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<document id="BD8935AA4C41C4AB571EC9AD19C7B601" ID-DOI="10.5281/zenodo.294105" ID-GBIF-Dataset="ba12cd1a-dea5-48d8-bb50-978cebcef3d7" ID-ISSN="1175-5326" ID-Zenodo-Dep="294105" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460242242210" checkinUser="plazi" docAuthor="Lis, Jerzy A. &amp; Zack, Richard S." docDate="2010" docId="038E87ECFFEBFFB749FCFA5D2E0E1F5F" docLanguage="en" docName="zt02523p064.pdf" docOrigin="Zootaxa 2523" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Byrsinus varians Fabricius 1803" docType="treatment" docVersion="7" lastPageNumber="60" masterDocId="FFB7FF94FFEAFFB4496BFF972C421D13" masterDocTitle="A review of burrower bugs (Hemiptera: Heteroptera: Cydnidae sensu lato) of Guam" masterLastPageNumber="64" masterPageNumber="57" pageNumber="58" updateTime="1698241056831" updateUser="plazi">
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<mods:title id="8840B507EC92498CF7D010136AEC806C">A review of burrower bugs (Hemiptera: Heteroptera: Cydnidae sensu lato) of Guam</mods:title>
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<mods:namePart id="650AAEE92825EC82A256D67C159C3C13">Lis, Jerzy A.</mods:namePart>
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<treatment id="038E87ECFFEBFFB749FCFA5D2E0E1F5F" ID-DOI="http://doi.org/10.5281/zenodo.6211209" ID-GBIF-Taxon="119389926" ID-Zenodo-Dep="6211209" LSID="urn:lsid:plazi:treatment:038E87ECFFEBFFB749FCFA5D2E0E1F5F" httpUri="http://treatment.plazi.org/id/038E87ECFFEBFFB749FCFA5D2E0E1F5F" lastPageId="3" lastPageNumber="60" pageId="1" pageNumber="58">
<subSubSection id="C33D6571FFEBFFB549FCFA5D2D711B14" pageId="1" pageNumber="58" type="nomenclature">
<paragraph id="8B9836FAFFEBFFB549FCFA5D2F0C18F7" blockId="1.[151,846,1482,1543]" box="[151,846,1482,1508]" pageId="1" pageNumber="58">
<heading id="D0D08196FFEBFFB549FCFA5D2F0C18F7" bold="true" box="[151,846,1482,1508]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
<emphasis id="B953EAE8FFEBFFB549FCFA5D2F0C18F7" bold="true" box="[151,846,1482,1508]" pageId="1" pageNumber="58">
<taxonomicName id="4C274D79FFEBFFB549FCFA5D2E7D18F7" authority="Fabricius, 1803" authorityName="Fabricius" authorityYear="1803" box="[151,575,1482,1508]" class="Insecta" family="Cydnidae" genus="Byrsinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B953EAE8FFEBFFB549FCFA5D2D2618F7" bold="true" box="[151,356,1482,1508]" italics="true" pageId="1" pageNumber="58">Byrsinus varians</emphasis>
(Fabricius, 1803)
</taxonomicName>
, first territory record
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9836FAFFEBFFB549FCFA7A2D711B14" blockId="1.[151,846,1482,1543]" box="[151,307,1517,1543]" pageId="1" pageNumber="58">
(
<figureCitation id="131C2A7FFFEBFFB549F4FA7A2D4C1B14" box="[159,270,1517,1543]" captionStart="FIGURES 1 8" captionStartId="2.[151,270,1880,1904]" captionTargetBox="[151,1436,197,1848]" captionTargetId="figure@2.[151,1436,197,1850]" captionTargetPageId="2" captionText="FIGURES 1 8. Head in two dorsal views. 1, 3, 5, 7 — showing head shape and puncturation, 2, 4, 6, 8 — showing head vestiture." httpUri="https://zenodo.org/record/294106/files/figure.png" pageId="1" pageNumber="58">Figs. 12</figureCitation>
,
