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<document id="3799D34685B0847C344E3E815D6A48D4" ID-CLB-Dataset="26165" ID-DOI="10.11646/zootaxa.4640.1" ID-GBIF-Dataset="1c74afbc-0c31-4b2b-b975-af79979613ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="3342242" ID-ZooBank="434B2D49-EC2C-4246-8703-42C6CFD31B87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563526702478" checkinUser="plazi" docAuthor="Grand, Daniel, Marinov, Milen, Jourdan, Herve, Cook, Carl, Rouys, Sophie, Mille, Christian &amp; Theuerkauf, Jörn" docDate="2019" docId="03913978FFFAFFDFFF1ABE4A654CFABC" docLanguage="en" docName="zootaxa.4640.1.1.pdf" docOrigin="Zootaxa 4640 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Rhyothemis phyllis subsp. apicalis Kirby 1889" docType="treatment" docVersion="12" lastPageNumber="92" masterDocId="FFA84100FFA0FF84FF8DB837610AFFD7" masterDocTitle="Distribution, habitats, phenology and conservation of New Caledonian Odonata" masterLastPageNumber="112" masterPageNumber="1" pageNumber="91" updateTime="1698745996437" updateUser="ExternalLinkService">
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<mods:title id="B5F3C8E9D828C7436BAA544594A0A780">Distribution, habitats, phenology and conservation of New Caledonian Odonata</mods:title>
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<paragraph id="8B87886EFFFAFFDEFF1ABE4A6376F940" blockId="90.[151,636,1661,1724]" box="[151,636,1661,1688]" pageId="90" pageNumber="91">
<heading id="D0CF3F02FFFAFFDEFF1ABE4A6376F940" bold="true" box="[151,636,1661,1688]" fontSize="11" level="1" pageId="90" pageNumber="91" reason="1">
<taxonomicName id="4C38F3EDFFFAFFDEFF1ABE4A6376F940" ID-CoL="7J376" authority="Kirby, 1889" authorityName="Kirby" authorityYear="1889" box="[151,636,1661,1688]" class="Insecta" family="Libellulidae" genus="Rhyothemis" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="subSpecies" species="phyllis" subSpecies="apicalis">
<emphasis id="B94C547CFFFAFFDEFF1ABE4A6376F940" bold="true" box="[151,636,1661,1688]" pageId="90" pageNumber="91">
<emphasis id="B94C547CFFFAFFDEFF1ABE4A60E8F94F" bold="true" box="[151,482,1661,1688]" italics="true" pageId="90" pageNumber="91">Rhyothemis phyllis apicalis</emphasis>
<bibRefCitation id="EFA9F59FFFFAFFDEFE64BE4A6376F940" author="Kirby, W." box="[489,636,1661,1687]" pageId="90" pageNumber="91" pagination="249 - 348" refId="ref33490" refString="Kirby, W. (1889) A revision of the subfamily Libellulinae, with descriptions of new genera and species. Transactions of the Zoological Society of London, 12 (9), 249 - 348. https: // doi. org / 10.1111 / j. 1096 - 3642.1889. tb 00016. x" type="journal article" year="1889">Kirby, 1889</bibRefCitation>
</emphasis>
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<paragraph id="8B87886EFFFAFFDEFF1ABE9661F3F96B" blockId="90.[151,636,1661,1724]" box="[151,249,1697,1724]" pageId="90" pageNumber="91">
(
<figureCitation id="130394EBFFFAFFDEFF2DBE9661FBF96B" box="[160,241,1697,1724]" captionStart="FIGURE 98" captionStartId="92.[151,250,1994,2019]" captionTargetBox="[151,1436,265,1974]" captionTargetId="figure@92.[151,1436,265,1974]" captionTargetPageId="92" captionText="FIGURE 98. Rhyothemis phyllis apicalis: a) male, b) female." httpUri="https://zenodo.org/record/3342430/files/figure.png" pageId="90" pageNumber="91">Fig. 98</figureCitation>
)
</paragraph>
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<subSubSection id="C322DBE5FFFAFFDEFF1ABED1643CF929" box="[151,1334,1765,1790]" pageId="90" pageNumber="91" type="reference_group">
