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<document id="793E06471F4F4831CD44F672CE68687A" ID-DOI="10.3853/j.0067-1975.55.2003.1378" ID-ISSN="2201-4349" ID-Zenodo-Dep="10092588" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629577951457" checkinUser="felipe" docAuthor="Valentine, James L. &amp; Brock, Glenn A." docDate="2003" docId="039C661BAB78FF92FE907782A71A1C64" docLanguage="en" docName="RecAustMus.55.2.231-244.pdf" docOrigin="Records of the Australian Museum 55 (2)" docSource="https://journals.australian.museum/valentine-and-brock-2003-rec-aust-mus-552-231244/" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Orbaspina Valentine &amp; Brock, 2003, n.gen." docType="treatment" docVersion="3" lastPageNumber="237" masterDocId="FFA51E63AB7EFF94FFED7433A2741D6C" masterDocTitle="A New Siphonotretid Brachiopod from the Silurian of Central-Western New South Wales, Australia" masterLastPageNumber="244" masterPageNumber="231" pageNumber="237" updateTime="1699558620517" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="E2FC7086E4D3505172E2D53CD806F642">A New Siphonotretid Brachiopod from the Silurian of Central-Western New South Wales, Australia</mods:title>
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<mods:namePart id="0AF263798370F6BF019B76160632BC50">Valentine, James L.</mods:namePart>
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<mods:namePart id="17131AE44E354AEB6B8B3A6234EBFD6A">Brock, Glenn A.</mods:namePart>
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<mods:title id="D426B432E6D55040977EEE79045B54D8">Records of the Australian Museum</mods:title>
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<mods:date id="B62C28E2F9BB1D176A6132810FA1C32C">2003</mods:date>
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<mods:identifier id="2E04D8BE2A8E8B90F08DE937660DB0E1" type="ISSN">2201-4349</mods:identifier>
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<treatment id="039C661BAB78FF92FE907782A71A1C64" LSID="urn:lsid:plazi:treatment:039C661BAB78FF92FE907782A71A1C64" httpUri="http://treatment.plazi.org/id/039C661BAB78FF92FE907782A71A1C64" lastPageNumber="237" pageId="6" pageNumber="237">
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<paragraph id="8B8AD70DAB78FF92FE907782A0371EA5" blockId="6.[381,579,945,969]" box="[381,579,945,969]" pageId="6" pageNumber="237">
<heading id="D0C26061AB78FF92FE907782A0371EA5" box="[381,579,945,969]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="237" reason="2">
<emphasis id="B9410B1FAB78FF92FE907782A0371EA5" bold="true" box="[381,579,945,969]" pageId="6" pageNumber="237">
<taxonomicName id="4C35AC8EAB78FF92FE907782A3821EA5" authorityName="Valentine &amp; Brock" authorityYear="2003" box="[381,502,945,969]" class="Lingulata" family="Siphonotretidae" genus="Orbaspina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Linguliporida" pageId="6" pageNumber="237" phylum="Brachiopoda" rank="genus" status="gen. nov.">
<emphasis id="B9410B1FAB78FF92FE907782A3821EA5" bold="true" box="[381,502,945,969]" italics="true" pageId="6" pageNumber="237">Orbaspina</emphasis>
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<subSubSection id="C32F8486AB78FF92FF4B77D2A365195F" pageId="6" pageNumber="237" type="type_taxon">
<paragraph id="8B8AD70DAB78FF92FF4B77D2A365195F" blockId="6.[166,794,993,1075]" pageId="6" pageNumber="237">
<materialsCitation id="3B5DDD50AB78FF92FF4B77D2A379195F" country="Australia" location="Llandovery" municipality="Wenlock" pageId="6" pageNumber="237" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis id="B9410B1FAB78FF92FF4B77D2A3361E95" bold="true" box="[166,322,993,1017]" pageId="6" pageNumber="237">
