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<document id="24AE45676CBC2B524CD15E1C55579EA0" ID-CLB-Dataset="21707" ID-DOI="10.11646/zootaxa.4790.2.10" ID-GBIF-Dataset="3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889408" ID-ZooBank="7F4A9B99-C12E-473B-B258-7BE5697F4986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859385935" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="038E87D4FF86FFA2FF157E35FB9F7761" docLanguage="en" docName="zootaxa.4790.2.10.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus lobissimus Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="365" masterDocId="FFB7FFACFF81FFA5FF827F44FFB87066" masterDocTitle="Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="374" masterPageNumber="358" pageNumber="365" updateTime="1698842324292" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B3DFDDA48D138C59499D9686BDA020C2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1F3F33D3E02D3FE2C1404E565CEFFA3E">
<mods:title id="783AF2F556CD26A494286303C7805912">Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="BAB7DC970613B08FE99BB3547F8BAB35" type="personal">
<mods:role id="F8AB9FB98126514118143E9279C2D1D4">
<mods:roleTerm id="346A793FA690DE4EAFF35524A2E7304E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4DCF322B4D533A6D034830E83A28D93D">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1D7E961DE72840D5708EF5EED13A6FF7">text</mods:typeOfResource>
<mods:relatedItem id="38C48AE6BE733C6BC98CCE7D2882F131" type="host">
<mods:titleInfo id="DFF18825EFCC567D6AC08F7EEAA379C1">
<mods:title id="48214C35A0315131FAFD14AC804B4EA9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C8CD51F13A4791EB56324E39D22CDDDF">
<mods:date id="74A7B906AE34F849D96FCBB6D4532B8B">2020</mods:date>
<mods:detail id="4C9008E2C8F83E7821A76A71236623A4" type="pubDate">
<mods:number id="8AA9747AB960A80251CBAFA6D180941A">2020-06-11</mods:number>
</mods:detail>
<mods:detail id="7AD10B68C1DE9460202C411CBD922812" type="volume">
<mods:number id="F51D9328E7F322DF3E57E12075A898D4">4790</mods:number>
</mods:detail>
<mods:detail id="D60A99702AFB75B629BF4CF2B1616D62" type="issue">
<mods:number id="BF72AEF26D704A4D16FC5F0EE7780DEB">2</mods:number>
</mods:detail>
<mods:extent id="04A49FF2C50E8D1A746CE0F05E46A493" unit="page">
<mods:start id="E307EE8F575F479874788A345C72412F">358</mods:start>
<mods:end id="1031C8A7419BC7322ADAC53BCC62F9FF">374</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="356631AB4BE5242D0B8AF3A7BA004F85">journal article</mods:classification>
<mods:identifier id="27D71EDD514D5FEB055017688FFDEDF2" type="CLB-Dataset">21707</mods:identifier>
<mods:identifier id="8C27D2B557292A14050CF725DE9802E1" type="DOI">10.11646/zootaxa.4790.2.10</mods:identifier>
<mods:identifier id="1AF7DA7A47A3FC796A2F0F2FC396C246" type="GBIF-Dataset">3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb</mods:identifier>
<mods:identifier id="EB92AF4B2492A5BCD808E25BBFF861C7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="00FD221650FC8B20DAD68872E589C601" type="Zenodo-Dep">3889408</mods:identifier>
<mods:identifier id="D4EB684C40DAE1FD0ABBAA00CB705947" type="ZooBank">7F4A9B99-C12E-473B-B258-7BE5697F4986</mods:identifier>
</mods:mods>