<figureCitation id="131C2A7FFFEBFFB54870FA7A2D681B14" box="[283,298,1517,1543]" captionStart="FIGURES 9 12" captionStartId="4.[151,269,1645,1669]" captionTargetBox="[151,1434,346,1622]" captionTargetId="figure@4.[151,1436,346,1623]" captionTargetPageId="4" captionText="FIGURES 9 12. Evaporative areas on meso- and metapleuron. 9 — Byrsinus varians, 10 — Fromundus biimpressus, 11 — F. pygmaeus, 12 — Rhytidoporus indentatus." httpUri="https://zenodo.org/record/294107/files/figure.png" pageId="1" pageNumber="58">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33D6571FFEBFFB549FCF9AF2FE11A47" pageId="1" pageNumber="58" type="reference_group">
<paragraph id="8B9836FAFFEBFFB549FCF9AF2F551B5C" blockId="1.[151,791,1592,1615]" box="[151,791,1592,1615]" pageId="1" pageNumber="58">
<treatmentCitationGroup id="AB3711D4FFEBFFB549FCF9AF2F551B5C" box="[151,791,1592,1615]" pageId="1" pageNumber="58">
<taxonomicName id="4C274D79FFEBFFB549FCF9AF2E321B5C" authority="Froeschner, 1976: 232" authorityName="Froeschner" authorityPageNumber="232" authorityYear="1976" box="[151,624,1592,1615]" class="Diplopoda" family="Dalodesmidae" genus="Microporus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="shiromai">
<emphasis id="B953EAE8FFEBFFB549FCF9AF2D3A1B5C" box="[151,376,1592,1615]" italics="true" pageId="1" pageNumber="58">Microporus shiromai</emphasis>
<treatmentCitation id="0A8610EBFFEBFFB54814F9AF2E321B5C" author="Froeschner" box="[383,624,1592,1615]" page="232" pageId="1" pageNumber="58" year="1976">
<bibRefCitation id="EFB64B0BFFEBFFB54814F9AF2E791B5C" author="Froeschner" box="[383,571,1592,1615]" pageId="1" pageNumber="63" refString="Froeschner, R. C. (1976) The burrowing bugs of Hawaii, with description of a new species (Hemiptera: Cydnidae)." type="book" year="1976">Froeschner, 1976</bibRefCitation>
: 232
</treatmentCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A2605793FFEBFFB54B11F9AF2F501B5C" box="[634,786,1592,1615]" pageId="1" pageNumber="58">new synonym</taxonomicNameLabel>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9836FAFFEBFFB549FCF9EA2FE11A47" blockId="1.[151,1437,1661,2035]" pageId="1" pageNumber="58">
<emphasis id="B953EAE8FFEBFFB549FCF9EA2DFF1B84" bold="true" box="[151,445,1661,1687]" pageId="1" pageNumber="58">Diagnostic characters:</emphasis>
Body from pale brown to blackish brown,
<quantity id="4CDF9B1FFFEBFFB54ABEF9EA282B1B84" box="[981,1129,1661,1687]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.5" metricValueMin="3.4" pageId="1" pageNumber="58" unit="mm" value="3.95" valueMax="4.5" valueMin="3.4">3.44.5 mm</quantity>
in length. Head sparsely punctured; clypeus subapically with two peglike setae (
<figureCitation id="131C2A7FFFEBFFB54A43F9352FD81BAF" box="[808,922,1698,1724]" captionStart="FIGURES 1 8" captionStartId="2.[151,270,1880,1904]" captionTargetBox="[151,1436,197,1848]" captionTargetId="figure@2.[151,1436,197,1850]" captionTargetPageId="2" captionText="FIGURES 1 8. Head in two dorsal views. 1, 3, 5, 7 — showing head shape and puncturation, 2, 4, 6, 8 — showing head vestiture." httpUri="https://zenodo.org/record/294106/files/figure.png" pageId="1" pageNumber="58">Figs. 12</figureCitation>
); each paraclypeus with a submarginal row of 1012 setigerous punctures (78 peglike setae, and 35 hairlike setae); 2nd antennal segment shorter than 3rd segment. Pronotum moderately punctured, each lateral margin with numerous (1940) submarginal setigerous punctures. Each costal margin with 1124 setigerous punctures. Evaporative areas on meso- and metapleuron small, almost indistinct (
<figureCitation id="131C2A7FFFEBFFB548DCF8AD2DBD1A47" box="[439,511,1850,1876]" captionStart="FIGURES 9 12" captionStartId="4.[151,269,1645,1669]" captionTargetBox="[151,1434,346,1622]" captionTargetId="figure@4.[151,1436,346,1623]" captionTargetPageId="4" captionText="FIGURES 9 12. Evaporative areas on meso- and metapleuron. 9 — Byrsinus varians, 10 — Fromundus biimpressus, 11 — F. pygmaeus, 12 — Rhytidoporus indentatus." httpUri="https://zenodo.org/record/294107/files/figure.png" pageId="1" pageNumber="58">Fig. 9</figureCitation>
). Paramere described in
<bibRefCitation id="EFB64B0BFFEBFFB54A4AF8AD2FDD1A47" author="Lis" box="[801,927,1850,1876]" pageId="1" pageNumber="64" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae). Upper Silesian Museum, Bytom, 349 pp." type="book" year="1994">Lis (1994)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6571FFEBFFB549AEF8F72FFF1AE0" pageId="1" pageNumber="58" type="materials_examined">
<paragraph id="8B9836FAFFEBFFB549AEF8F72FFF1AE0" blockId="1.[151,1437,1661,2035]" pageId="1" pageNumber="58">
<emphasis id="B953EAE8FFEBFFB549AEF8F72DFD1A69" bold="true" box="[197,447,1888,1914]" pageId="1" pageNumber="58">Material examined:</emphasis>
<collectingCountry id="F330766AFFEBFFB548ACF8F72E561A69" box="[455,532,1888,1914]" name="Guam" pageId="1" pageNumber="58">Guam</collectingCountry>
: Mangilao Village, University of
<collectingCountry id="F330766AFFEBFFB54ACFF8F72FAE1A69" box="[932,1004,1888,1914]" name="Guam" pageId="1" pageNumber="58">Guam</collectingCountry>
campus, Marine Biol. Lab. Area, at lights,
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<geoCoordinate id="EE13503DFFEBFFB548E3F81F2E6F1AB1" box="[392,557,1925,1954]" direction="east" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="144.79855">E144o47.913</geoCoordinate>
,
<specimenCount id="9D21FD73FFEBFFB54B51F81F2EE01AB1" box="[570,674,1928,1954]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
<date id="FF99103AFFEBFFB54BC1F81F2F0E1AB1" box="[682,844,1928,1954]" pageId="1" pageNumber="58" value="2004-05-17">17 May 2004</date>
,
<specimenCount id="9D21FD73FFEBFFB54A31F81F2F801AB1" box="[858,962,1928,1954]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
<date id="FF99103AFFEBFFB54AA1F81F282E1AB1" box="[970,1132,1928,1954]" pageId="1" pageNumber="58" value="2004-05-18">18 May 2004</date>
,
<specimenCount id="9D21FD73FFEBFFB54D12F81F28AE1AB1" box="[1145,1260,1928,1954]" pageId="1" pageNumber="58" type="female">2 females</specimenCount>
<date id="FF99103AFFEBFFB54D9EF81F29D51AB1" box="[1269,1431,1928,1954]" pageId="1" pageNumber="58" value="2004-05-19">19 May 2004</date>
, R.S. Zack collr.;
<collectingCountry id="F330766AFFEBFFB54836F8262DEB1AD8" box="[349,425,1969,1995]" name="Guam" pageId="1" pageNumber="58">Guam</collectingCountry>
: Mangilao Village, Fadian Point, University of
<collectingCountry id="F330766AFFEBFFB54ABFF826285E1AD8" box="[980,1052,1969,1995]" name="Guam" pageId="1" pageNumber="58">Guam</collectingCountry>
aquaculture center,
<geoCoordinate id="EE13503DFFEBFFB54C6CF82629D91AD8" box="[1287,1435,1965,1995]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="1" value="13.4431">N13o26.586</geoCoordinate>
<geoCoordinate id="EE13503DFFEBFFB549FCF84E2D781AE0" box="[151,314,2006,2035]" direction="east" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="144.8196">E144o49.176</geoCoordinate>
,
<specimenCount id="9D21FD73FFEBFFB5482EF84E2DDD1AE0" box="[325,415,2009,2035]" pageId="1" pageNumber="58" type="male">5 males</specimenCount>
<specimenCount id="9D21FD73FFEBFFB548CCF84E2E671AE0" box="[423,549,2009,2035]" pageId="1" pageNumber="58" type="female">11 females</specimenCount>
<date id="FF99103AFFEBFFB54B46F84E2EF81AE0" box="[557,698,2009,2035]" pageId="1" pageNumber="58" value="2005-08-08">8 Aug 2005</date>
, R.S. Zack, MV light.
</paragraph>
</subSubSection>
<caption id="DF586672FFE8FFB649FCF8CF2CB81A83" httpUri="https://zenodo.org/record/294106/files/figure.png" pageId="2" pageNumber="59" targetBox="[151,1436,197,1848]" targetPageId="2">
<paragraph id="8B9836FAFFE8FFB649FCF8CF2CB81A83" blockId="2.[151,1436,1880,1936]" pageId="2" pageNumber="59">
<emphasis id="B953EAE8FFE8FFB649FCF8CF2D061A63" bold="true" box="[151,324,1880,1904]" pageId="2" pageNumber="59">FIGURES 18.</emphasis>
Head in two dorsal views. 1, 3, 5, 7—showing head shape and puncturation, 2, 4, 6, 8—showing head vestiture.
</paragraph>
</caption>
<subSubSection id="C33D6571FFE9FFB749AEFF002D931DEF" pageId="3" pageNumber="60" type="distribution">
<paragraph id="8B9836FAFFE9FFB749AEFF002D931DEF" blockId="3.[151,1437,151,588]" pageId="3" pageNumber="60">
<emphasis id="B953EAE8FFE9FFB749AEFF002D2D1DA2" bold="true" box="[197,367,151,177]" pageId="3" pageNumber="60">Distribution:</emphasis>
<collectingCountry id="F330766AFFE9FFB74813FF0F2E481DA1" box="[376,522,152,178]" name="Bangladesh" pageId="3" pageNumber="60">Bangladesh</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74B7CFF0F2E2C1DA1" box="[535,622,152,178]" name="Brunei" pageId="3" pageNumber="60">Brunei</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74B10FF0F2E931DA1" box="[635,721,152,178]" name="Myanmar" pageId="3" pageNumber="60">Burma</collectingCountry>
, Chagos Archipelago,
<collectingCountry id="F330766AFFE9FFB74A81FF0F28291DA1" box="[1002,1131,152,178]" name="Cambodia" pageId="3" pageNumber="60">Cambodia</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74D13FF0F28831DA1" box="[1144,1217,152,178]" name="China" pageId="3" pageNumber="60">China</collectingCountry>
(Hainan,
<collectingCountry id="F330766AFFE9FFB74C56FF0F29DA1DA1" box="[1341,1432,152,178]" name="China" pageId="3" pageNumber="60">Tianjin</collectingCountry>
, Neimenggu, Ningxia),
<collectingCountry id="F330766AFFE9FFB748D8FF2A2DB71DC4" box="[435,501,189,215]" name="India" pageId="3" pageNumber="60">India</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74B69FF2A2E3E1DC4" box="[514,636,189,215]" name="Indonesia" pageId="3" pageNumber="60">Indonesia</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74BE1FF2A2E911DC4" box="[650,723,189,215]" name="Japan" pageId="3" pageNumber="60">Japan</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74B8BFF2A2F6C1DC4" box="[736,814,189,215]" name="South Korea" pageId="3" pageNumber="60">Korea</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74A50FF2A2F381DC4" box="[827,890,189,215]" name="Laos" pageId="3" pageNumber="60">Laos</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74AECFF2A2FB91DC4" box="[903,1019,189,215]" name="Malaysia" pageId="3" pageNumber="60">Malaysia</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74D6CFF2A28921DC4" box="[1031,1232,189,215]" name="Maldives" pageId="3" pageNumber="60">Maldive Islands</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74DB5FF2A292E1DC4" box="[1246,1388,189,215]" name="Philippines" pageId="3" pageNumber="60">Philippines</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74C13FF2A2CA11DEF" name="Sri Lanka" pageId="3" pageNumber="60">Sri Lanka</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB74984FF752D181DEF" box="[239,346,226,252]" name="Thailand" pageId="3" pageNumber="60">Thailand</collectingCountry>