<typeStatus id="548E69AFAB78FF92FF4B77D2A2951E95" box="[166,225,993,1017]" pageId="6" pageNumber="237">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C35AC8EAB78FF92FEB877D1A03D1E95" box="[341,585,994,1017]" class="Lingulata" family="Siphonotretidae" genus="Orbaspina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Linguliporida" pageId="6" pageNumber="237" phylum="Brachiopoda" rank="species" species="gelasinus" status="n.gen. and n.sp.">
<emphasis id="B9410B1FAB78FF92FEB877D1A03D1E95" box="[341,585,994,1017]" italics="true" pageId="6" pageNumber="237">Orbaspina gelasinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272B664AB78FF92FDB877D1A1601E95" box="[597,788,994,1017]" pageId="6" pageNumber="237" rank="species">n.gen. and n.sp.</taxonomicNameLabel>
,
<location id="8EEA81D6AB78FF92FF4B77CCA356197A" LSID="urn:lsid:plazi:treatment:039C661BAB78FF92FE907782A71A1C64:8EEA81D6AB78FF92FF4B77CCA356197A" box="[166,290,1023,1046]" country="Australia" municipality="Wenlock" name="Llandovery" pageId="6" pageNumber="237" stateProvince="New South Wales">Llandovery</location>
to
<collectingMunicipality id="6BEE4D77AB78FF92FEAC77CCA3D7197A" box="[321,419,1023,1046]" pageId="6" pageNumber="237">Wenlock</collectingMunicipality>
, central-western
<collectingRegion id="49F119EFAB78FF92FDBA77CCA162197A" box="[599,790,1023,1046]" country="Australia" name="New South Wales" pageId="6" pageNumber="237">New South Wales</collectingRegion>
,
<collectingCountry id="F322979DAB78FF92FF4B702FA379195F" box="[166,269,1052,1075]" name="Australia" pageId="6" pageNumber="237">Australia</collectingCountry>
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.
</paragraph>
</subSubSection>
<subSubSection id="C32F8486AB78FF92FF4B7063A0FB1859" pageId="6" pageNumber="237" type="diagnosis">
<paragraph id="8B8AD70DAB78FF92FF4B7063A0FB1859" blockId="6.[166,794,1104,1333]" pageId="6" pageNumber="237">
<emphasis id="B9410B1FAB78FF92FF4B7063A36C1904" bold="true" box="[166,280,1104,1128]" pageId="6" pageNumber="237">Diagnosis</emphasis>
. A siphonotretid with large, keyhole-shaped pedicle foramen extending forward through resorption to form an elongate, broadly triangular, pedicle track; pedicle track covered posteriorly by a concave plate and anteriorly by short “listrum-like” plate; tubular hollow spines of uniform size arranged in concentric rows close to valve margin; post larval shell bearing numerous subcircular dimples loosely arranged in concentric rows.
</paragraph>
</subSubSection>
<subSubSection id="C32F8486AB78FF92FF4B7162A3ED18B2" pageId="6" pageNumber="237" type="etymology">
<paragraph id="8B8AD70DAB78FF92FF4B7162A3ED18B2" blockId="6.[166,794,1361,1502]" pageId="6" pageNumber="237">
<emphasis id="B9410B1FAB78FF92FF4B7162A3561805" bold="true" box="[166,290,1361,1385]" pageId="6" pageNumber="237">Etymology</emphasis>
.
<taxonomicName id="4C35AC8EAB78FF92FEDF7161A31F1805" authorityName="Hall" authorityYear="1867" box="[306,363,1362,1385]" class="Agaricomycetes" family="Camaenidae" genus="Orba" kingdom="Fungi" order="Polyporales" pageId="6" pageNumber="237" phylum="Basidiomycota" rank="genus">
<emphasis id="B9410B1FAB78FF92FEDF7161A31F1805" box="[306,363,1362,1385]" italics="true" pageId="6" pageNumber="237">Orba</emphasis>
</taxonomicName>
L., orphan, refers to the occurrence of this taxon after the supposed demise of the siphonotretids during the end-Ordovician extinction event;
<emphasis id="B9410B1FAB78FF92FD5F71BEA09B18C8" box="[690,751,1421,1444]" italics="true" pageId="6" pageNumber="237">spina</emphasis>
L., thorny, in reference to the post-larval shell ornament of tubular hollow spines.
