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<document id="CE27ABFF4C659499F5119359492005C4" ID-CLB-Dataset="8783" ID-DOI="10.11646/zootaxa.4908.3.1" ID-GBIF-Dataset="6206d142-3b21-4c39-afca-75b602420619" ID-ISSN="1175-5326" ID-Zenodo-Dep="4441762" ID-ZooBank="F2774810-3C77-426E-A1BC-0F42F21E7F9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1610699400433" checkinUser="plazi" docAuthor="Snegovaya, Nataly Yu. &amp; Cokendolpher, James C." docDate="2021" docId="039C113F0958FFA098EBDFDEFC21F8BC" docLanguage="en" docName="zootaxa.4908.3.1.pdf" docOrigin="Zootaxa 4908 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homolophus kozlovi Snegovaya &amp; Cokendolpher 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="333" masterDocId="FFA569470978FF80987CDD6CFFAEFFF1" masterDocTitle="Further studies on harvestman genus Homolophus (Opiliones: Phalangiidae), with descriptions of two new species" masterLastPageNumber="353" masterPageNumber="301" pageNumber="333" updateTime="1698924489785" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="73B3A13FDD43FB003D74E7B12F89423A">Further studies on harvestman genus Homolophus (Opiliones: Phalangiidae), with descriptions of two new species</mods:title>
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<mods:namePart id="8C59FFB22549A46B104175240DFB10C6">Snegovaya, Nataly Yu.</mods:namePart>
<mods:affiliation id="C4541755561F237FA27573D6CD61A6F0">Institute of Zoology, Azerbaijan Academy of Sciences, pr. 1128, kv. 504, Baku AZ 1073, Azerbaijan.</mods:affiliation>
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<mods:namePart id="A66F2419519C12A4582F1CD0AC3BF2B3">Cokendolpher, James C.</mods:namePart>
<mods:affiliation id="377E1E1C8F9785C4DFA37490D4077A73">Invertebrate Zoology, Museum of Texas Tech University, Lubbock, Texas 79415, USA.</mods:affiliation>
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<treatment id="039C113F0958FFA098EBDFDEFC21F8BC" ID-DOI="http://doi.org/10.5281/zenodo.4450804" ID-GBIF-Taxon="177009874" ID-Zenodo-Dep="4450804" LSID="urn:lsid:plazi:treatment:039C113F0958FFA098EBDFDEFC21F8BC" httpUri="http://treatment.plazi.org/id/039C113F0958FFA098EBDFDEFC21F8BC" lastPageNumber="333" pageId="32" pageNumber="333">
<subSubSection id="C32FF3A20958FFA098EBDFDEFE5BFD3C" box="[151,501,690,717]" pageId="32" pageNumber="333" type="nomenclature">
<paragraph id="8B8AA0290958FFA098EBDFDEFE5BFD3C" blockId="32.[151,501,690,753]" box="[151,501,690,717]" pageId="32" pageNumber="333">
<heading id="D0C217450958FFA098EBDFDEFE5BFD3C" bold="true" box="[151,501,690,717]" fontSize="11" level="1" pageId="32" pageNumber="333" reason="1">
<emphasis id="B9417C3B0958FFA098EBDFDEFE5BFD3C" bold="true" box="[151,501,690,717]" pageId="32" pageNumber="333">
<taxonomicName id="4C35DBAA0958FFA098EBDFDEFE3EFD3C" authority="Snegovaya &amp; Cokendolpher, 2021" authorityName="Snegovaya &amp; Cokendolpher" authorityYear="2021" box="[151,400,690,717]" class="Arachnida" family="Phalangiidae" genus="Homolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="kozlovi" status="sp. nov.">
<emphasis id="B9417C3B0958FFA098EBDFDEFE3EFD3C" bold="true" box="[151,400,690,717]" italics="true" pageId="32" pageNumber="333">Homolophus kozlovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272C1400958FFA099EBDFDFFE5BFD3C" box="[407,501,691,717]" pageId="32" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32FF3A20958FFA098EBDFBBFECCFD00" box="[151,354,727,753]" pageId="32" pageNumber="333" type="description">
<paragraph id="8B8AA0290958FFA098EBDFBBFECCFD00" blockId="32.[151,501,690,753]" box="[151,354,727,753]" pageId="32" pageNumber="333">
<figureCitation id="130EBCAC0958FFA098EBDFBBFEBEFD00" box="[151,272,727,753]" captionStart="FIGURE 14" captionStartId="22.[151,250,1974,1998]" captionTargetBox="[179,1408,1081,1941]" captionTargetId="figure-56@22.[173,1415,1074,1950]" captionTargetPageId="22" captionText="FIGURE 14. Distribution of some Homolophus species in the Palearctic: Central and Inner Asia. Data for pale colored symbols taken from literature; identifications not verified." figureDoi="http://doi.org/10.5281/zenodo.4441798" httpUri="https://zenodo.org/record/4441798/files/figure.png" pageId="32" pageNumber="333">Figures 14</figureCitation>
