132 lines
16 KiB
XML
132 lines
16 KiB
XML
<document id="1939D49E31B25EF01D91B884D6221340" ID-CLB-Dataset="29301" ID-DOI="10.11646/zootaxa.4450.4.8" ID-GBIF-Dataset="eb84af6a-57be-44a2-9524-d61b4ced603b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1444988" ID-ZooBank="738C1F16-F3D4-4E0F-A866-C0B5FC4781EE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538635213848" checkinUser="plazi" docAuthor="Chen, Zhi-Teng" docDate="2018" docId="0398C353FFF96F1DEFA5FE6BFEE77C08" docLanguage="en" docName="zootaxa.4450.4.8.pdf" docOrigin="Zootaxa 4450 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Largusoperla dewalti Chen 2018" docType="treatment" docVersion="7" lastPageNumber="496" masterDocId="FFA1BB2BFFF86F1CEF32FFC5FF8C7A61" masterDocTitle="Key to the fossil genus Largusoperla (Plecoptera: Perlidae), with description of two new species from mid-Cretaceous Burmese amber" masterLastPageNumber="500" masterPageNumber="495" pageNumber="496" updateTime="1698564498798" updateUser="ExternalLinkService">
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<mods:mods id="A372DF99B4FB2738D19848ACD7A821FC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2E79A7D803361C0283631816DE8C2EC0">
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<mods:title id="E97E4B1C33EF4A5275BDBB0CAF469068">Key to the fossil genus Largusoperla (Plecoptera: Perlidae), with description of two new species from mid-Cretaceous Burmese amber</mods:title>
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<mods:name id="5FCC8F36E1B1B52BA2501B3F2483A115" type="personal">
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<mods:roleTerm id="3B22D451A7BC5D847243FA731C968CDE">Author</mods:roleTerm>
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<mods:namePart id="2C24D291A447495ED425EEAEAA1B737D">Chen, Zhi-Teng</mods:namePart>
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<mods:typeOfResource id="8FFF885628A6DD4DEE806685C01FAED5">text</mods:typeOfResource>
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<mods:title id="D54F7FE4D9256D6C0C8BFF59D0F0A9A5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E663972FC6560874A1532243FB386DDC">
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<mods:date id="579DF68E7A54E459EAAF05A29799956B">2018</mods:date>
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<mods:number id="5FA6633094946B4F06D998D0187BA11F">2018-07-26</mods:number>
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<mods:detail id="27347DDA26235CC45861243B009DE6CA" type="volume">
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<mods:number id="BF3B39052A5104827DB21D27DF00BC9E">4450</mods:number>
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<mods:detail id="3CC9B6D9B6D212AC4A7A5E398638FD5B" type="issue">
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<mods:number id="D98453A87E4F2C2A3E5DB27765CA9611">4</mods:number>
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<mods:start id="C0FB781C1C0D0B084CBA2486B169453D">495</mods:start>
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<mods:identifier id="5D7437FFF23FBFC8E192C422F9672618" type="DOI">10.11646/zootaxa.4450.4.8</mods:identifier>
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<mods:identifier id="3A623E137250DA07E085C199FBF69CE9" type="GBIF-Dataset">eb84af6a-57be-44a2-9524-d61b4ced603b</mods:identifier>
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<mods:identifier id="7D191A39CFDD31BB547CCC9F5140DDD8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F12C7B864C6943B41FE07178874D0199" type="Zenodo-Dep">1444988</mods:identifier>
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<mods:identifier id="DED28E27EBDE20AFE4E20883808A6470" type="ZooBank">738C1F16-F3D4-4E0F-A866-C0B5FC4781EE</mods:identifier>
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<treatment id="0398C353FFF96F1DEFA5FE6BFEE77C08" ID-DOI="http://doi.org/10.5281/zenodo.5966270" ID-GBIF-Taxon="148462499" ID-Zenodo-Dep="5966270" LSID="urn:lsid:plazi:treatment:0398C353FFF96F1DEFA5FE6BFEE77C08" httpUri="http://treatment.plazi.org/id/0398C353FFF96F1DEFA5FE6BFEE77C08" lastPageNumber="496" pageId="1" pageNumber="496">
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<subSubSection id="C32B21CEFFF96F1DEFA5FE6BFE8A7B88" pageId="1" pageNumber="496" type="nomenclature">
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<paragraph id="8B8E7245FFF96F1DEFA5FE6BFDC17BA9" blockId="1.[151,589,430,490]" box="[151,589,430,456]" pageId="1" pageNumber="496">
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<heading id="D0C6C529FFF96F1DEFA5FE6BFDC17BA9" bold="true" box="[151,589,430,456]" fontSize="11" level="1" pageId="1" pageNumber="496" reason="1">
