S2 reaching about 1/5
<superScript id="7C42301CBD37FF906399FBFBFEF94A4B" attach="left" box="[254,267,1102,1116]" fontSize="6" pageId="11" pageNumber="359">th</superScript>
of the segment posteriorly; S2 with a small yellow marking dorsally at the anterior border; S37 with blueyellow crescent shaped rings at the anterior border, dorsal markings broader posteriorly, extended more ventrally; S8 marked yellow only ventrally at the anterior border; S910 unmarked.
</paragraph>
<paragraph id="8B889D54BD37FF9063A0FB0EFB524AEE" blockId="11.[151,1437,526,1309]" pageId="11" pageNumber="359">
<emphasis id="B9434146BD37FF9063A0FB0EFE574AC2" box="[199,421,1211,1237]" italics="true" pageId="11" pageNumber="359">Caudal appendages</emphasis>
(
<figureCitation id="130C81D1BD37FF9062D5FB0EFDEF4AC2" box="[434,541,1211,1237]" captionStart="FIGURE 7" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-19@10.[173,1414,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Protosticta khasia sp. nov. Paratype female (IBC-BX522): (a) lateral habitus; (b) anterior view of head; (c) dorsal view of prothorax; (df) caudal appendages, scale bars 0.5 mm; (d) ventral view; (e) lateral view; (f) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231729" httpUri="https://zenodo.org/record/11231729/files/figure.png" pageId="11" pageNumber="359">Fig. 7df</figureCitation>
). Cerci triangular, curved and sharply pointed at apices. Ovipositor black, broader at base, laterally dark brown. Style brown; terebra posteriorly pointed, short, grayish brown.
</paragraph>
<paragraph id="8B889D54BD37FF9063A0FAB6FB544B0A" blockId="11.[151,1437,526,1309]" box="[199,1190,1282,1309]" pageId="11" pageNumber="359">Measurements. Abdomen + caudal appendages = 40.2, FW = 26.927, HW = 26.326.5.</paragraph>
</subSubSection>
<caption id="DF48CDDCBD37FF9063FFF8C5FE8349E2" ID-DOI="http://doi.org/10.5281/zenodo.11231733" ID-Zenodo-Dep="11231733" httpUri="https://zenodo.org/record/11231733/files/figure.png" pageId="11" pageNumber="359" startId="11.[152,255,1904,1928]" targetBox="[151,1435,1333,1881]" targetPageId="11" targetType="figure">
<paragraph id="8B889D54BD37FF9063FFF8C5FE8349E2" blockId="11.[151,1437,1904,2037]" pageId="11" pageNumber="359">
<emphasis id="B9434146BD37FF9063FFF8C5FEE9499F" bold="true" box="[152,283,1904,1928]" pageId="11" pageNumber="359">FIGURE 8.</emphasis>
Caudal appendages of
<taxonomicName id="4C37E6D7BD37FF906175F8C4FD47499F" box="[530,693,1905,1928]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="359" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9434146BD37FF906175F8C4FD70499F" box="[530,642,1905,1928]" italics="true" pageId="11" pageNumber="359">Protosticta</emphasis>
spp.
</taxonomicName>
All scale bars 0.5 mm.
<taxonomicName id="4C37E6D7BD37FF9060D3F8C4FB6D499E" authorityName="Terzani &amp; Carletti" authorityYear="1998" box="[948,1183,1905,1929]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="359" phylum="Arthropoda" rank="species" species="damacornu">
<emphasis id="B9434146BD37FF9060D3F8C4FB6D499E" box="[948,1183,1905,1929]" italics="true" pageId="11" pageNumber="359">Protosticta damacornu</emphasis>
</taxonomicName>
: (a) ventro-lateral view.