<paragraph id="8B87886EFFFAFFDEFF1ABED1643CF929" blockId="90.[151,1334,1765,1790]" box="[151,1334,1765,1790]" pageId="90" pageNumber="91">
<treatmentCitationGroup id="AB28AF40FFFAFFDEFF1ABED1643CF929" box="[151,1334,1765,1790]" pageId="90" pageNumber="91">
<taxonomicName id="4C38F3EDFFFAFFDEFF1ABED160CFF929" ID-CoL="9X3CK" authority="Sulzer, 1776" authorityName="Sulzer" authorityYear="1776" box="[151,453,1765,1790]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="phyllis">
<emphasis id="B94C547CFFFAFFDEFF1ABED16037F929" box="[151,317,1766,1790]" italics="true" pageId="90" pageNumber="91">Libellula phyllis</emphasis>
Sulzer, 1776
</taxonomicName>
;
<taxonomicName id="4C38F3EDFFFAFFDEFE42BED162ACF929" ID-CoL="9X4B3" authority="Selys-Longchamps, 1878" authorityName="Selys-Longchamps" authorityYear="1878" box="[463,934,1765,1790]" class="Insecta" family="Libellulidae" genus="Rhyothemis" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="snelleni">
<emphasis id="B94C547CFFFAFFDEFE42BED16391F929" box="[463,667,1766,1790]" italics="true" pageId="90" pageNumber="91">Rhyothemis snelleni</emphasis>
<bibRefCitation id="EFA9F59FFFFAFFDEFD2CBED262ACF929" author="Selys-Longchamps, E. de" box="[673,934,1765,1790]" pageId="90" pageNumber="91" pagination="289 - 323" refId="ref35964" refString="Selys-Longchamps, E. de. (1878) Odonates de la Region de la Nouvelle Guinee. Mitteilungen aus dem K ˆ niglich Zoologischen Museum zu Dresden, 3, 289 - 323." type="journal article" year="1878">Selys-Longchamps, 1878</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C38F3EDFFFAFFDEFC3DBED16438F929" ID-CoL="6X7H2" authority="Kirby, 1889: 319" authorityName="Kirby" authorityPageNumber="319" authorityYear="1889" box="[944,1330,1765,1790]" class="Insecta" family="Libellulidae" genus="Rhyothemis" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B94C547CFFFAFFDEFC3DBED16576F929" box="[944,1148,1766,1790]" italics="true" pageId="90" pageNumber="91">Rhyothemis apicalis</emphasis>
<treatmentCitation id="0A99AE7FFFFAFFDEFB0EBED26438F929" author="Kirby, W." box="[1155,1330,1765,1790]" page="319" pageId="90" pageNumber="91" year="1889">
<bibRefCitation id="EFA9F59FFFFAFFDEFB0EBED26438F929" author="Kirby, W." box="[1155,1330,1765,1790]" pageId="90" pageNumber="91" pagination="249 - 348" refId="ref33490" refString="Kirby, W. (1889) A revision of the subfamily Libellulinae, with descriptions of new genera and species. Transactions of the Zoological Society of London, 12 (9), 249 - 348. https: // doi. org / 10.1111 / j. 1096 - 3642.1889. tb 00016. x" type="journal article" year="1889">Kirby, 1889: 319</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
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<paragraph id="8B87886EFFFAFFDEFF1ABF1C6357F82D" blockId="90.[151,1437,1835,2043]" pageId="90" pageNumber="91">
<emphasis id="B94C547CFFFAFFDEFF1ABF1C6023F891" bold="true" box="[151,297,1835,1862]" pageId="90" pageNumber="91">Distribution</emphasis>
(
<figureCitation id="130394EBFFFAFFDEFEB4BF1B6087F891" box="[313,397,1836,1862]" captionStart="FIGURE 99" captionStartId="93.[151,250,1091,1116]" captionTargetBox="[151,1405,166,1071]" captionTargetId="figure@93.[151,1405,166,1071]" captionTargetPageId="93" captionText="FIGURE 99. Records of Rhyothemis phyllis apicalis." httpUri="https://zenodo.org/record/3342432/files/figure.png" pageId="90" pageNumber="91">Fig. 99</figureCitation>
). Regional endemic subspecies.
