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<document id="E75ED33251452BB1096E8B2D919E2B75" ID-CLB-Dataset="29301" ID-DOI="10.11646/zootaxa.4450.4.8" ID-GBIF-Dataset="eb84af6a-57be-44a2-9524-d61b4ced603b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1444988" ID-ZooBank="738C1F16-F3D4-4E0F-A866-C0B5FC4781EE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538635213848" checkinUser="plazi" docAuthor="Chen, Zhi-Teng" docDate="2018" docId="0398C353FFF96F1FEFA5F978FD3C7F8A" docLanguage="en" docName="zootaxa.4450.4.8.pdf" docOrigin="Zootaxa 4450 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Largusoperla borisi Chen 2018" docType="treatment" docVersion="7" lastPageNumber="498" masterDocId="FFA1BB2BFFF86F1CEF32FFC5FF8C7A61" masterDocTitle="Key to the fossil genus Largusoperla (Plecoptera: Perlidae), with description of two new species from mid-Cretaceous Burmese amber" masterLastPageNumber="500" masterPageNumber="495" pageNumber="496" updateTime="1698564498798" updateUser="ExternalLinkService">
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<mods:title id="79D7FBF1EF1DC322F240256A0981FEE0">Key to the fossil genus Largusoperla (Plecoptera: Perlidae), with description of two new species from mid-Cretaceous Burmese amber</mods:title>
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<mods:namePart id="A467F7BBF16E4BA7D75C27E34FA7C8D0">Chen, Zhi-Teng</mods:namePart>
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<mods:title id="AEA307BF5F931004AD86FE263783FFF2">Zootaxa</mods:title>
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<mods:date id="2B32610BA035B8954625AA7456C45187">2018</mods:date>
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<mods:number id="6133984D77B5E0F829503F71E83A867D">2018-07-26</mods:number>
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<mods:identifier id="D8816A8A47C78FC2C4DC548879D3083C" type="DOI">10.11646/zootaxa.4450.4.8</mods:identifier>
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<treatment id="0398C353FFF96F1FEFA5F978FD3C7F8A" ID-DOI="http://doi.org/10.5281/zenodo.5966272" ID-GBIF-Taxon="148462498" ID-Zenodo-Dep="5966272" LSID="urn:lsid:plazi:treatment:0398C353FFF96F1FEFA5F978FD3C7F8A" httpUri="http://treatment.plazi.org/id/0398C353FFF96F1FEFA5F978FD3C7F8A" lastPageId="3" lastPageNumber="498" pageId="1" pageNumber="496">
<subSubSection id="C32B21CEFFF96F1DEFA5F978FE8A7C99" pageId="1" pageNumber="496" type="nomenclature">
<paragraph id="8B8E7245FFF96F1DEFA5F978FDB77CB6" blockId="1.[151,571,1725,1784]" box="[151,571,1725,1751]" pageId="1" pageNumber="496">
<heading id="D0C6C529FFF96F1DEFA5F978FDB77CB6" bold="true" box="[151,571,1725,1751]" fontSize="11" level="1" pageId="1" pageNumber="496" reason="1">
<emphasis id="B945AE57FFF96F1DEFA5F978FDB77CB6" bold="true" box="[151,571,1725,1751]" pageId="1" pageNumber="496">
<taxonomicName id="4C3109C6FFF96F1DEFA5F978FE097CB6" ID-CoL="8NGPJ" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[151,389,1725,1751]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="499" phylum="Arthropoda" rank="species" species="borisi" status="sp. nov.">
<emphasis id="B945AE57FFF96F1DEFA5F978FE097CB6" bold="true" box="[151,389,1725,1751]" italics="true" pageId="1" pageNumber="496">LargUSoperla borISI</emphasis>
</taxonomicName>
Chen, sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E7245FFF96F1DEFA5F91AFE8A7C99" blockId="1.[151,571,1725,1784]" box="[151,262,1759,1784]" pageId="1" pageNumber="496">