</paragraph>
</subSubSection>
<subSubSection id="C32F8486AB78FF92FF4B71C8A71A1C64" pageId="6" pageNumber="237" type="discussion">
<paragraph id="8B8AD70DAB78FF92FF4B71C8A16E1AC1" blockId="6.[166,794,1531,1994]" pageId="6" pageNumber="237">
<emphasis id="B9410B1FAB78FF92FF4B71C8A37A1B7F" bold="true" box="[166,270,1531,1555]" pageId="6" pageNumber="237">Remarks</emphasis>
. Features considered to be of generic significance among the siphonotretids include the nature of the pedicle foramen and tube, the pseudointerarea of both valves and the presence of tubular hollow spines (Holmer &amp; Popov, 2000). Based on these features,
<taxonomicName id="4C35AC8EAB78FF92FDEE7242A0021BE4" authorityName="Valentine &amp; Brock" authorityYear="2003" box="[515,630,1649,1672]" class="Lingulata" family="Siphonotretidae" genus="Orbaspina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Linguliporida" pageId="6" pageNumber="237" phylum="Brachiopoda" rank="genus" status="gen. nov.">
<emphasis id="B9410B1FAB78FF92FDEE7242A0021BE4" box="[515,630,1649,1672]" italics="true" pageId="6" pageNumber="237">Orbaspina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272B664AB78FF92FD937242A0B41BE4" box="[638,704,1649,1672]" pageId="6" pageNumber="237" rank="genus">n.gen.</taxonomicNameLabel>
appears closest morphologically to those genera grouped together by
<bibRefCitation id="EFA4AAFCAB78FF92FF217298A3F51BAE" author="Havlieek" box="[204,385,1707,1731]" firstAuthor="Havlieek" pageId="6" pageNumber="237" pagination="9 - 82" refId="ref8866" refString="Havlieek, V., 1982. Lingulacea, Paterinacea, and Siphonotretacea (Brachiopoda) in the Lower Ordovician sequence of Bohemia. Sbornik Geologickych ved, Paleontologie 25: 9 - 82." type="journal article" year="1982">Havlíěek (1982)</bibRefCitation>
and
<bibRefCitation id="EFA4AAFCAB78FF92FE517298A0BB1BAE" author="Popov &amp; Holmer" box="[444,719,1707,1730]" firstAuthor="Popov" pageId="6" pageNumber="237" pagination="1 - 156" refId="ref9827" refString="Popov, L. E., &amp; L. E. Holmer, 1994. Cambrian-Ordovician lingulate brachiopods from Scandinavia, Kazakhstan, and South Ural Mountains. Fossils and Strata 35: 1 - 156." type="journal article" year="1994">Popov &amp; Holmer (1994)</bibRefCitation>
in the subfamily
<taxonomicName id="4C35AC8EAB78FF92FEFA72FAA3B81B8C" authorityName="Havlíček" authorityYear="1982" box="[279,460,1737,1760]" class="Lingulata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonotretida" pageId="6" pageNumber="237" phylum="Brachiopoda" rank="subFamily" subFamily="Schizamboninae">Schizamboninae</taxonomicName>
. The most characteristic feature of these siphonotretids is the large pedicle foramen that extends forward through resorption, forming an elongate triangular pedicle track that may be covered posteriorly by a plate. However, the dimpled post-larval shell of
<taxonomicName id="4C35AC8EAB78FF92FD44730DA16E1A39" authorityName="Valentine &amp; Brock" authorityYear="2003" box="[681,794,1854,1877]" class="Lingulata" family="Siphonotretidae" genus="Orbaspina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Linguliporida" pageId="6" pageNumber="237" phylum="Brachiopoda" rank="genus" status="gen. nov.">
<emphasis id="B9410B1FAB78FF92FD44730DA16E1A39" box="[681,794,1854,1877]" italics="true" pageId="6" pageNumber="237">Orbaspina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272B664AB78FF92FF4B7368A29D1A1E" box="[166,233,1883,1906]" pageId="6" pageNumber="237" rank="genus">n.gen.</taxonomicNameLabel>
is unknown in any previously described Cambrian or Ordovician siphonotretid. This feature is present on a number of post-Ordovician siphonotretids including,
</paragraph>