<treatment id="038E87D4FF86FFA2FF157E35FB9F7761" ID-DOI="http://doi.org/10.5281/zenodo.5918722" ID-GBIF-Taxon="164501158" ID-Zenodo-Dep="5918722" LSID="urn:lsid:plazi:treatment:038E87D4FF86FFA2FF157E35FB9F7761" httpUri="http://treatment.plazi.org/id/038E87D4FF86FFA2FF157E35FB9F7761" lastPageNumber="365" pageId="7" pageNumber="365">
<subSubSection id="C33D6549FF86FFA2FF157E35FDFA71ED" box="[151,578,369,396]" pageId="7" pageNumber="365" type="nomenclature">
<paragraph id="8B9836C2FF86FFA2FF157E35FDFA71ED" blockId="7.[151,578,369,431]" box="[151,578,369,396]" pageId="7" pageNumber="365">
<heading id="D0D081AEFF86FFA2FF157E35FDFA71ED" bold="true" box="[151,578,369,396]" fontSize="11" level="1" pageId="7" pageNumber="365" reason="1">
<emphasis id="B953EAD0FF86FFA2FF157E35FDFA71ED" bold="true" box="[151,578,369,396]" pageId="7" pageNumber="365">
<taxonomicName id="4C274D41FF86FFA2FF157E35FE1B71EA" ID-CoL="8Y4QM" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,419,369,396]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="lobissimus" status="sp. nov.">
<emphasis id="B953EAD0FF86FFA2FF157E35FE1B71EA" bold="true" box="[151,419,369,396]" italics="true" pageId="7" pageNumber="365">Hydrochus lobissimus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26057ABFF86FFA2FE307E35FDFA71ED" box="[434,578,369,395]" pageId="7" pageNumber="365" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF86FFA2FF157ED1FF4771C9" box="[151,255,404,431]" pageId="7" pageNumber="365" type="description">
<paragraph id="8B9836C2FF86FFA2FF157ED1FF4771C9" blockId="7.[151,578,369,431]" box="[151,255,404,431]" pageId="7" pageNumber="365">
<figureCitation id="131C2A47FF86FFA2FF157ED1FF5B71C9" box="[151,227,405,431]" captionStart="FIGURE 1" captionStartId="2.[151,250,2019,2044]" captionTargetBox="[192,1405,448,1963]" captionTargetId="figure@2.[151,1435,417,1993]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3889410" httpUri="https://zenodo.org/record/3889410/files/figure.png" pageId="7" pageNumber="365">Figs. 1</figureCitation>
,
<figureCitation id="131C2A47FF86FFA2FF737ED0FF4771C9" box="[241,255,404,431]" captionStart="FIGURE 5" captionStartId="8.[151,250,1762,1787]" captionTargetBox="[182,1429,196,1719]" captionTargetId="figure@8.[151,1435,181,1736]" captionTargetPageId="8" captionText="FIGURE 5. Hydrochus lobissimus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889418" httpUri="https://zenodo.org/record/3889418/files/figure.png" pageId="7" pageNumber="365">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF86FFA2FF157E99FD0D72A9" pageId="7" pageNumber="365" type="materials_examined">
<paragraph id="8B9836C2FF86FFA2FF157E99FD0D72A9" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">
<emphasis id="B953EAD0FF86FFA2FF157E99FEF27191" bold="true" box="[151,330,476,503]" pageId="7" pageNumber="365">Type Material.</emphasis>
<materialsCitation id="3B4F3C9FFF86FFA2FED07E99FEF5727D" ID-GBIF-Occurrence="2632886335" collectingDate="1977-06-11" collectionCode="NMNH" collectorName="P. J. Spangler &amp; D. R. Givens" country="Ecuador" location="Limoncocha" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