<taxonomicName id="4C37E6D7BD37FF9063F0F820FEA349BA" authority="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte, 2024" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[151,337,1941,1965]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="359" phylum="Arthropoda" rank="species" species="khasia" status="sp. nov.">
<emphasis id="B9434146BD37FF9063F0F820FEA349BA" box="[151,337,1941,1965]" italics="true" pageId="11" pageNumber="359">Protosticta khasia</emphasis>
</taxonomicName>
<emphasis id="B9434146BD37FF90623EF820FE5E49BA" bold="true" box="[345,428,1941,1965]" pageId="11" pageNumber="359">
<taxonomicNameLabel id="A270FC3DBD37FF90623EF820FE5E49BA" box="[345,428,1941,1965]" pageId="11" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (b) ventral view; (c) dorsal view; (d) lateral view.
<taxonomicName id="4C37E6D7BD37FF9060A1F820FB7049BA" authorityName="Kennedy" authorityYear="1936" box="[966,1154,1941,1965]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="359" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD37FF9060A1F820FB7049BA" box="[966,1154,1941,1965]" italics="true" pageId="11" pageNumber="359">Protosticta fraseri</emphasis>
</taxonomicName>
: (e) dorsal view; (f) lateral view.
<taxonomicName id="4C37E6D7BD37FF9063B1F80CFE4149C6" authorityName="Laidlaw" authorityYear="1917" box="[214,435,1977,2001]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="359" phylum="Arthropoda" rank="species" species="himalaica">
<emphasis id="B9434146BD37FF9063B1F80CFE4149C6" box="[214,435,1977,2001]" italics="true" pageId="11" pageNumber="359">Protosticta himalaica</emphasis>
</taxonomicName>
(redrawn from
<bibRefCitation id="EFA6E0A5BD37FF90613CF80CFD2849C6" author="Fraser, F. C." box="[603,730,1977,2001]" pageId="11" pageNumber="359" refId="ref8709" refString="Fraser, F. C. (1933) The fauna of British India, including Ceylon and Burma. Odonata. Vol. I. Taylor &amp; Francis Ltd., London, 423 pp." type="book" year="1933">Fraser 1933</bibRefCitation>
): (g) dorsal view; (h) lateral view.
<taxonomicName id="4C37E6D7BD37FF90672EF80CFADF49C7" box="[1097,1325,1977,2000]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="359" phylum="Arthropoda" rank="species" species="samstensis">
<emphasis id="B9434146BD37FF90672EF80CFADF49C7" box="[1097,1325,1977,2000]" italics="true" pageId="11" pageNumber="359">Protosticta samstensis</emphasis>
</taxonomicName>
: (i) dorsal view; (j) lateral view.
</paragraph>
</caption>
<caption id="DF48CDDCBD30FF9763FFF9D8FB184892" ID-DOI="http://doi.org/10.5281/zenodo.11231735" ID-Zenodo-Dep="11231735" box="[152,1258,1645,1670]" httpUri="https://zenodo.org/record/11231735/files/figure.png" pageId="12" pageNumber="360" startId="12.[152,255,1645,1669]" targetBox="[154,1434,181,1621]" targetPageId="12" targetType="figure">
<paragraph id="8B889D54BD30FF9763FFF9D8FB184892" blockId="12.[152,1258,1645,1670]" box="[152,1258,1645,1670]" pageId="12" pageNumber="360">
<emphasis id="B9434146BD30FF9763FFF9D8FEEA4892" bold="true" box="[152,280,1645,1669]" pageId="12" pageNumber="360">FIGURE 9.</emphasis>
<taxonomicName id="4C37E6D7BD30FF976278F9DBFE254892" authority="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte, 2024" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[287,471,1645,1669]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="360" phylum="Arthropoda" rank="species" species="khasia" status="sp. nov.">
<emphasis id="B9434146BD30FF976278F9DBFE254892" box="[287,471,1645,1669]" italics="true" pageId="12" pageNumber="360">Protosticta khasia</emphasis>
</taxonomicName>
<emphasis id="B9434146BD30FF9762BAF9D8FDDC4892" bold="true" box="[477,558,1645,1669]" pageId="12" pageNumber="360">
<taxonomicNameLabel id="A270FC3DBD30FF9762BAF9D8FDDC4892" box="[477,558,1645,1669]" pageId="12" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) paratype female (IBC-BX522) in situ; (b) habitat at type locality.