<figureCitation id="130A6EC0FFF96F1DEFA5F91AFE8F7C99" box="[151,259,1759,1784]" captionStart="FIGURE 56" captionStartId="4.[151,250,1236,1258]" captionTargetBox="[151,1436,208,1202]" captionTargetId="figure@4.[150,1436,208,1202]" captionTargetPageId="4" captionText="FIGURE 56. Largusoperla borisi, sp. nov. Adult habitus, 5, dorsal view; 6, ventral view." httpUri="https://zenodo.org/record/1444994/files/figure.png" pageId="1" pageNumber="496">Figs. 56</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B21CEFFF96F1FEFA5F8E2FCAA7EE2" lastPageId="3" lastPageNumber="498" pageId="1" pageNumber="496" type="description">
<paragraph id="8B8E7245FFF96F1DEFA5F8E2FA027D21" blockId="1.[151,1436,1831,2037]" box="[151,1422,1831,1856]" pageId="1" pageNumber="496">
<emphasis id="B945AE57FFF96F1DEFA5F8E2FEA57D21" bold="true" box="[151,297,1831,1856]" pageId="1" pageNumber="496">Description.</emphasis>
Macropterous; body length (excluding antennae and cerci) ca. 8.0 mm, generally brown (
<figureCitation id="130A6EC0FFF96F1DEA27F8E2FA0D7D21" box="[1301,1409,1831,1856]" captionStart="FIGURE 56" captionStartId="4.[151,250,1236,1258]" captionTargetBox="[151,1436,208,1202]" captionTargetId="figure@4.[150,1436,208,1202]" captionTargetPageId="4" captionText="FIGURE 56. Largusoperla borisi, sp. nov. Adult habitus, 5, dorsal view; 6, ventral view." httpUri="https://zenodo.org/record/1444994/files/figure.png" pageId="1" pageNumber="496">Figs. 5, 6</figureCitation>
).
</paragraph>
<paragraph id="8B8E7245FFF96F1DEFF5F889FE127DCC" blockId="1.[151,1436,1831,2037]" pageId="1" pageNumber="496">
<emphasis id="B945AE57FFF96F1DEFF5F889FE8B7D04" bold="true" box="[199,263,1868,1893]" pageId="1" pageNumber="496">Head</emphasis>
(
<figureCitation id="130A6EC0FFF96F1DEE24F889FED77D04" box="[278,347,1868,1893]" captionStart="FIGURE 56" captionStartId="4.[151,250,1236,1258]" captionTargetBox="[151,1436,208,1202]" captionTargetId="figure@4.[150,1436,208,1202]" captionTargetPageId="4" captionText="FIGURE 56. Largusoperla borisi, sp. nov. Adult habitus, 5, dorsal view; 6, ventral view." httpUri="https://zenodo.org/record/1444994/files/figure.png" pageId="1" pageNumber="496">Fig. 5</figureCitation>
): Head rounded, mostly covered with a dark stigma, compound eyes dark. Antennae slender and dark brown, with about 46 segments. Maxillary palps slender and three-segmented, longer than the threesegmented labial palps.
</paragraph>
<paragraph id="8B8E7245FFF96F1EEFF5F87DFCBD7AB4" blockId="1.[151,1436,1831,2037]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="497" pageId="1" pageNumber="496">
<emphasis id="B945AE57FFF96F1DEFF5F87DFECC7DB0" bold="true" box="[199,320,1976,2001]" pageId="1" pageNumber="496">Pronotum</emphasis>
(
<figureCitation id="130A6EC0FFF96F1DEE63F87DFE1A7DB0" box="[337,406,1976,2001]" captionStart="FIGURE 56" captionStartId="4.[151,250,1236,1258]" captionTargetBox="[151,1436,208,1202]" captionTargetId="figure@4.[150,1436,208,1202]" captionTargetPageId="4" captionText="FIGURE 56. Largusoperla borisi, sp. nov. Adult habitus, 5, dorsal view; 6, ventral view." httpUri="https://zenodo.org/record/1444994/files/figure.png" pageId="1" pageNumber="496">Fig. 5</figureCitation>
): Pronotum slightly narrower than head, rugose, mostly brown except for the pale median band, near reverse trapezoidal in shape. Meso- and metathorax sclerotized, nearly as wide as pronotum. Legs mostly dark brown except for the pale joints; tibial spurs present; first two tarsal segments shortest with conspicuous euplantulae; apex of the claw dark sclerotized.