<typeStatus id="549C8860FF86FFA2FED07E99FE027191" box="[338,442,477,503]" pageId="7" pageNumber="365" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="F3307652FF86FFA2FDA77E99FD0C7191" box="[549,692,477,503]" name="Ecuador" pageId="7" pageNumber="365">ECUADOR</collectingCountry>
:
<collectingRegion id="49E3F820FF86FFA2FD427E99FD457191" box="[704,765,477,503]" country="Ecuador" name="Napo" pageId="7" pageNumber="365">Napo</collectingRegion>
<location id="8EF86019FF86FFA2FC877E99FC2C7191" LSID="urn:lsid:plazi:treatment:038E87D4FF86FFA2FF157E35FB9F7761:8EF86019FF86FFA2FC877E99FC2C7191" box="[773,916,477,503]" country="Ecuador" name="Limoncocha" pageId="7" pageNumber="365" stateProvince="Napo">Limoncocha</location>
,
<date id="FF991002FF86FFA2FC227E99FB827191" box="[928,1082,477,503]" pageId="7" pageNumber="365" value="1977-06-11">
<collectingDate id="EFDDE9EAFF86FFA2FC227E99FB827191" box="[928,1082,477,503]" pageId="7" pageNumber="365" value="1977-06-11">11 June 1977</collectingDate>
</date>
,
<collectorName id="26D25314FF86FFA2FBC57E99FB597191" box="[1095,1249,477,503]" pageId="7" pageNumber="365">P. J. Spangler</collectorName>
&amp;
<collectorName id="26D25314FF86FFA2FA857E99FA247191" box="[1287,1436,477,503]" pageId="7" pageNumber="365">D. R. Givens</collectorName>
#115” (
<collectionCode id="ED36AE07FF86FFA2FF6B7D45FEFD727D" box="[233,325,513,539]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3C9FFF86FFA2FED57D45FC2B727D" ID-GBIF-Occurrence="2632886341" box="[343,915,512,539]" collectingDate="1977-06-11" collectionCode="NMNH" collectorName="P. J. Spangler &amp; D. R. Givens" country="Ecuador" location="Same" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="B953EAD0FF86FFA2FED57D45FDB5727D" bold="true" box="[343,525,512,539]" pageId="7" pageNumber="365">
<typeStatus id="549C8860FF86FFA2FED57D45FE77727D" box="[343,463,513,539]" pageId="7" pageNumber="365" type="paratype">Paratypes</typeStatus>
(17):
</emphasis>
<location id="8EF86019FF86FFA2FD917D45FDE9727D" LSID="urn:lsid:plazi:treatment:038E87D4FF86FFA2FF157E35FB9F7761:8EF86019FF86FFA2FD917D45FDE9727D" box="[531,593,513,539]" name="Same" pageId="7" pageNumber="365">Same</location>
data as holotype (4
<collectionCode id="ED36AE07FF86FFA2FCB27D45FC33727D" box="[816,907,513,539]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF86FFA2FC1F7D45FC1B7259" ID-GBIF-Occurrence="2632886314" collectingDate="1999-09-05" collectingDateMax="1999-09-10" collectingDateMin="1999-09-05" collectionCode="TAMU" collectorName="E. G. Riley" country="Ecuador" elevation="215" latitude="-0.67444444" location="Estacion Cientifica Yasuni" longLatPrecision="21" longitude="-76.64722" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<collectingRegion id="49E3F820FF86FFA2FC1F7D45FC64727D" box="[925,988,513,539]" country="Ecuador" name="Napo" pageId="7" pageNumber="365">Napo</collectingRegion>
,
<location id="8EF86019FF86FFA2FC657D45FAA9727D" LSID="urn:lsid:plazi:treatment:038E87D4FF86FFA2FF157E35FB9F7761:8EF86019FF86FFA2FC657D45FAA9727D" box="[999,1297,513,539]" country="Ecuador" latitude="-0.67444444" longLatPrecision="21" longitude="-76.64722" name="Estacion Cientifica Yasuni" pageId="7" pageNumber="365" stateProvince="Napo">Estacion Cientifica Yasuni</location>
, elev.
<quantity id="4CDF9B27FF86FFA2FAD47D44FA24727D" box="[1366,1436,512,539]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="365" unit="m" value="215.0">
<elevation id="000AD1F1FF86FFA2FAD47D44FA24727D" box="[1366,1436,512,539]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="365" unit="m" value="215.0">215 m</elevation>
</quantity>
(
<geoCoordinate id="EE135005FF86FFA2FF227D61FEA57259" box="[160,285,549,575]" degrees="0" direction="south" minutes="40" orientation="latitude" pageId="7" pageNumber="365" precision="15" seconds="28" value="-0.67444444">0°40’28’’S</geoCoordinate>
,
<geoCoordinate id="EE135005FF86FFA2FEAB7D60FE787259" box="[297,448,548,575]" degrees="76" direction="west" minutes="38" orientation="longitude" pageId="7" pageNumber="365" precision="15" seconds="50" value="-76.64722">76°38’50’’W</geoCoordinate>
),
<date id="FF991002FF86FFA2FE567D60FDDD7259" box="[468,613,548,575]" pageId="7" pageNumber="365" value="1999-09-05" valueMax="1999-09-10" valueMin="1999-09-05">
<collectingDate id="EFDDE9EAFF86FFA2FE567D60FDDD7259" box="[468,613,548,575]" pageId="7" pageNumber="365" value="1999-09-05" valueMax="1999-09-10" valueMin="1999-09-05">5-10.ix.1999</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF86FFA2FD207D61FC987259" box="[674,800,549,575]" pageId="7" pageNumber="365">E. G. Riley</collectorName>
(3
<collectionCode id="ED36AE07FF86FFA2FCC77D61FC237259" box="[837,923,549,575]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="7" pageNumber="365" type="Herbarium">TAMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF86FFA2FC2F7D61FE507205" ID-GBIF-Occurrence="2632886311" collectingDate="1977-06-03" collectionCode="NMNH" collectorName="W. E. Steiner" country="Ecuador" latitude="-1.55" location="Limoncocha" longLatPrecision="1308" longitude="-75.583336" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<location id="8EF86019FF86FFA2FC2F7D61FB847259" LSID="urn:lsid:plazi:treatment:038E87D4FF86FFA2FF157E35FB9F7761:8EF86019FF86FFA2FC2F7D61FB847259" box="[941,1084,549,575]" country="Ecuador" latitude="-1.55" longLatPrecision="1308" longitude="-75.583336" name="Limoncocha" pageId="7" pageNumber="365" stateProvince="Napo">Limoncocha</location>
, (
<geoCoordinate id="EE135005FF86FFA2FBD37D61FB187259" box="[1105,1184,549,575]" degrees="1" direction="south" minutes="33" orientation="latitude" pageId="7" pageNumber="365" precision="925" value="-1.55">1°33’S</geoCoordinate>
,
<geoCoordinate id="EE135005FF86FFA2FB2F7D60FAAF7259" box="[1197,1303,548,575]" degrees="75" direction="west" minutes="35" orientation="longitude" pageId="7" pageNumber="365" precision="925" value="-75.583336">75°35’W</geoCoordinate>
),
<date id="FF991002FF86FFA2FAA97D61FA207259" box="[1323,1432,549,575]" pageId="7" pageNumber="365" value="1977-06-03">
<collectingDate id="EFDDE9EAFF86FFA2FAA97D61FA207259" box="[1323,1432,549,575]" pageId="7" pageNumber="365" value="1977-06-03">3.vi.1977</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF86FFA2FF4A7D0DFEE67205" box="[200,350,585,611]" pageId="7" pageNumber="365">W. E. Steiner</collectorName>
(1
<collectionCode id="ED36AE07FF86FFA2FE047D0DFE597205" box="[390,481,585,611]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF86FFA2FE767D0DFBD77205" ID-GBIF-Occurrence="2632886333" box="[500,1135,585,611]" collectingDate="1977-06-14" collectionCode="NMNH" collectorName="W. E. Steiner" country="Ecuador" latitude="-1.55" location="Limoncocha" longLatPrecision="1308" longitude="-75.583336" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
same locality,
<date id="FF991002FF86FFA2FD157D0DFCA97205" box="[663,785,585,611]" pageId="7" pageNumber="365" value="1977-06-14">
<collectingDate id="EFDDE9EAFF86FFA2FD157D0DFCA97205" box="[663,785,585,611]" pageId="7" pageNumber="365" value="1977-06-14">14.vi.1977</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF86FFA2FCCC7D0DFC5C7205" box="[846,996,585,611]" pageId="7" pageNumber="365">W. E. Steiner</collectorName>
(1
<collectionCode id="ED36AE07FF86FFA2FB8E7D0DFBDF7205" box="[1036,1127,585,611]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF86FFA2FBF97D0DFE5272E1" ID-GBIF-Occurrence="2632886317" collectingDate="1977-06-15" collectionCode="NMNH" collectorName="W. E. Steiner" country="Ecuador" latitude="-1.55" location="Limoncocha" longLatPrecision="1308" longitude="-75.583336" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
same locality,
<date id="FF991002FF86FFA2FA9F7D0CFA2F7205" box="[1309,1431,584,611]" pageId="7" pageNumber="365" value="1977-06-15">
<collectingDate id="EFDDE9EAFF86FFA2FA9F7D0CFA2F7205" box="[1309,1431,584,611]" pageId="7" pageNumber="365" value="1977-06-15">15.vi.1977</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF86FFA2FF4A7D29FEE772E1" box="[200,351,621,647]" pageId="7" pageNumber="365">W. E. Steiner</collectorName>
(1
<collectionCode id="ED36AE07FF86FFA2FE057D29FE5A72E1" box="[391,482,621,647]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF86FFA2FE747D29FEA072CD" ID-GBIF-Occurrence="2632886340" collectingDate="1974-04-10" collectionCode="NMNH" collectorName="Boyce A. Drummond" country="Ecuador" elevation="300" location="Rio" municipality="Limoncocha" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<collectingMunicipality id="6BFCACB8FF86FFA2FE747D29FD3D72E1" box="[502,645,621,647]" pageId="7" pageNumber="365">Limoncocha</collectingMunicipality>
, on
<location id="8EF86019FF86FFA2FD347D29FD6672E1" LSID="urn:lsid:plazi:treatment:038E87D4FF86FFA2FF157E35FB9F7761:8EF86019FF86FFA2FD347D29FD6672E1" box="[694,734,621,647]" country="Ecuador" municipality="Limoncocha" name="Rio" pageId="7" pageNumber="365" stateProvince="Napo">Rio</location>
<collectingRegion id="49E3F820FF86FFA2FD657D29FC9E72E1" box="[743,806,621,647]" country="Ecuador" name="Napo" pageId="7" pageNumber="365">Napo</collectingRegion>
, elev.
<quantity id="4CDF9B27FF86FFA2FCED7D29FC0272E1" box="[879,954,621,647]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="365" unit="m" value="300.0">
<elevation id="000AD1F1FF86FFA2FCED7D29FC0272E1" box="[879,954,621,647]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="365" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FF991002FF86FFA2FC457D29FB8772E1" box="[967,1087,621,647]" pageId="7" pageNumber="365" value="1974-04-10">
<collectingDate id="EFDDE9EAFF86FFA2FC457D29FB8772E1" box="[967,1087,621,647]" pageId="7" pageNumber="365" value="1974-04-10">10.iv.1974</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF86FFA2FBFC7D29FACC72E1" box="[1150,1396,621,647]" pageId="7" pageNumber="365">Boyce A. Drummond</collectorName>
, III (6
<collectionCode id="ED36AE07FF86FFA2FF347DD5FEA972CD" box="[182,273,657,683]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF86FFA2FEA17DD5FBA372CD" ID-GBIF-Occurrence="2632886328" box="[291,1051,657,683]" collectingDate="1974-04-23" collectionCode="NMNH" collectorName="Boyce A. Drummond" country="Ecuador" elevation="300" location="Rio" municipality="Limoncocha" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
same locality,
<date id="FF991002FF86FFA2FE467DD5FD8472CD" box="[452,572,657,683]" pageId="7" pageNumber="365" value="1974-04-23">
<collectingDate id="EFDDE9EAFF86FFA2FE467DD5FD8472CD" box="[452,572,657,683]" pageId="7" pageNumber="365" value="1974-04-23">23.iv.1974</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF86FFA2FDFA7DD5FCD472CD" box="[632,876,657,683]" pageId="7" pageNumber="365">Boyce A. Drummond</collectorName>