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<emphasis id="B945AE57FFF96F1DEFA5FE6BFDC17BA9" bold="true" box="[151,589,430,456]" pageId="1" pageNumber="496">
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<taxonomicName id="4C3109C6FFF96F1DEFA5FE6BFE1B7BA9" ID-CoL="8NH3Y" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[151,407,430,456]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="498" phylum="Arthropoda" rank="species" species="dewalti" status="sp. nov.">
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<emphasis id="B945AE57FFF96F1DEFA5FE6BFE1B7BA9" bold="true" box="[151,407,430,456]" italics="true" pageId="1" pageNumber="496">LargUSoperla dewaltI</emphasis>
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</taxonomicName>
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Chen, sp. nov.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8E7245FFF96F1DEFA5FE14FE8A7B88" blockId="1.[151,589,430,490]" box="[151,262,465,490]" pageId="1" pageNumber="496">
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<figureCitation id="130A6EC0FFF96F1DEFA5FE14FE8E7B88" box="[151,258,465,490]" captionStart-0="FIGURE 1–3" captionStart-1="FIGURE 4" captionStartId-0="2.[151,250,1861,1883]" captionStartId-1="3.[151,250,961,983]" captionTargetBox-0="[155,1431,270,1836]" captionTargetBox-1="[150,1422,337,940]" captionTargetId-0="figure@2.[150,1436,265,1840]" captionTargetId-1="figure@3.[150,1436,337,940]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1–3. Largusoperla dewalti, sp. nov. Adult habitus, 1, dorsal view; 2, ventral view 3, dorsal view." captionText-1="FIGURE 4. Largusoperla dewalti, sp. nov. Terminalia, ventral view" httpUri-0="https://zenodo.org/record/1444990/files/figure.png" httpUri-1="https://zenodo.org/record/1444992/files/figure.png" pageId="1" pageNumber="496">Figs. 1–4</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B21CEFFF96F1DEFA5FDDCFE447F63" pageId="1" pageNumber="496" type="description">
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<paragraph id="8B8E7245FFF96F1DEFA5FDDCFA037850" blockId="1.[151,1437,537,1642]" box="[151,1423,537,562]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFA5FDDCFEA57853" bold="true" box="[151,297,537,562]" pageId="1" pageNumber="496">Description.</emphasis>
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Macropterous; body length (excluding antennae and cerci) ca.
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<quantity id="4CC9DFA0FFF96F1DECDEFDDCFBC87850" box="[1004,1092,537,561]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="496" unit="mm" value="9.1">9.1 mm</quantity>
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, generally brown (
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<figureCitation id="130A6EC0FFF96F1DEA27FDDCFA0D7850" box="[1301,1409,537,562]" captionStart="FIGURE 1–3" captionStartId="2.[151,250,1861,1883]" captionTargetBox="[155,1431,270,1836]" captionTargetId="figure@2.[150,1436,265,1840]" captionTargetPageId="2" captionText="FIGURE 1–3. Largusoperla dewalti, sp. nov. Adult habitus, 1, dorsal view; 2, ventral view 3, dorsal view." httpUri="https://zenodo.org/record/1444990/files/figure.png" pageId="1" pageNumber="496">Figs. 1, 2</figureCitation>
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).
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</paragraph>
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<paragraph id="8B8E7245FFF96F1DEFF5FDF8FF7A78FF" blockId="1.[151,1437,537,1642]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5FDF8FE8B7837" bold="true" box="[199,263,573,598]" pageId="1" pageNumber="496">Head</emphasis>
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(
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<figureCitation id="130A6EC0FFF96F1DEE24FDF8FE087837" box="[278,388,573,598]" captionStart="FIGURE 1–3" captionStartId="2.[151,250,1861,1883]" captionTargetBox="[155,1431,270,1836]" captionTargetId="figure@2.[150,1436,265,1840]" captionTargetPageId="2" captionText="FIGURE 1–3. Largusoperla dewalti, sp. nov. Adult habitus, 1, dorsal view; 2, ventral view 3, dorsal view." httpUri="https://zenodo.org/record/1444990/files/figure.png" pageId="1" pageNumber="496">Figs. 1–3</figureCitation>
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): Head oblate, mostly covered with a dark stigma, compound eyes dark. Antennae hairy and dark brown, with at least 29 segments. Maxillary palps not preserved; labial palps three-segmented, basal segment thickest.