<paragraph id="8B8AD70DAB78FF92FF5F7380A71A1C64" blockId="6.[166,794,1531,1994]" lastBlockId="6.[839,1467,153,264]" pageId="6" pageNumber="237">
<taxonomicName id="4C35AC8EAB78FF92FF5F7380A0361AA6" authority="Morris (1849)" authorityName="Morris" authorityYear="1849" box="[178,578,1971,1994]" class="Lingulata" family="Siphonotretidae" genus="Siphonotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Linguliporida" pageId="6" pageNumber="237" phylum="Brachiopoda" rank="species" species="anglica">
<emphasis id="B9410B1FAB78FF92FF5F7380A3D51AA6" box="[178,417,1971,1994]" italics="true" pageId="6" pageNumber="237">Siphonotreta anglica</emphasis>
<bibRefCitation id="EFA4AAFCAB78FF92FE457380A0361AA6" author="Morris" box="[424,578,1971,1994]" firstAuthor="Morris" pageId="6" pageNumber="237" pagination="315 - 321" refId="ref9722" refString="Morris, J., 1849. Note on the genus Siphonotreta, with a description of a new species. Annals and Magazine of Natural History 4: 315 - 321." type="journal article" year="1849">Morris (1849)</bibRefCitation>
</taxonomicName>
, and Siphonotretine sp. and Schizambonine sp. A
<bibRefCitation id="EFA4AAFCAB78FF92FB6274AAA75E1DDC" author="Mergl" box="[1167,1322,153,176]" firstAuthor="Mergl" pageId="6" pageNumber="237" pagination="345 - 351" refId="ref9545" refString="Mergl, M., 2001 a. Extinction of some lingulate brachiopod families: new stratigraphical data from the Silurian and Devonian of central Bohemia. In Brachiopods Past and Present, ed. C. H. C. Brunton, L. R. M. Cocks &amp; S. M. Long. Systematics Association Special Volume 63: 345 - 351." type="journal article" year="2001" yearSuffix="a">Mergl (2001a</bibRefCitation>
,b). The short “listrum-like” plate anteriorly covering the pedicle track of
<taxonomicName id="4C35AC8EAB78FF92FC8574E7A1811D87" box="[872,1013,212,235]" class="Lingulata" family="Siphonotretidae" genus="Orbaspina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Linguliporida" pageId="6" pageNumber="237" phylum="Brachiopoda" rank="species" species="gelasinus" status="n.gen. and n.sp.">
<emphasis id="B9410B1FAB78FF92FC8574E7A1811D87" box="[872,1013,212,235]" italics="true" pageId="6" pageNumber="237">O. gelasinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272B664AB78FF92FC1274E0A6C61D86" box="[1023,1202,211,234]" pageId="6" pageNumber="237" rank="species">n.gen. and n.sp.</taxonomicNameLabel>
(
<figureCitation id="130ECB88AB78FF92FB2874E0A76C1D86" box="[1221,1304,211,234]" captionStart="Fig" captionStartId="9.[121,157,1301,1322]" captionTargetBox="[143,1408,190,1256]" captionTargetId="figure-10@9.[129,1422,166,1269]" captionTargetPageId="9" captionText="Fig. 6. Orbaspina gelasinus n.gen. and n.sp.: ad, paratype AMF120612, ventral valve from sample BM 14.30, external view (a), detail of spines on anterior slope (b), detail of larval shell (c), internal view (d); e, f, paratypeAMF120613, ventral valve from sample BM 14.30, internal view (e), detail of pseudointerarea (f); g, paratype AMF122213, ventral valve from sample BM 13.80, internal view, note the “listrum-like” plate covering the anterior portion of the pedicle track as indicated by arrow; h, paratypeAMF122214, ventral valve from sample BM 9.90, external view showing common state of presentation of recovered ventral valves; i, paratype AMF122215, fragment from sample BM 9.30, external view. Scale bars = 500 µm (g); 100 µm (a, cf,h, i); 10 µm (b)." figureDoi="http://doi.org/10.5281/zenodo.10092602" httpUri="https://zenodo.org/record/10092602/files/figure.png" pageId="6" pageNumber="237">
Fig.
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) has not been previously documented among the siphonotretids.
</paragraph>
</subSubSection>
</treatment>
</document>