,
<figureCitation id="130EBCAC0958FFA09960DFBBFECCFD00" box="[284,354,727,753]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStartId-0="33.[151,250,1923,1947]" captionStartId-1="34.[151,250,988,1012]" captionTargetBox-0="[159,1431,181,1898]" captionTargetBox-1="[170,1436,181,965]" captionTargetId-0="figure-17@33.[151,1436,181,1899]" captionTargetId-1="figure-22@34.[151,1436,181,965]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="FIGURE 23. Homolophus kozlovi sp. nov., male: A, body, dorsal; B, ocularium, dorsal; C, femur I, dorsal; D, right pedipalp, ectal; E, right pedipalp, mesal; F, right chelicera, ectal; G, right chelicera, mesal; H, penis, dorsal; I, penis, lateral; J, penis glans, dorsal; K, penis glans, lateral." captionText-1="FIGURE 24. Famous scientists: (left) Pyotr Kuzmich Kozlov (18631935), a Russian traveler, geographer, ethnographer, archeologist, and explorer of Mongolia and Xîzàng (Tibet), for which Homolophus kozlovi sp. nov. is named; (right) Dmitry A. Milko, an entomologist, researcher, and zoologist from Kyrgyzstan for which Homolophus milkoi sp. nov. is named." figureDoi-0="http://doi.org/10.5281/zenodo.4441823" figureDoi-1="http://doi.org/10.5281/zenodo.4441825" httpUri-0="https://zenodo.org/record/4441823/files/figure.png" httpUri-1="https://zenodo.org/record/4441825/files/figure.png" pageId="32" pageNumber="333">2324</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32FF3A20958FFA098EBDE72FB9AFC73" pageId="32" pageNumber="333" type="diagnosis">
<paragraph id="8B8AA0290958FFA098EBDE72FB9AFC73" blockId="32.[151,1437,798,1869]" pageId="32" pageNumber="333">
<emphasis id="B9417C3B0958FFA098EBDE72FE4AFCC9" bold="true" box="[151,484,798,825]" pageId="32" pageNumber="333">Diagnosis and Comparisons</emphasis>
. With diamond shaped glans (in anterior view) of penis. New species most similar to
<taxonomicName id="4C35DBAA0958FFA098EBDE2FFD97FCAC" authority="(Roewer, 1911)" authorityName="Snegovaya &amp; Cokendolpher" authorityYear="2021" baseAuthorityName="Roewer" baseAuthorityYear="1911" box="[151,569,835,861]" class="Arachnida" family="Phalangiidae" genus="Homolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="almasyi">
<emphasis id="B9417C3B0958FFA098EBDE2FFE2AFCAC" box="[151,388,835,861]" italics="true" pageId="32" pageNumber="333">Homolophus almasyi</emphasis>
(
<bibRefCitation id="EFA4DDD80958FFA099EFDE2FFD9FFCAC" author="Roewer, C. F." box="[403,561,835,861]" pageId="32" pageNumber="333" pagination="1 - 106" refId="ref32561" refString="Roewer, C. F. (1911) Ubersicht der Genera der Subfamilie der Phalangiini der Opiliones Palpatores nebst Beschreibung einiger neuer Gattungen und Arten. Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten, 77 (Supplementheft 2), 1 - 106, pls. 1 - 3." type="journal article" year="1911">Roewer, 1911</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C35DBAA0958FFA09A0CDE2FFC14FCAC" authority="Gricenko, 1979 a" authorityName="Gricenko" authorityYear="1979" box="[624,954,835,861]" class="Arachnida" family="Phalangiidae" genus="Opilio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="asiaticus">
<emphasis id="B9417C3B0958FFA09A0CDE2FFD58FCAD" box="[624,758,835,860]" italics="true" pageId="32" pageNumber="333">H. asiaticus</emphasis>
<bibRefCitation id="EFA4DDD80958FFA09A81DE2FFC14FCAC" author="Gricenko, N. I. &amp; H. I" box="[765,954,835,861]" pageId="32" pageNumber="333" pagination="28 - 38" refId="ref31057" refString="Gricenko, N. I. [Gрitiнko, H. I.] (1979 a) Сiнokoсtы (Opiliones) iZ aZiatсkoй caсti СССР [Senokoscy (Opiliones) iz azi- atskoj chasti SSSR-Harvestmen (Opiliones) from the Asian part of USSR]. In: Balashov, Y. S. (Ed.), Fauna i ecologiya paukoobraznykh [The Fauna and Ecology of Arachnida]. Trudy Zoologicheskogo Instituta AN SSSR [Proceedings of the Zoological Institute Academy of Sciences of the USSR, Leningrad], 85, pp. 28 - 38. [in Russian]" type="journal article" year="1979">Gricenko, 1979a</bibRefCitation>
</taxonomicName>
, but differs by having significantly smaller body size (
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long) and length of legs (femur II about twice as long as body).
</paragraph>
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<subSubSection id="C32FF3A20958FFA098BBDEE7FC4EFC1C" pageId="32" pageNumber="333" type="materials_examined">
<paragraph id="8B8AA0290958FFA098BBDEE7FF62FC38" blockId="32.[151,1437,798,1869]" pageId="32" pageNumber="333">
<materialsCitation id="3B5DAA740958FFA098BBDEE7FF62FC38" ID-GBIF-Occurrence="3019649327" country="Kyrgyzstan" elevation="3962" latitude="31.973778" location="Chu-Pod River" longLatPrecision="1" longitude="96.988495" municipality="Gor-ly Pass" pageId="32" pageNumber="333" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548E1E8B0958FFA098BBDEE7FEACFC54" box="[199,258,907,933]" pageId="32" pageNumber="333">Type</typeStatus>
Locality
</emphasis>
.
<collectingCountry id="F322E0B90958FFA09905DEE7FE54FC54" box="[377,506,907,933]" name="Kyrgyzstan" pageId="32" pageNumber="333">Kyrgyzstan</collectingCountry>
, headwaters of
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,
<collectingMunicipality id="6BEE3A530958FFA09B1DDEE7FC48FC54" box="[865,998,907,933]" pageId="32" pageNumber="333">Gor-ly Pass</collectingMunicipality>
,
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,
<geoCoordinate id="EE01C6EE0958FFA09CE2DEE6FA92FC54" box="[1182,1340,906,933]" degrees="96" direction="east" minutes="59" orientation="longitude" pageId="32" pageNumber="333" precision="1" seconds="18.6" value="96.988495">96°5918.6”E</geoCoordinate>
,
<elevation id="0018471A0958FFA09D3BDEE7FF62FC38" metricMagnitude="3" metricUnit="m" metricValue="3.9619999999999997" pageId="32" pageNumber="333" unit="m" value="3962.0">
<quantity id="4CCD0DCC0958FFA09D3BDEE7FA32FC54" box="[1351,1436,907,933]" metricMagnitude="3" metricUnit="m" metricValue="3.9619999999999997" pageId="32" pageNumber="333" unit="m" value="3962.0">3962 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
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<emphasis id="B9417C3B0958FFA098BBDEBFFEDFFC1D" bold="true" box="[199,369,978,1005]" pageId="32" pageNumber="333">Type Material</emphasis>
.
<materialsCitation id="3B5DAA740958FFA09901DEBFFC72FC1C" ID-GBIF-Occurrence="3019649376" box="[381,988,979,1005]" collectingDate="1900-08" collectionCode="ZIN" pageId="32" pageNumber="333" specimenCount="2" typeStatus="holotype">
<date id="FF8B86E90958FFA09901DEBFFDA3FC1C" box="[381,525,979,1005]" pageId="32" pageNumber="333" value="1900-08">
<collectingDate id="EFCF7F010958FFA09901DEBFFDA3FC1C" box="[381,525,979,1005]" pageId="32" pageNumber="333" value="1900-08">August 1900</collectingDate>
</date>
(male
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and male
<typeStatus id="548E1E8B0958FFA09B4DDEBFFC38FC1C" box="[817,918,979,1005]" pageId="32" pageNumber="333" type="paratype">Paratype</typeStatus>
,
<collectionCode id="ED2438EC0958FFA09BDDDEBFFC7AFC1C" box="[929,980,979,1005]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="32" pageNumber="333" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32FF3A20958FFA098BBDE9AFBABFBE0" box="[199,1029,1014,1041]" pageId="32" pageNumber="333" type="distribution">
<paragraph id="8B8AA0290958FFA098BBDE9AFBABFBE0" blockId="32.[151,1437,798,1869]" box="[199,1029,1014,1041]" pageId="32" pageNumber="333">
<emphasis id="B9417C3B0958FFA098BBDE9AFEF7FBE1" bold="true" box="[199,345,1014,1040]" pageId="32" pageNumber="333">Distribution</emphasis>
. Only known from the
<typeStatus id="548E1E8B0958FFA09A1CDE9BFD3EFBE0" box="[608,656,1015,1041]" pageId="32" pageNumber="333">type</typeStatus>
locality, in
<collectingCountry id="F322E0B90958FFA09B68DE9BFC3BFBE0" box="[788,917,1015,1041]" name="Kyrgyzstan" pageId="32" pageNumber="333">Kyrgyzstan</collectingCountry>
(
<figureCitation id="130EBCAC0958FFA09BD8DE9BFC56FBE0" box="[932,1016,1015,1041]" captionStart="FIGURE 14" captionStartId="22.[151,250,1974,1998]" captionTargetBox="[179,1408,1081,1941]" captionTargetId="figure-56@22.[173,1415,1074,1950]" captionTargetPageId="22" captionText="FIGURE 14. Distribution of some Homolophus species in the Palearctic: Central and Inner Asia. Data for pale colored symbols taken from literature; identifications not verified." figureDoi="http://doi.org/10.5281/zenodo.4441798" httpUri="https://zenodo.org/record/4441798/files/figure.png" pageId="32" pageNumber="333">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32FF3A20958FFA098BBD977FE4CFB34" pageId="32" pageNumber="333" type="etymology">
<paragraph id="8B8AA0290958FFA098BBD977FE4CFB34" blockId="32.[151,1437,798,1869]" pageId="32" pageNumber="333">
<emphasis id="B9417C3B0958FFA098BBD977FEE6FBC4" bold="true" box="[199,328,1051,1077]" pageId="32" pageNumber="333">Etymology</emphasis>
. The new species is named in honor of Pyotr Kuzmich Kozlov (
<figureCitation id="130EBCAC0958FFA09C52D977FB2CFBC4" box="[1070,1154,1051,1077]" captionStart="FIGURE 24" captionStartId="34.[151,250,988,1012]" captionTargetBox="[170,1436,181,965]" captionTargetId="figure-22@34.[151,1436,181,965]" captionTargetPageId="34" captionText="FIGURE 24. Famous scientists: (left) Pyotr Kuzmich Kozlov (18631935), a Russian traveler, geographer, ethnographer, archeologist, and explorer of Mongolia and Xîzàng (Tibet), for which Homolophus kozlovi sp. nov. is named; (right) Dmitry A. Milko, an entomologist, researcher, and zoologist from Kyrgyzstan for which Homolophus milkoi sp. nov. is named." figureDoi="http://doi.org/10.5281/zenodo.4441825" httpUri="https://zenodo.org/record/4441825/files/figure.png" pageId="32" pageNumber="333">Fig. 24</figureCitation>
), a famous Russian traveler, geographer, ethnographer, archeologist, and explorer of
<collectingCountry id="F322E0B90958FFA09B3AD953FC1DFBA8" box="[838,947,1087,1113]" name="Mongolia" pageId="32" pageNumber="333">Mongolia</collectingCountry>
and Tibet (Xîzàng). He took part in six expeditions mounted by the Imperial Russian Geographical Society. Three of these he organized and led. Specimens collected by him and others on expeditions were studied for this publication. For further details about his life see
<bibRefCitation id="EFA4DDD80958FFA098EBD9C7FE70FB34" author="Andreev, A. I. &amp; Yusupova, T. I." box="[151,478,1194,1221]" pageId="32" pageNumber="333" pagination="1863 - 1935" refId="ref30406" refString="Andreev, A. I., &amp; Yusupova, T. I. (2015) Pyotr Kuz'mich Kozlov: 1863 - 1935. In: Lorimer, H. &amp; Withers, C. W. (Eds.), Geographers Biobibliographical Studies. Vol. 34. Bloomsbury Academic, London, pp. 127 - 164. https: // doi. org / 10.5040 / 9781474251402. ch- 005" type="book chapter" year="2015">Andreev &amp; Yusupova (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32FF3A20958FFA098BBD9A2FE3FF8D8" pageId="32" pageNumber="333" type="description">