, III (1
<collectionCode id="ED36AE07FF86FFA2FC3A7DD5FBAB72CD" box="[952,1043,657,683]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF86FFA2FBA47DD5FD0972A9" ID-GBIF-Occurrence="2632886316" collectingDate="1969-03-25" collectingDateMax="1969-03-31" collectingDateMin="1969-03-25" collectionCode="NMNH" collectorName="P. &amp; P. Spangler" country="Ecuador" latitude="0.033333335" location="Santa Cecilia" longLatPrecision="1308" longitude="-76.96667" pageId="7" pageNumber="365" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<location id="8EF86019FF86FFA2FBA47DD5FB0572CD" LSID="urn:lsid:plazi:treatment:038E87D4FF86FFA2FF157E35FB9F7761:8EF86019FF86FFA2FBA47DD5FB0572CD" box="[1062,1213,657,683]" country="Ecuador" latitude="0.033333335" longLatPrecision="1308" longitude="-76.96667" name="Santa Cecilia" pageId="7" pageNumber="365" stateProvince="Napo">Santa Cecilia</location>
, (
<geoCoordinate id="EE135005FF86FFA2FB507DD5FAA172CD" box="[1234,1305,657,683]" degrees="0" direction="north" minutes="2" orientation="latitude" pageId="7" pageNumber="365" precision="925" value="0.033333335">0°2’N</geoCoordinate>
,
<geoCoordinate id="EE135005FF86FFA2FAA77DD4FA3772CD" box="[1317,1423,656,683]" degrees="76" direction="west" minutes="58" orientation="longitude" pageId="7" pageNumber="365" precision="925" value="-76.96667">76°58’W</geoCoordinate>
),
<date id="FF991002FF86FFA2FF157DF0FE8072A9" box="[151,312,692,719]" pageId="7" pageNumber="365" value="1969-03-25" valueMax="1969-03-31" valueMin="1969-03-25">
<collectingDate id="EFDDE9EAFF86FFA2FF157DF0FE8072A9" box="[151,312,692,719]" pageId="7" pageNumber="365" value="1969-03-25" valueMax="1969-03-31" valueMin="1969-03-25">25-31.iii.1969</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF86FFA2FEF67DF1FD9272A9" box="[372,554,693,719]" pageId="7" pageNumber="365">P. &amp; P. Spangler</collectorName>
(1
<collectionCode id="ED36AE07FF86FFA2FDCD7DF1FD1272A9" box="[591,682,693,719]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="365" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF86FFA2FF457D9CFE0073E2" pageId="7" pageNumber="365" type="diagnosis">
<paragraph id="8B9836C2FF86FFA2FF457D9CFE0073E2" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">
<emphasis id="B953EAD0FF86FFA2FF457D9CFE6A7295" bold="true" box="[199,466,728,755]" pageId="7" pageNumber="365">Differential Diagnosis.</emphasis>
Among known Ecuadorian species, differentiated from all by the deep cervical groove behind the eyes. This species is a member of the
<taxonomicName id="4C274D41FF86FFA2FD4A7DBAFCF97371" authorityName="Perkins" authorityYear="2019" box="[712,833,765,791]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="B953EAD0FF86FFA2FD4A7DBAFCF97371" box="[712,833,765,791]" italics="true" pageId="7" pageNumber="365">H. collaris</emphasis>
</taxonomicName>
group (
<emphasis id="B953EAD0FF86FFA2FC1E7DBAFC627371" box="[924,986,766,791]" italics="true" pageId="7" pageNumber="365">sensu</emphasis>
<bibRefCitation id="EFB64B33FF86FFA2FC617DB9FB307371" author="Perkins, P. D." box="[995,1160,765,791]" pageId="7" pageNumber="365" pagination="1 - 59" refId="ref7294" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins 2019c</bibRefCitation>
); differentiated from the three other species in the group (
<taxonomicName id="4C274D41FF86FFA2FD867C66FD61735D" authority="Perkins" authorityName="Perkins" authorityYear="2019" box="[516,729,801,827]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="B953EAD0FF86FFA2FD867C66FDC4735D" box="[516,636,801,827]" italics="true" pageId="7" pageNumber="365">H. collaris</emphasis>
Perkins
</taxonomicName>
,