</paragraph>
<caption id="DF4E22CDFFFA6F1EEFA5F880FB5F7D3A" box="[151,1235,1861,1883]" httpUri="https://zenodo.org/record/1444990/files/figure.png" pageId="2" pageNumber="497" startId="2.[151,250,1861,1883]" targetBox="[155,1431,270,1836]" targetPageId="2">
<paragraph id="8B8E7245FFFA6F1EEFA5F880FB5F7D3A" blockId="2.[151,1235,1861,1883]" box="[151,1235,1861,1883]" pageId="2" pageNumber="497">
<emphasis id="B945AE57FFFA6F1EEFA5F880FEA07D3B" bold="true" box="[151,300,1861,1883]" pageId="2" pageNumber="497">FIGURE 13.</emphasis>
<taxonomicName id="4C3109C6FFFA6F1EEE00F880FD807D3B" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[306,524,1861,1882]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="dewalti" status="sp. nov.">
<emphasis id="B945AE57FFFA6F1EEE00F880FD807D3B" box="[306,524,1861,1882]" italics="true" pageId="2" pageNumber="497">Largusoperla dewalti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A276132CFFFA6F1EED2BF880FDE57D3B" box="[537,617,1861,1882]" pageId="2" pageNumber="497" rank="species">
<emphasis id="B945AE57FFFA6F1EED2BF880FDE57D3B" bold="true" box="[537,617,1861,1882]" pageId="2" pageNumber="497">sp. nov.</emphasis>
</taxonomicNameLabel>
Adult habitus, 1, dorsal view; 2, ventral view 3, dorsal view.
</paragraph>
</caption>
<paragraph id="8B8E7245FFFA6F1FEFF5F842FCB77B7C" blockId="2.[151,1436,1927,2024]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="498" pageId="2" pageNumber="497">
<emphasis id="B945AE57FFFA6F1EEFF5F842FE9F7DC1" bold="true" box="[199,275,1927,1952]" pageId="2" pageNumber="497">Wings</emphasis>
(
<figureCitation id="130A6EC0FFFA6F1EEE10F842FE017DC1" box="[290,397,1927,1952]" captionStart="FIGURE 56" captionStartId="4.[151,250,1236,1258]" captionTargetBox="[151,1436,208,1202]" captionTargetId="figure@4.[150,1436,208,1202]" captionTargetPageId="4" captionText="FIGURE 56. Largusoperla borisi, sp. nov. Adult habitus, 5, dorsal view; 6, ventral view." httpUri="https://zenodo.org/record/1444994/files/figure.png" pageId="2" pageNumber="497">Figs. 5, 6</figureCitation>
): Wings hyaline, veins dark brown. Left forewing: length ca. 8.0 mm; ScP about three-fourth of forewing length, reaching ScP+RA beyond ra-rp; apical area between anterior margin and RA with one crossvein; about 11 crossveins present between ScP and the anterior margin; RP originated from about one-third of RA and with three branches; M forked at near apical one-third of the wing length; ra-rp not connected with the opposite crossvein rp-ma; CuA forked basal to the fork of M, with three branches; CuP simple; area between M and CuA with six crossveins; area between CuA and CuP with four crossveins in left forewing, five in right forewing; AA1 simple, AA2 forked. Hind wings not visible.