<paragraph id="8B8AA0290958FFA098BBD9A2FC6AF8F4" blockId="32.[151,1437,798,1869]" pageId="32" pageNumber="333">
<emphasis id="B9417C3B0958FFA098BBD9A2FEFFFB19" bold="true" box="[199,337,1230,1256]" pageId="32" pageNumber="333">Description</emphasis>
. Medium-sized harvestmen, male body rectangular, with rounded corners; medium sclerotization of body cuticle, less slender, 1.43 times longer than wide; length
<quantity id="4CCD0DCC0958FFA09B01D99FFC79FAFC" box="[893,983,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="32" pageNumber="333" unit="mm" value="4.3">4.3 mm</quantity>
, width 3.0 mm; with transverse rows of tiny to small denticles along borders of tergites, sand color with light and dark spots and splotches (almost all dark areas are where muscles are attached on other side of exoskeleton- see
<figureCitation id="130EBCAC0958FFA09BD1D857FBAAFAA4" box="[941,1028,1339,1365]" captionStart="FIGURE 1" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[212,1360,194,1788]" captionTargetId="figure-22@4.[212,1375,181,1799]" captionTargetPageId="4" captionText="FIGURE 1. Homolophus anatomy, males: A, kozlovi sp. nov., body, dorsal showing muscle attachment sites in black (color photograph Fig. 23); B, gobiensis outline of body showing denticles (compare Fig. 21A); C, betpakdalensis denticles (compare Fig. 12A); D, pallens denticles (compare Fig 30C); E, chemerisi leg I metatarsus with characteristic thickly packed “brushes” of short setae at higher magnification (see insert); F, albofasciatus denticles (compare Fig. 3A); G, nordenskioeldi (from Murmansk) penis, dorsal; muscle presence more evident in magnified sections. Scale lines: AD, F = 1 mm; E = 0.5 mm, insert greatly magnified, G = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4441764" httpUri="https://zenodo.org/record/4441764/files/figure.png" pageId="32" pageNumber="333">Fig. 1A</figureCitation>
); indistinct central longitudinal light colored stripe extends length of body. Ocularium low, round, small, 1.2 times its length from anterior margin of cephalothorax; with 89 denticles on each side, posterior part of ocularium with double rows of these denticles. Preocular area and lateral sides near ocularium with few black-tipped denticles. Legs long, cylindrical in cross-section, segments either with very small, poorly visible denticles, or only setae except for Fe with longitudinal rows of small denticles. Fe not thickened. Leg lengths (mm): I 4.8 + 1.5 + 4.5 + 5.6 + 11.0 = 27.4, II 9.0 + 2.0 + 8.0 + 4.8 + 26.0 = 49.8, III 5.3 + 1.4 + 4.5 + 6.5 + 12.1 = 29.8, IV 7.0 + 1.5 + 5.8 + 6.0 + 21 = 41.3. Pedipalp segment lengths (mm): 1.4 + 0.6 + 1.0 + 1.5 = 4.5. Pedipalps medium-sized to slender, Fe dorsally and ventrally without large denticles, remaining segments with various kinds of setae; Ta ventrally with row of sensory microdenticles. Chelicerae not enlarged, with faint tabby marks, no denticles, only with various kinds of setae. Cheliceral basal segment length
<quantity id="4CCD0DCC0958FFA09D05DB12FF6DF94C" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="333" unit="mm" value="1.5">1.5 mm</quantity>
ventrally with slight rounded bulge, distal segment length
<quantity id="4CCD0DCC0958FFA09B21DBCFFC18F94C" box="[861,950,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="32" pageNumber="333" unit="mm" value="1.7">1.7 mm</quantity>
. Penis wide at base, truncus dorsoventrally flattened, bowed, recurved in lateral view, tapers to glans; glans narrow at base, expanding distally, diamond shaped in anterior view; medium length (mm): truncus 3.0, glans 0.4, stylus 0.1.
</paragraph>
<paragraph id="8B8AA0290958FFA098BBDA63FE3FF8D8" blockId="32.[151,1437,798,1869]" box="[199,401,1807,1833]" pageId="32" pageNumber="333">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C32FF3A20958FFA098BBDA5EFC21F8BC" box="[199,911,1842,1869]" pageId="32" pageNumber="333" type="discussion">
<paragraph id="8B8AA0290958FFA098BBDA5EFC21F8BC" blockId="32.[151,1437,798,1869]" box="[199,911,1842,1869]" pageId="32" pageNumber="333">
<emphasis id="B9417C3B0958FFA098BBDA5EFEF8F8BD" bold="true" box="[199,342,1842,1868]" pageId="32" pageNumber="333">Comments.</emphasis>
Specimen over 100 years old. Colors likely faded.
</paragraph>
</subSubSection>
</treatment>
</document>