<taxonomicName id="4C274D41FF86FFA2FD667C66FC64735D" authority="Perkins" authorityName="Perkins" authorityYear="2019" box="[740,988,801,827]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="conjunctus">
<emphasis id="B953EAD0FF86FFA2FD667C66FCC6735D" box="[740,894,802,827]" italics="true" pageId="7" pageNumber="365">H. conjunctus</emphasis>
Perkins
</taxonomicName>
, and
<taxonomicName id="4C274D41FF86FFA2FB947C66FABA735A" authority="Knisch" authorityName="Knisch" box="[1046,1282,801,828]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="purpureus">
<emphasis id="B953EAD0FF86FFA2FB947C66FB12735D" box="[1046,1194,802,827]" italics="true" pageId="7" pageNumber="365">H. purpureus</emphasis>
Knisch
</taxonomicName>
) by the larger and more closely spaced elytral punctures, the smaller body size (ca. 2.93 vs.
<quantity id="4CDF9B27FF86FFA2FC767C01FB2E7339" box="[1012,1174,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="3.565" metricValueMax="3.8" metricValueMin="3.33" pageId="7" pageNumber="365" unit="mm" value="3.565" valueMax="3.8" valueMin="3.33">3.33-3.80 mm</quantity>
), and the male genitalia (
<figureCitation id="131C2A47FF86FFA2FF227C2DFF5C73E5" box="[160,228,872,899]" captionStart="FIGURE 5" captionStartId="8.[151,250,1762,1787]" captionTargetBox="[182,1429,196,1719]" captionTargetId="figure@8.[151,1435,181,1736]" captionTargetPageId="8" captionText="FIGURE 5. Hydrochus lobissimus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889418" httpUri="https://zenodo.org/record/3889418/files/figure.png" pageId="7" pageNumber="365">Fig. 5</figureCitation>
; described below).
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF86FFA2FF457CC8FC767685" pageId="7" pageNumber="365" type="description">
<paragraph id="8B9836C2FF86FFA2FF457CC8FEC0743D" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">
<emphasis id="B953EAD0FF86FFA2FF457CC8FEE973C1" bold="true" box="[199,337,908,935]" pageId="7" pageNumber="365">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="131C2A47FF86FFA2FDDF7CC9FD1A73C1" box="[605,674,908,935]" captionStart="FIGURE 5" captionStartId="8.[151,250,1762,1787]" captionTargetBox="[182,1429,196,1719]" captionTargetId="figure@8.[151,1435,181,1736]" captionTargetPageId="8" captionText="FIGURE 5. Hydrochus lobissimus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889418" httpUri="https://zenodo.org/record/3889418/files/figure.png" pageId="7" pageNumber="365">Fig. 5</figureCitation>
). Size:
<typeStatus id="549C8860FF86FFA2FD757CC9FCE173C1" box="[759,857,909,935]" pageId="7" pageNumber="365" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.93/1.10; head width 0.71; pronotum l/w 0.71/0.63; PA 0.61; PB 0.54; elytra 1.83/1.10. Body size moderate (ca.
<quantity id="4CDF9B27FF86FFA2FF157C91FF477389" box="[151,255,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="2.93" pageId="7" pageNumber="365" unit="mm" value="2.93">2.93 mm</quantity>
). Dorsum grey to brown, with bronze or greenish, strong, iridescence, elytra with black spots. Punctation on elytra ca. 2-3x that of pronotum. Elytra interstriae ca. 0.5-1x strial puncture diameter. Interstria 9
<superScript id="7C529B8AFF86FFA2FB757CBCFABC7460" attach="left" box="[1271,1284,1016,1030]" fontSize="6" pageId="7" pageNumber="365">th</superScript>
more convex than others, overhanging 10
<superScript id="7C529B8AFF86FFA2FE507B58FE67744C" attach="left" box="[466,479,1052,1066]" fontSize="6" pageId="7" pageNumber="365">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C529B8AFF86FFA2FCF17B58FC38744C" attach="left" box="[883,896,1052,1066]" fontSize="6" pageId="7" pageNumber="365">th</superScript>
well raised, rather elongate, ca. eight punctures in row on each side.