<taxonomicName id="4C38F3EDFFFAFFDEFD75BF1B62B4F891" box="[760,958,1836,1862]" class="Insecta" genus="Ryothemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="phyllis">
<emphasis id="B94C547CFFFAFFDEFD75BF1B62B4F891" box="[760,958,1836,1862]" italics="true" pageId="90" pageNumber="91">Ryothemis phyllis</emphasis>
</taxonomicName>
, which includes about a dozen subspecies, is widespread in south-east Asia, north-eastern
<collectingCountry id="F32FC8FEFFFAFFDEFD28BF676206F8BD" box="[677,780,1872,1898]" name="Australia" pageId="90" pageNumber="91">Australia</collectingCountry>
and the western Pacific Islands (
<bibRefCitation id="EFA9F59FFFFAFFDEFBF7BF676409F8BD" author="Tsuda, S." box="[1146,1283,1872,1898]" pageId="90" pageNumber="91" refId="ref36559" refString="Tsuda, S. (2000) A distributional list of world Odonata. Osaka, Japan, 430 pp." type="book" year="2000">Tsuda 2000</bibRefCitation>
). The subspecies
<taxonomicName id="4C38F3EDFFFAFFDEFF44BF426098F859" box="[201,402,1908,1934]" class="Insecta" genus="Ryothemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="subSpecies" species="phyllis" subSpecies="apicalis">
<emphasis id="B94C547CFFFAFFDEFF44BF426098F859" box="[201,402,1908,1934]" italics="true" pageId="90" pageNumber="91">R. phyllis apicalis</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F32FC8FEFFFAFFDEFDBEBF426385F858" box="[563,655,1909,1935]" name="Vanuatu" pageId="90" pageNumber="91">Vanuatu</collectingCountry>
and
<collectingCountry id="F32FC8FEFFFAFFDEFD49BF42627BF859" box="[708,881,1908,1935]" name="New Caledonia" pageId="90" pageNumber="91">New Caledonia</collectingCountry>
where it is recorded from more than 25 localities, in sometimes large populations. One specimen from Nouméa is preserved in the
<collectionCode id="ED2910ABFFFAFFDEFBAABFAF657CF865" box="[1063,1142,1944,1970]" country="New Caledonia" httpUri="http://grbio.org/cool/crvj-94ys" name="Office de la Recherche Scientifique et Technique d'Outre-Mer" pageId="90" pageNumber="91">ONNC</collectionCode>
collection and others collected in
<quantity id="4CC0258BFFFAFFDEFF70BF8B605BF801" box="[253,337,1980,2006]" metricMagnitude="1" metricUnit="m" metricValue="5.07238" pageId="90" pageNumber="91" unit="in" value="1997.0">1997 in</quantity>
the municipality of La Foa (Pocquereux valley) and in
<quantity id="4CC0258BFFFAFFDEFC34BF8B6507F801" box="[953,1037,1980,2006]" metricMagnitude="1" metricUnit="m" metricValue="5.097779999999999" pageId="90" pageNumber="91" unit="in" value="2007.0">2007 in</quantity>
the municipality of Poya (Beaupré) are deposited in the
<collectionCode id="ED2910ABFFFAFFDEFEF4BFD760EAF82D" box="[377,480,2016,2042]" pageId="90" pageNumber="91">CXMNC</collectionCode>
collection.