</paragraph>
<caption id="DF4E22CDFFFB6F1FEFA5FC04FCC279B6" box="[151,846,961,983]" httpUri="https://zenodo.org/record/1444992/files/figure.png" pageId="3" pageNumber="498" startId="3.[151,250,961,983]" targetBox="[150,1422,337,940]" targetPageId="3">
<paragraph id="8B8E7245FFFB6F1FEFA5FC04FCC279B6" blockId="3.[151,846,961,983]" box="[151,846,961,983]" pageId="3" pageNumber="498">
<emphasis id="B945AE57FFFB6F1FEFA5FC04FE9E79B7" bold="true" box="[151,274,961,983]" pageId="3" pageNumber="498">FIGURE 4.</emphasis>
<taxonomicName id="4C3109C6FFFB6F1FEE2BFC04FE7F79B7" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[281,499,961,982]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="dewalti" status="sp. nov.">
<emphasis id="B945AE57FFFB6F1FEE2BFC04FE7F79B7" box="[281,499,961,982]" italics="true" pageId="3" pageNumber="498">Largusoperla dewalti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A276132CFFFB6F1FEECDFC04FDDC79B7" box="[511,592,961,982]" pageId="3" pageNumber="498" rank="species">
<emphasis id="B945AE57FFFB6F1FEECDFC04FDDC79B7" bold="true" box="[511,592,961,982]" pageId="3" pageNumber="498">sp. nov.</emphasis>
</taxonomicNameLabel>
Terminalia, ventral view
</paragraph>
</caption>
<paragraph id="8B8E7245FFFB6F1FEFF5FC3BFCAA7EE2" blockId="3.[151,1437,1022,1515]" pageId="3" pageNumber="498">
<emphasis id="B945AE57FFFB6F1FEFF5FC3BFEB07E76" bold="true" box="[199,316,1022,1047]" pageId="3" pageNumber="498">Abdomen</emphasis>
(
<figureCitation id="130A6EC0FFFB6F1FEE79FC3BFE1C7E76" box="[331,400,1022,1047]" captionStart="FIGURE 7" captionStartId="4.[151,250,2012,2034]" captionTargetBox="[150,1406,1323,1971]" captionTargetId="figure@4.[150,1436,1323,1971]" captionTargetPageId="4" captionText="FIGURE 7. Largusoperla borisi, sp. nov. Terminalia, ventral view" httpUri="https://zenodo.org/record/1444996/files/figure.png" pageId="3" pageNumber="498">Fig. 7</figureCitation>
): Abdomen near half the length of the body, lateral coloration darker; posterolateral margins not extended. Sternum 9 extended backwards, hammer not clear visible. Paraproct strongly sclerotized, long triangular, inner margin concave, outer margin convex, apex obtuse. Cerci pale brown, hairy and about 10- segmented, each segment with several long apical bristles.
</paragraph>
</subSubSection>
<subSubSection id="C32B21CEFFFB6F1FEFF5FB4BFA0F7EAA" pageId="3" pageNumber="498" type="materials_examined">
<paragraph id="8B8E7245FFFB6F1FEFF5FB4BFA0F7EAA" blockId="3.[151,1437,1022,1515]" pageId="3" pageNumber="498">
<emphasis id="B945AE57FFFB6F1FEFF5FB4BFEF47EC6" bold="true" box="[199,376,1166,1191]" pageId="3" pageNumber="498">Type material.</emphasis>
Holotype male, No. CZT-PLE-MA4. This specimen is mostly wizened; apex of wings not clear visible, dorsal aspect of abdomen is not clear; wings are overlapped so only the forewings are clear visible.