</paragraph>
<paragraph id="8B9836C2FF86FFA2FF457B21FE6874C5" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">Head with deep curving cervical groove from side to side behind eyes, groove punctate, cervical area behind groove impunctate, shining.</paragraph>
<paragraph id="8B9836C2FF86FFA2FF457BE9FCD07569" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">Pronotum longer than wide (as ca. 71/63), widest at anterior margin, narrowed at base, sides very slightly arcuate, smooth; anterior margin markedly arcuate, fitting into cervical groove; depressions, especially anterior, deep, slightly more densely punctate than reliefs between depressions.</paragraph>
<paragraph id="8B9836C2FF86FFA2FF457A5DFAFA7531" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">Elytra with spaces between strial punctures ca. 0.5–1x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with distinct angulation, impunctate area of apices rather large.</paragraph>
<paragraph id="8B9836C2FF86FFA2FF457A25FF5E75F9" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae. Legs dark brown.</paragraph>
<paragraph id="8B9836C2FF86FFA2FF457AEDFABD7611" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">
Male genitalia (
<figureCitation id="131C2A47FF86FFA2FEF97AEDFE7875A5" box="[379,448,1448,1475]" captionStart="FIGURE 5" captionStartId="8.[151,250,1762,1787]" captionTargetBox="[182,1429,196,1719]" captionTargetId="figure@8.[151,1435,181,1736]" captionTargetPageId="8" captionText="FIGURE 5. Hydrochus lobissimus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889418" httpUri="https://zenodo.org/record/3889418/files/figure.png" pageId="7" pageNumber="365">Fig. 5</figureCitation>
) general characters: genitalia slender; parameres laterally expanded apically in dorsal/ ventral views, tips wider in dorsal/ventral views than lateral view, extending very slightly beyond distal end of aedeagus, each paramere with very distinct pwsa slightly before midlength; basal piece shorter than parameres, lateral margins parallel for most of length, orifice strongly sclerotized, in lateral view hooking toward dorsad; aedeagus distal to adbl narrow in dorsal/ventral views, slightly wider at midlength than at distal end of adbl, tapering apically, lateral margins narrowly sclerotized in distal 1/2, in lateral view with proximal ½ of avm emarginate.
</paragraph>
<paragraph id="8B9836C2FF86FFA2FF4579C5FD9476D9" blockId="7.[151,1437,476,1799]" pageId="7" pageNumber="365">Dorsal surface: adtl lacking; pdmm approximating alm; adbl very large, widest part of aedeagus, much wider than paramere area adjacent to adbl.</paragraph>
<paragraph id="8B9836C2FF86FFA2FF15798DFC767685" blockId="7.[151,1437,476,1799]" box="[151,974,1737,1763]" pageId="7" pageNumber="365">Ventral surface: pvmm basally very narrowly separated, overlapping alm.</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF86FFA2FF4579A9FB9F7761" box="[199,1063,1773,1799]" pageId="7" pageNumber="365" type="etymology">
<paragraph id="8B9836C2FF86FFA2FF4579A9FB9F7761" blockId="7.[151,1437,476,1799]" box="[199,1063,1773,1799]" pageId="7" pageNumber="365">
<emphasis id="B953EAD0FF86FFA2FF4579A9FEF67761" bold="true" box="[199,334,1773,1799]" pageId="7" pageNumber="365">Etymology.</emphasis>
Named in reference to the very large aedeagus dorsal basal lobe.
</paragraph>
</subSubSection>
</treatment>
</document>
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