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</paragraph>
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<paragraph id="8B8E7245FFF96F1DEFF5FD6CFE947933" blockId="1.[151,1437,537,1642]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5FD6CFECC78A3" bold="true" box="[199,320,681,706]" pageId="1" pageNumber="496">Pronotum</emphasis>
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(
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<figureCitation id="130A6EC0FFF96F1DEE7CFD6CFE1E78A3" box="[334,402,681,706]" captionStart="FIGURE 1–3" captionStartId="2.[151,250,1861,1883]" captionTargetBox="[155,1431,270,1836]" captionTargetId="figure@2.[150,1436,265,1840]" captionTargetPageId="2" captionText="FIGURE 1–3. Largusoperla dewalti, sp. nov. Adult habitus, 1, dorsal view; 2, ventral view 3, dorsal view." httpUri="https://zenodo.org/record/1444990/files/figure.png" pageId="1" pageNumber="496">Fig. 3</figureCitation>
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): Pronotum nearly trapezoidal, rugose and mostly brown, much narrower than head, anterior margin wider than posterior margin, median area not visible. Meso- and metathorax sclerotized, nearly as wide as pronotum. Legs mostly dark brown except for the pale joints; tibia with several thick ventral spines, the paired tibial spurs present; first two tarsal segments shortest with conspicuous euplantulae; apex of the claw dark sclerotized.
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</paragraph>
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<paragraph id="8B8E7245FFF96F1DEFF5FC98FDDB7E2F" blockId="1.[151,1437,537,1642]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5FC98FE9F7917" bold="true" box="[199,275,861,886]" pageId="1" pageNumber="496">Wings</emphasis>
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(
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<figureCitation id="130A6EC0FFF96F1DEE17FC98FE187917" box="[293,404,861,886]" captionStart="FIGURE 1–3" captionStartId="2.[151,250,1861,1883]" captionTargetBox="[155,1431,270,1836]" captionTargetId="figure@2.[150,1436,265,1840]" captionTargetPageId="2" captionText="FIGURE 1–3. Largusoperla dewalti, sp. nov. Adult habitus, 1, dorsal view; 2, ventral view 3, dorsal view." httpUri="https://zenodo.org/record/1444990/files/figure.png" pageId="1" pageNumber="496">Figs. 1–2</figureCitation>
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): Wings hyaline, veins dark brown. Left forewing: length ca. 11.0 mm; ScP about threefourth of forewing length, reaching ScP+RA beyond ra-rp; apical area between anterior margin and RA with one crossvein; about nine crossveins present between ScP and the anterior margin; RP originated from about one-third of RA and with three branches; M forked at near half of the wing length; ra-rp not connected with the opposite crossvein rp-ma; CuA forked basal to the fork of M, with three branches; CuP simple; area between M and CuA with six crossveins; area between CuA and CuP with five crossveins; AA1 simple, AA2 forked. Hind wings covered by forewings, veins not visible.