</paragraph>
<caption id="DF47D8E6FFFBFFDFFF1ABC05639CFB9D" box="[151,662,1074,1099]" httpUri="https://zenodo.org/record/3342428/files/figure.png" pageId="91" pageNumber="92" startId="91.[151,250,1074,1099]" targetBox="[178,1408,166,1054]" targetPageId="91">
<paragraph id="8B87886EFFFBFFDFFF1ABC05639CFB9D" blockId="91.[151,662,1074,1099]" box="[151,662,1074,1099]" pageId="91" pageNumber="92">
<emphasis id="B94C547CFFFBFFDFFF1ABC05602AFB9C" bold="true" box="[151,288,1074,1099]" pageId="91" pageNumber="92">FIGURE 97.</emphasis>
Records of
<taxonomicName id="4C38F3EDFFFBFFDFFE16BC04639AFB9C" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[411,656,1075,1099]" class="Insecta" family="Libellulidae" genus="Rhyothemis" kingdom="Animalia" order="Odonata" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="graphiptera">
<emphasis id="B94C547CFFFBFFDFFE16BC04639AFB9C" box="[411,656,1075,1099]" italics="true" pageId="91" pageNumber="92">Rhyothemis graphiptera</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B87886EFFFBFFDFFF4ABC4F604BFB60" blockId="91.[151,1436,1144,1387]" pageId="91" pageNumber="92">
<emphasis id="B94C547CFFFBFFDFFF4ABC4F6020FB44" bold="true" box="[199,298,1144,1171]" pageId="91" pageNumber="92">Habitat.</emphasis>
This species is encountered in about the same habitats as
<taxonomicName id="4C38F3EDFFFBFFDFFC3FBC4D655DFB44" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[946,1111,1145,1171]" class="Insecta" family="Libellulidae" genus="Rhyothemis" kingdom="Animalia" order="Odonata" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="graphiptera">
<emphasis id="B94C547CFFFBFFDFFC3FBC4D655DFB44" box="[946,1111,1145,1171]" italics="true" pageId="91" pageNumber="92">R. graphiptera</emphasis>
</taxonomicName>
and, like this species, avoids ultramafic soil.
</paragraph>
<paragraph id="8B87886EFFFBFFDFFF4ABCF763E0FA90" blockId="91.[151,1436,1144,1387]" pageId="91" pageNumber="92">
<emphasis id="B94C547CFFFBFFDFFF4ABCF760D4FB0C" bold="true" box="[199,478,1216,1243]" pageId="91" pageNumber="92">Biology and behaviour.</emphasis>
This species does not seem particularly territorial. Males seek females flying along banks at the edge of the water. Mating takes place on low vegetation and lasts a few minutes, then the female lays her eggs unaccompanied in areas of dense aquatic vegetation. During a rapid and frenetic flight, she releases her eggs, touching the surface of the water several times.
</paragraph>
<paragraph id="8B87886EFFFBFFDFFF4ABD67654CFABC" blockId="91.[151,1436,1144,1387]" box="[199,1094,1360,1387]" pageId="91" pageNumber="92">
<emphasis id="B94C547CFFFBFFDFFF4ABD676013FABC" bold="true" box="[199,281,1360,1387]" pageId="91" pageNumber="92">Status.</emphasis>
<emphasis id="B94C547CFFFBFFDFFEADBD66634CFABC" box="[288,582,1361,1387]" italics="true" pageId="91" pageNumber="92">
<taxonomicName id="4C38F3EDFFFBFFDFFEADBD6660ECFABC" box="[288,486,1361,1387]" class="Insecta" genus="Ryothemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="phyllis">Ryothemis phyllis</taxonomicName>
<taxonomicName id="4C38F3EDFFFBFFDFFE60BD66634CFABC" authorityName="Kirby" authorityYear="1889" box="[493,582,1361,1387]" class="Insecta" family="Libellulidae" genus="Rhyothemis" kingdom="Animalia" order="Odonata" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="apicalis">apicalis</taxonomicName>
</emphasis>
does not appear threatened, we rank it as LC.
</paragraph>
</subSubSection>
</treatment>
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