,
<collectingCountry id="F330766AFFE9FFB7480EFF752D8E1DEF" box="[357,460,226,252]" name="Vietnam" pageId="3" pageNumber="60">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6571FFE9FFB749AEFE902E0E1F5F" pageId="3" pageNumber="60" type="discussion">
<paragraph id="8B9836FAFFE9FFB749AEFE902E0E1F5F" blockId="3.[151,1437,151,588]" pageId="3" pageNumber="60">
<emphasis id="B953EAE8FFE9FFB749AEFE902D031C32" bold="true" box="[197,321,263,289]" pageId="3" pageNumber="60">Remarks:</emphasis>
When describing
<taxonomicName id="4C274D79FFE9FFB74B71FE9F2F561C32" box="[538,788,264,289]" class="Diplopoda" family="Dalodesmidae" genus="Microporus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="shiromai">
<emphasis id="B953EAE8FFE9FFB74B71FE9F2F561C32" box="[538,788,264,289]" italics="true" pageId="3" pageNumber="60">Microporus shiromai</emphasis>
</taxonomicName>
as a new species from Hawaii,
<bibRefCitation id="EFB64B0BFFE9FFB74DE6FE9F29241C31" author="Froeschner" box="[1165,1382,264,290]" pageId="3" pageNumber="63" refString="Froeschner, R. C. (1976) The burrowing bugs of Hawaii, with description of a new species (Hemiptera: Cydnidae)." type="book" year="1976">Froeschner (1976)</bibRefCitation>
was aware that this species was introduced into Hawaii from elsewhere; however, he decided to describe it as a new because, as he wrote, “this recently collected species needs a name, under which accumulating data can be stored.” Moreover, he stated that “its occurrence on the island represents a recent introduction,” and all the morphological characters “suggest an Old World origin of the species, but give no clue to a particular part thereof.” At present, it is clear that
<taxonomicName id="4C274D79FFE9FFB74B58FE552E831CC8" box="[563,705,450,476]" class="Diplopoda" family="Dalodesmidae" genus="Microporus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="shiromai">
<emphasis id="B953EAE8FFE9FFB74B58FE552E091CC8" box="[563,587,450,475]" italics="true" pageId="3" pageNumber="60">M</emphasis>
.
<emphasis id="B953EAE8FFE9FFB74B31FE552E831CC8" box="[602,705,450,475]" italics="true" pageId="3" pageNumber="60">shiromai</emphasis>
</taxonomicName>
has all of the characters of the Oriental
<taxonomicName id="4C274D79FFE9FFB74DFEFE55294C1CC8" box="[1173,1294,450,476]" class="Insecta" family="Cydnidae" genus="Byrsinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B953EAE8FFE9FFB74DFEFE5528E51CC8" box="[1173,1191,450,475]" italics="true" pageId="3" pageNumber="60">B</emphasis>
.
<emphasis id="B953EAE8FFE9FFB74DDDFE55294C1CC8" box="[1206,1294,450,475]" italics="true" pageId="3" pageNumber="60">varians</emphasis>
</taxonomicName>
, and should be regarded as its junior synonym. Unfortunately, the
<typeStatus id="549C8858FFE9FFB74A2DFE7F2F3E1F11" box="[838,892,488,514]" pageId="3" pageNumber="60">type</typeStatus>
specimens were not found in Froeschners collection (personal information from Dr. A. Wolski), and therefore the present synonymization is based only upon the original species description.
</paragraph>
</subSubSection>
</treatment>
</document>