</paragraph>
</caption>
<subSubSection id="C32DCEDFBD30FF9663F0F962FBA04FBA" lastPageId="13" lastPageNumber="361" pageId="12" pageNumber="360" type="diagnosis">
<paragraph id="8B889D54BD30FF9763F0F962FEE248E6" blockId="12.[151,272,1751,1777]" box="[151,272,1751,1777]" pageId="12" pageNumber="360">
<heading id="D0C02A38BD30FF9763F0F962FEE248E6" bold="true" box="[151,272,1751,1777]" fontSize="11" level="1" pageId="12" pageNumber="360" reason="1">
<emphasis id="B9434146BD30FF9763F0F962FEE248E6" bold="true" box="[151,272,1751,1777]" pageId="12" pageNumber="360">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B889D54BD30FF9763F0F8AAFE3449B1" blockId="12.[151,1437,1823,2030]" pageId="12" pageNumber="360">
Male of this species can be distinguished at once from all its congeners by the unique shape of paraproct (deeply incised, wrench-shaped apices, inner arm curved slightly upwards, sharply pointed). The abdominal markings (S9 with a M shaped blue marking on the dorsal margin) and markings of prothorax will help to distinguish both sexes from other
<taxonomicName id="4C37E6D7BD30FF976272F839FE3449B1" box="[277,454,1932,1958]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="360" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9434146BD30FF976272F839FE6249B2" box="[277,400,1932,1957]" italics="true" pageId="12" pageNumber="360">Protosticta</emphasis>
spp.
</taxonomicName>
</paragraph>
<paragraph id="8B889D54BD30FF9663A0F81AFEE24F56" blockId="12.[151,1437,1823,2030]" lastBlockId="13.[151,1437,151,429]" lastPageId="13" lastPageNumber="361" pageId="12" pageNumber="360">
The only species in Northeast
<collectingCountry id="F320DDC4BD30FF97617BF81AFDA449DE" box="[540,598,1967,1993]" name="India" pageId="12" pageNumber="360">India</collectingCountry>
that remotely resembles this new species in terms of caudal appendages is
<taxonomicName id="4C37E6D7BD30FF9763F0F861FEC649FA" authorityName="Terzani &amp; Carletti" authorityYear="1998" box="[151,308,2003,2029]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="360" phylum="Arthropoda" rank="species" species="damacornu">
<emphasis id="B9434146BD30FF9763F0F861FEC649FA" box="[151,308,2003,2029]" italics="true" pageId="12" pageNumber="360">P. damacornu</emphasis>
</taxonomicName>
. From
<taxonomicName id="4C37E6D7BD30FF9762E0F861FDD149FA" authorityName="Terzani &amp; Carletti" authorityYear="1998" box="[391,547,2003,2029]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="360" phylum="Arthropoda" rank="species" species="damacornu">
<emphasis id="B9434146BD30FF9762E0F861FDD149FA" box="[391,547,2003,2029]" italics="true" pageId="12" pageNumber="360">P. damacornu</emphasis>
</taxonomicName>
,
<taxonomicName id="4C37E6D7BD30FF976155F861FD6549FA" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[562,663,2003,2029]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="360" phylum="Arthropoda" rank="species" species="khasia">
<emphasis id="B9434146BD30FF976155F861FD6549FA" box="[562,663,2003,2029]" italics="true" pageId="12" pageNumber="360">P. khasia</emphasis>
</taxonomicName>
male can be distinguished by (a) shape of cerci (apices prominently bi-lobed in
<taxonomicName id="4C37E6D7BD31FF96627AFF22FE494EA6" authorityName="Terzani &amp; Carletti" authorityYear="1998" box="[285,443,151,177]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="361" phylum="Arthropoda" rank="species" species="damacornu">
<emphasis id="B9434146BD31FF96627AFF22FE494EA6" box="[285,443,151,177]" italics="true" pageId="13" pageNumber="361">P. damacornu</emphasis>
</taxonomicName>
(