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</paragraph>
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<paragraph id="8B8E7245FFF96F1DEFF5FB9CFE447F63" blockId="1.[151,1437,537,1642]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5FB9CFEB07E13" bold="true" box="[199,316,1113,1138]" pageId="1" pageNumber="496">Abdomen</emphasis>
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(
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<figureCitation id="130A6EC0FFF96F1DEE79FB9CFE1C7E13" box="[331,400,1113,1138]" captionStart="FIGURE 4" captionStartId="3.[151,250,961,983]" captionTargetBox="[150,1422,337,940]" captionTargetId="figure@3.[150,1436,337,940]" captionTargetPageId="3" captionText="FIGURE 4. Largusoperla dewalti, sp. nov. Terminalia, ventral view" httpUri="https://zenodo.org/record/1444992/files/figure.png" pageId="1" pageNumber="496">Fig. 4</figureCitation>
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): Abdomen approximately ½ length of the body, generally brown, laterally with dark marks; sterna 6-8 with a pale oval mark; posterolateral margins of each segment slightly extruded. Sternum 9 strongly extended backwards, posteromedially with an elliptical hammer. Paraproct thick, hairy and finger-shaped, unevenly sclerotized with a banded pattern, apex rounded. Cerci hairy and pale,10-segmented, each segment with several dark apical bristles.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B21CEFFF96F1DEFF5FAC8FC397F0F" pageId="1" pageNumber="496" type="materials_examined">
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<paragraph id="8B8E7245FFF96F1DEFF5FAC8FC397F0F" blockId="1.[151,1437,537,1642]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5FAC8FEF97F47" bold="true" box="[199,373,1293,1318]" pageId="1" pageNumber="496">Type material.</emphasis>
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Holotype male, No. CZT-PLE-MA3. This specimen is relatively well preserved except for the apex of the antennae, forelegs and right mid-leg, and the maxillary palps. The dorsal aspect of abdomen is not clear; wings are overlapping with only the left forewing clearly visible.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B21CEFFF96F1DEFF5FABCFCBD7F9F" pageId="1" pageNumber="496" type="etymology">
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<paragraph id="8B8E7245FFF96F1DEFF5FABCFF697FBB" blockId="1.[151,1437,537,1642]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5FABCFEC27FF3" bold="true" box="[199,334,1401,1426]" pageId="1" pageNumber="496">Etymology.</emphasis>
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The new species is named in honor of Dr. R. Edward DeWalt,
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<collectingRegion id="49F5BCA7FFF96F1DEB01FABCFB0E7FF3" box="[1075,1154,1401,1426]" country="United States of America" name="Illinois" pageId="1" pageNumber="496">Illinois</collectingRegion>
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Natural History Survey, Champaign,
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<collectingRegion id="49F5BCA7FFF96F1DEE14FA58FEF47FD7" box="[294,376,1437,1462]" country="United States of America" name="Illinois" pageId="1" pageNumber="496">Illinois</collectingRegion>
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,
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<collectingCountry id="F32632D5FFF96F1DEEB7FA5BFE317FD7" box="[389,445,1438,1462]" name="United States of America" pageId="1" pageNumber="496">USA</collectingCountry>
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for his enthusiasm for the study of the
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<taxonomicName id="4C3109C6FFF96F1DEC48FA58FC7C7FD7" authorityName="Burmeister" authorityYear="1839" box="[890,1008,1437,1462]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="496" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
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and for his generous assistance to the author.
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</paragraph>
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<paragraph id="8B8E7245FFF96F1DEFF5FA20FCBD7F9F" blockId="1.[151,1437,537,1642]" box="[199,817,1509,1534]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5FA20FEFD7F9F" bold="true" box="[199,369,1509,1534]" pageId="1" pageNumber="496">Syninclusions.</emphasis>
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Plant remains and unknown organisms.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B21CEFFF96F1DEFF5F9CCFEE77C08" pageId="1" pageNumber="496" type="discussion">
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<paragraph id="8B8E7245FFF96F1DEFF5F9CCFEE77C08" blockId="1.[151,1437,537,1642]" pageId="1" pageNumber="496">
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<emphasis id="B945AE57FFF96F1DEFF5F9CCFEB77C43" bold="true" box="[199,315,1545,1570]" pageId="1" pageNumber="496">Remarks.</emphasis>
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The new species exhibits typical head pattern, body color, similar wing venation and terminalia structures of other known species of
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<taxonomicName id="4C3109C6FFF96F1DED09F9E8FD5F7C24" authorityName="Chen, Wang & Du" authorityYear="2018" box="[571,723,1581,1605]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="496" phylum="Arthropoda" rank="genus">
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<emphasis id="B945AE57FFF96F1DED09F9E8FD5F7C24" box="[571,723,1581,1605]" italics="true" pageId="1" pageNumber="496">Largusoperla</emphasis>
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</taxonomicName>
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but
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<taxonomicName id="4C3109C6FFF96F1DEC35F9EBFCF47C24" authorityName="Chen" authorityYear="2018" box="[775,888,1581,1605]" class="Insecta" family="Perlidae" genus="Largusoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="496" phylum="Arthropoda" rank="species" species="dewalti">
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<emphasis id="B945AE57FFF96F1DEC35F9EBFCF47C24" box="[775,888,1581,1605]" italics="true" pageId="1" pageNumber="496">L. dewalti</emphasis>
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</taxonomicName>
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is easily distinguished by the finger-shaped and banded paraprocts.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |