1437 lines
173 KiB
XML
1437 lines
173 KiB
XML
<document id="7DF0872BF7724D7EA1F78F45176B4C84" ID-CLB-Dataset="299644" ID-DOI="10.3897/zse.100.117859" ID-GBIF-Dataset="f17d3dc6-151e-4b2c-bfe6-1e3978daa701" ID-ZooBank="15CA9ED4-C24C-4AD7-81AC-F699FA953FE8" ID-publisher-id="117859" URI-arpha="35315993-8E70-5007-A8AE-277D7EFC9889" URI-zoobank="http://zoobank.org/15CA9ED4-C24C-4AD7-81AC-F699FA953FE8" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1720542344403" checkinUser="pensoft" docAuthor="Yuan, Zi-Ming, Jiang, Wei & Sha, Zhong-Li" docDate="2024" docId="F5D4F37E18E35DD2B528EBEE01F48D06" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 965-987" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/117859/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Gothus Yuan, Jiang & Sha, 2024, gen. nov." docType="treatment" docUuid="240FAE30-235A-49A8-995D-6209A7F68991" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="987" masterDocId="353159938E705007A8AE277D7EFC9889" masterDocTitle="Morphological and molecular evidence for Gothus teemo gen. et sp. nov., a new xanthid crab (Crustacea, Brachyura, Xanthoidea) from coral reefs in the South China Sea, with a review of the taxonomy of Actaeodes consobrinus (A. Milne-Edwards, 1867)" masterLastPageNumber="987" masterPageNumber="965" pageNumber="965" updateTime="1720578290841" updateUser="ExternalLinkService">
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<mods:title id="E926294F0683AF33079E674BB3C433A9">Morphological and molecular evidence for Gothus teemo gen. et sp. nov., a new xanthid crab (Crustacea, Brachyura, Xanthoidea) from coral reefs in the South China Sea, with a review of the taxonomy of Actaeodes consobrinus (A. Milne-Edwards, 1867)</mods:title>
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<mods:name id="916866EEC577399998632024219CBB9A" type="personal">
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<mods:namePart id="1AC138AE3E17E25A0EE63B20D75235BA">Yuan, Zi-Ming</mods:namePart>
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<mods:nameIdentifier id="AD39E42801E4B026755C8C2B582A5803" type="ORCID">https://orcid.org/0009-0008-0009-2693</mods:nameIdentifier>
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<mods:affiliation id="A5A67CB1EB1D3F47DEFCAB4D2E0D4E7B">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China & Laoshan Laboratory, Qingdao 266237, China & Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China</mods:affiliation>
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<mods:name id="9FB4E189F0E6AC60873316C5DAB3C81F" type="personal">
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<mods:namePart id="1EC22490613EC7817A689DA218B4521A">Jiang, Wei</mods:namePart>
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<mods:affiliation id="6921B96604A6FA39B4459939AC33063D">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China & Laoshan Laboratory, Qingdao 266237, China</mods:affiliation>
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<mods:namePart id="A3167A2A1FF0FE777EE0A12BD7E39D8F">Sha, Zhong-Li</mods:namePart>
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<mods:affiliation id="8D0D5F1669DB0B1BCDA1C685DB4538D8">Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China & Laoshan Laboratory, Qingdao 266237, China</mods:affiliation>
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<mods:title id="B570B9E464FD1309E6B9197B0253841B">Zoosystematics and Evolution</mods:title>
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<mods:date id="23D3CD8B9EA30B9FDB6BA44E1BA88798">2024</mods:date>
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<mods:number id="9E38B7B9106FA78ECF968C02C5CE9A7C">2024-07-09</mods:number>
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<mods:identifier id="9D0A84AB82CFECDB06949DCC2A9675E9" type="DOI">10.3897/zse.100.117859</mods:identifier>
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<treatment id="F5D4F37E18E35DD2B528EBEE01F48D06" ID-DOI="http://doi.org/10.5281/zenodo.12702516" ID-GBIF-Taxon="235025287" ID-Zenodo-Dep="12702516" ID-arpha="F5D4F37E-18E3-5DD2-B528-EBEE01F48D06" ID-zoobank="https://zoobank.org/240FAE30-235A-49A8-995D-6209A7F68991" LSID="urn:lsid:zoobank.org:act:240FAE30-235A-49A8-995D-6209A7F68991" LSID-ZBK="urn:lsid:zoobank.org:act:240FAE30-235A-49A8-995D-6209A7F68991" httpUri="http://treatment.plazi.org/id/F5D4F37E18E35DD2B528EBEE01F48D06">
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<subSubSection id="574DE33A89DE53A13967B290848C3F15" type="nomenclature">
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<paragraph id="DAC47542813D4A5E6B4C5712244EB14D">
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<taxonomicName id="173843FE1407794B89618CBB73A701F8" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="0C2BBCBE3F544A90563E8499BFD90749" italics="true">Gothus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AD28C097456ED87F77C9E16B3943D2CC" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="DA319560A1ACE2AB24F3FC7936528A8F" type="description">
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<paragraph id="9A3EF2F94F619766941603E323C65C71">
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<figureCitation id="DFDBEFC76082B13E7CD2736F9D1319C9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="B15C7092977D3997864954DBD0C8C8DE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A – F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2</figureCitation>
|
||
,
|
||
<figureCitation id="45C53F3C5F5C6EE6614D8A47C03D2D5F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3</figureCitation>
|
||
,
|
||
<figureCitation id="6EC45F83011B6335A34FF63DAE2A62CE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Gothus teemo sp. nov. A, B. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; C – E. Paratypes, 2 juvenile, CW 1.8–2.2 mm, CL 1.3–1.5 mm, MBM 287023; A – D. Live coloration; E. Habitat and substrate conditions. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure4" httpUri="https://binary.pensoft.net/fig/1088097">4</figureCitation>
|
||
,
|
||
<figureCitation id="9AA3013693F3163D61C7528EBD8FBC65" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Gothus teemo sp. nov., paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026, artistic illustration, displaying live coloration. Drawn by Fei Gao." figureDoi="10.3897/zse.100.117859.figure5" httpUri="https://binary.pensoft.net/fig/1088098">5</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1EAI" type="type_taxon">
|
||
<paragraph id="0744592335AD1CBBDE67C8A942729C47">
|
||
<heading id="F3A5E2E4B824CCCCA2FCCEC35570E739" reason="title">
|
||
<typeStatus id="96346893BF9CAC2A8C6DFFA895E6DAB8">Type</typeStatus>
|
||
species.