<figureCitation id="130C81D1BD31FF9662AAFF22FDE34EA6" box="[461,529,151,177]" captionStart="FIGURE 8" captionStartId="11.[152,255,1904,1928]" captionTargetBox="[151,1435,1333,1881]" captionTargetId="figure-462@11.[151,1435,1333,1881]" captionTargetPageId="11" captionText="FIGURE 8. Caudal appendages of Protosticta spp. All scale bars 0.5 mm. Protosticta damacornu: (a) ventro-lateral view. Protosticta khasia sp. nov.; (b) ventral view; (c) dorsal view; (d) lateral view. Protosticta fraseri: (e) dorsal view; (f) lateral view. Protosticta himalaica (redrawn from Fraser 1933): (g) dorsal view; (h) lateral view. Protosticta samstensis: (i) dorsal view; (j) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231733" httpUri="https://zenodo.org/record/11231733/files/figure.png" pageId="13" pageNumber="361">Fig. 8</figureCitation>
)), (b) structure of paraproct (
<figureCitation id="130C81D1BD31FF966000FF22FC5E4EA6" box="[871,940,151,177]" captionStart="FIGURE 8" captionStartId="11.[152,255,1904,1928]" captionTargetBox="[151,1435,1333,1881]" captionTargetId="figure-462@11.[151,1435,1333,1881]" captionTargetPageId="11" captionText="FIGURE 8. Caudal appendages of Protosticta spp. All scale bars 0.5 mm. Protosticta damacornu: (a) ventro-lateral view. Protosticta khasia sp. nov.; (b) ventral view; (c) dorsal view; (d) lateral view. Protosticta fraseri: (e) dorsal view; (f) lateral view. Protosticta himalaica (redrawn from Fraser 1933): (g) dorsal view; (h) lateral view. Protosticta samstensis: (i) dorsal view; (j) lateral view." figureDoi="http://doi.org/10.5281/zenodo.11231733" httpUri="https://zenodo.org/record/11231733/files/figure.png" pageId="13" pageNumber="361">Fig. 8</figureCitation>
), (c) S9 with small dorsal blue marking or black (S9 completely blue in
<taxonomicName id="4C37E6D7BD31FF9662BBFF0EFD844EC2" authorityName="Terzani &amp; Carletti" authorityYear="1998" box="[476,630,187,213]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="361" phylum="Arthropoda" rank="species" species="damacornu">
<emphasis id="B9434146BD31FF9662BBFF0EFD844EC2" box="[476,630,187,213]" italics="true" pageId="13" pageNumber="361">P. damacornu</emphasis>
</taxonomicName>
), and (d) markings of prothorax. From
<taxonomicName id="4C37E6D7BD31FF96674DFF0EFB624EC2" authorityName="Kennedy" authorityYear="1936" box="[1066,1168,187,213]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="361" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD31FF96674DFF0EFB624EC2" box="[1066,1168,187,213]" italics="true" pageId="13" pageNumber="361">P. fraseri</emphasis>
</taxonomicName>
, the male differs widely by the thoracic and abdominal markings and the complex shape of paraproct (thin and pointed in
<taxonomicName id="4C37E6D7BD31FF966796FF6AFAAB4EEE" authorityName="Kennedy" authorityYear="1936" box="[1265,1369,223,249]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="361" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD31FF966796FF6AFAAB4EEE" box="[1265,1369,223,249]" italics="true" pageId="13" pageNumber="361">P. fraseri</emphasis>
</taxonomicName>
). The dark thorax, marking on S9 along with the shape of paraproct will serve to distinguish
<taxonomicName id="4C37E6D7BD31FF9667EBFEB6FB014F0A" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[1164,1267,259,285]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="361" phylum="Arthropoda" rank="species" species="khasia">
<emphasis id="B9434146BD31FF9667EBFEB6FB014F0A" box="[1164,1267,259,285]" italics="true" pageId="13" pageNumber="361">P. khasia</emphasis>
</taxonomicName>
from all other congeners.