</paragraph>
</subSubSection>
<subSubSection id="C32B21CEFFFB6F1FEFF5FB13FCBD7F56" pageId="3" pageNumber="498" type="etymology">
<paragraph id="8B8E7245FFFB6F1FEFF5FB13FCD37F72" blockId="3.[151,1437,1022,1515]" pageId="3" pageNumber="498">
<emphasis id="B945AE57FFFB6F1FEFF5FB13FEC27E8E" bold="true" box="[199,334,1238,1263]" pageId="3" pageNumber="498">Etymology.</emphasis>
The new species is named in honor of Dr. Boris Kondratieff,
<collectingRegion id="49F5BCA7FFFB6F1FECCFFB13FBE97E8E" box="[1021,1125,1238,1263]" country="United States of America" name="Colorado" pageId="3" pageNumber="498">Colorado</collectingRegion>
State University,
<collectingRegion id="49F5BCA7FFFB6F1FEA1FFB13FA147E8E" box="[1325,1432,1238,1263]" country="United States of America" name="Colorado" pageId="3" pageNumber="498">Colorado</collectingRegion>
,
<collectingCountry id="F32632D5FFFB6F1FEFA5FB3EFF437F72" box="[151,207,1275,1299]" name="United States of America" pageId="3" pageNumber="498">USA</collectingCountry>
for his assistance in the authors studies of the
<taxonomicName id="4C3109C6FFFB6F1FEDD3FB3FFCD77F72" authorityName="Burmeister" authorityYear="1839" box="[737,859,1274,1299]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.
</paragraph>
<paragraph id="8B8E7245FFFB6F1FEFF5FADBFCBD7F56" blockId="3.[151,1437,1022,1515]" box="[199,817,1310,1335]" pageId="3" pageNumber="498">
<emphasis id="B945AE57FFFB6F1FEFF5FADBFEFD7F56" bold="true" box="[199,369,1310,1335]" pageId="3" pageNumber="498">Syninclusions.</emphasis>
Plant remains and unknown organisms.
</paragraph>
</subSubSection>
<subSubSection id="C32B21CEFFFB6F1FEFF5FA87FD3C7F8A" pageId="3" pageNumber="498" type="discussion">
<paragraph id="8B8E7245FFFB6F1FEFF5FA87FD3C7F8A" blockId="3.[151,1437,1022,1515]" pageId="3" pageNumber="498">
<emphasis id="B945AE57FFFB6F1FEFF5FA87FEB77F3A" bold="true" box="[199,315,1346,1371]" pageId="3" pageNumber="498">Remarks.</emphasis>
This new species also has similar head pattern, body color, wing venation and terminalia structures with other known
<taxonomicName id="4C3109C6FFFB6F1FEE57FAA3FE707F1F" authorityName="Chen, Wang &amp; Du" authorityYear="2018" box="[357,508,1382,1406]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="B945AE57FFFB6F1FEE57FAA3FE707F1F" box="[357,508,1382,1406]" italics="true" pageId="3" pageNumber="498">Largusoperla</emphasis>
</taxonomicName>
species. The strongly sclerotized and sub-triangular paraprocts of the new species is most similar to that of
<taxonomicName id="4C3109C6FFFB6F1FEE84FA4EFD9E7FC3" box="[438,530,1419,1442]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="arcus">
<emphasis id="B945AE57FFFB6F1FEE84FA4EFD9E7FC3" box="[438,530,1419,1442]" italics="true" pageId="3" pageNumber="498">L. arcus</emphasis>
</taxonomicName>
but can be distinguished by the concave inner margin and convex outer margin and obtuse apex of the paraprocts. The paraprocts of
<taxonomicName id="4C3109C6FFFB6F1FEDC4FA6AFCDE7FA7" box="[758,850,1455,1478]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="arcus">
<emphasis id="B945AE57FFFB6F1FEDC4FA6AFCDE7FA7" box="[758,850,1455,1478]" italics="true" pageId="3" pageNumber="498">L. arcus</emphasis>
</taxonomicName>
have a convex inner margin, concave outer margin and sharp apex (see fig.
<quantity id="4CC9DFA0FFFB6F1FEE95FA16FE6C7F8A" box="[423,480,1490,1515]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="3" pageNumber="498" unit="in" value="19.0">19 in</quantity>
Chen
<emphasis id="B945AE57FFFB6F1FED1BFA16FDED7F8B" box="[553,609,1490,1514]" italics="true" pageId="3" pageNumber="498">et al.</emphasis>
2018).
</paragraph>
</subSubSection>
</treatment>
</document>