</paragraph>
<paragraph id="8B889D54BD31FF9663A0FEFEFBA04FBA" blockId="13.[151,1437,151,429]" pageId="13" pageNumber="361">
Female
<taxonomicName id="4C37E6D7BD31FF966246FEFEFE744F72" authorityName="Joshi &amp; Sarkar &amp; Sawant &amp; Pawar &amp; Kunte" authorityYear="2024" box="[289,390,331,357]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="361" phylum="Arthropoda" rank="species" species="khasia">
<emphasis id="B9434146BD31FF966246FEFEFE744F72" box="[289,390,331,357]" italics="true" pageId="13" pageNumber="361">P. khasia</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName id="4C37E6D7BD31FF96619CFEFEFC5E4F72" box="[763,940,331,357]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="361" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9434146BD31FF96619CFEFEFC844F73" box="[763,886,331,356]" italics="true" pageId="13" pageNumber="361">Protosticta</emphasis>
spp.
</taxonomicName>
by the markings of prothorax (anterior lobe yellow with a brown median stripe, medial lobe brown, posterior margin of posterior lobe pale blue) and abdominal markings (S12 laterally yellow, base of S8 with a yellow dorsal markings) (
<figureCitation id="130C81D1BD31FF966094FE26FBB64FBA" box="[1011,1092,403,429]" captionStart="FIGURE 9" captionStartId="12.[152,255,1645,1669]" captionTargetBox="[154,1434,181,1621]" captionTargetId="figure-19@12.[154,1434,181,1621]" captionTargetPageId="12" captionText="FIGURE 9. Protosticta khasia sp. nov. (a) paratype female (IBC-BX522) in situ; (b) habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.11231735" httpUri="https://zenodo.org/record/11231735/files/figure.png" pageId="13" pageNumber="361">Fig. 9a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32DCEDFBD31FF9663F0FE4BFDFB4CDA" pageId="13" pageNumber="361" type="biology_ecology">
<paragraph id="8B889D54BD31FF9663F0FE4BFE564C0F" blockId="13.[151,420,510,537]" box="[151,420,510,537]" pageId="13" pageNumber="361">
<heading id="D0C02A38BD31FF9663F0FE4BFE564C0F" bold="true" box="[151,420,510,537]" fontSize="11" level="1" pageId="13" pageNumber="361" reason="1">
<emphasis id="B9434146BD31FF9663F0FE4BFE564C0F" bold="true" box="[151,420,510,537]" pageId="13" pageNumber="361">Habitat and behavior</emphasis>
</heading>
</paragraph>
<paragraph id="8B889D54BD31FF9663F0FDF2FDFB4CDA" blockId="13.[151,1437,582,717]" pageId="13" pageNumber="361">
<specimenCount id="9D3156DDBD31FF9663F0FDF2FEFA4C76" box="[151,264,583,609]" count="2" pageId="13" pageNumber="361" type="male">Two male</specimenCount>
individuals were observed&gt;
<quantity id="4CCF30B1BD31FF96613EFDF3FD744C76" box="[601,646,582,609]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="361" unit="m" value="5.0">5 m</quantity>
away from a small, shaded stream (
<figureCitation id="130C81D1BD31FF96674AFDF2FB714C76" box="[1069,1155,583,609]" captionStart="FIGURE 9" captionStartId="12.[152,255,1645,1669]" captionTargetBox="[154,1434,181,1621]" captionTargetId="figure-19@12.[154,1434,181,1621]" captionTargetPageId="12" captionText="FIGURE 9. Protosticta khasia sp. nov. (a) paratype female (IBC-BX522) in situ; (b) habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.11231735" httpUri="https://zenodo.org/record/11231735/files/figure.png" pageId="13" pageNumber="361">Fig. 9b</figureCitation>
). The female was found farther away in the forest. Individuals were observed perching a few feet high on vegetation and had the tendency of flying up into the canopy when disturbed. This species was only observed along one stream despite extensive surveys in the neighboring areas.
</paragraph>
</subSubSection>
</treatment>
</document>