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C15C68BE7D632AE1C78F5E93529DF0D8">
|
||
<taxonomicName id="007AA985F1D62B4FEDB02EF32E08F7BA" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
|
||
<emphasis id="2CF99AE5FCB6B84510504414690FDF2F" italics="true">Gothus teemo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="279D01D551D14571D99D6620DF0E48B8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, by present designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELFAI" type="diagnosis">
|
||
<paragraph id="6A968E576BCB2E11404966AABAFD0C5B">
|
||
<heading id="BA99306BDF34E96EA392FF4D3FE8803A" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="2B2789B02C96E2B94EF75EB456F93067">
|
||
Small species,
|
||
<abbrev id="ABBRID0ERFAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
under
|
||
<quantity id="970B740E837918400FC4ABEC2FBB7E1B" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
|
||
. Carapace broader than long, dorsal surface bearing round granules, regions clearly defined; front wide, not protruding, divided into two slightly triangular lobes by a V-shaped notch; frontal lobes and dorsal inner orbital angle separated by shallow depression; eyestalks densely granulated; area beneath outer orbital angle slightly concave, not forming a subhepatic cavity; anterolateral margin with four teeth, first tooth flattened, sometimes completely reduced to appear as three teeth; posterolateral margin almost straight; subhepatic region densely granulated.
|
||
</paragraph>
|
||
<paragraph id="7A099F8D21D44F5B3D77BA8FC846136B">Epistome central region with low median projection on posterior margin. Maxilliped 3 granulated, anterior edge of merus indented, external terminal angle expanded. Antennule folding transversely; basal segment of antenna subrectangular; contacting ventral external frontal margin and ventral internal orbital angle; antennal flagellum filling orbital hiatus.</paragraph>
|
||
<paragraph id="F2E2F776E2718CDD97848E8B29F51B20">Chelipeds symmetrical, merus short; carpi robust, surface granulated, aggregated into nodules; outer and dorsal surfaces of palm densely granulated; fingers elongated, with triangular teeth; tips sharp, crossing at extremities when closed; dorsal surface of movable finger with three granulated ridges. Fingers brownish-black, coloration of immovable finger extending onto inner and outer surfaces of palm in male.</paragraph>
|
||
<paragraph id="7F331BA26B5D4DB53C4A4F3DBC4359C7">Ambulatory legs with meri flattened, granulated along anterior and posterior edges; dactyli elongated, margins with granules and setae, terminal end chitinous, sharp, slightly curved backward, dactylo-propodal lock present but underdeveloped.</paragraph>
|
||
<paragraph id="48807F64B090F93B52797C5DEAE9EDED">Male thoracic sternum with sternites 1 and 2 completely fused, suture between sternites 2 and 3 straight, complete, sternites 3 and 4 mostly fused, suture between them visible only at margins, sternites 3 short, sternite 4 with central longitudinal groove, tubercle of sterno-pleonal lock located on posterior of sternite 5. Male pleon narrow, pleonites 3 to 5 completely fused, lateral margins of pleonite 6 slightly concave. Telson long, broad, truncated oval, base margin wider than terminal margin of pleonite 6.</paragraph>
|
||
<paragraph id="5D33D3CC146A982C447BE7A9BA045BD0">
|
||
<abbrev id="ABBRID0E2FAI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
slender, curving slightly outward, distal lobe spoon-shaped, long setae on inner subdistal part, small spines on outer part.
|
||
<abbrev id="ABBRID0E6FAI" xlink_title="second gonopod of male">G 2</abbrev>
|
||
not exceeding 1 / 6 length of
|
||
<abbrev id="ABBRID0EDGAI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
, distal lobe elongated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EHGAI" type="etymology">
|
||
<paragraph id="44EC47EBE6DF5424321AA5AE557C75F7">
|
||
<heading id="1BAEE6C81501634510CF0417E17C66B6" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="98FABA3D7C8B659ACE5F0EFE1DAF8A26">
|
||
The genus is named after the game of Go, alluding to the intermingled black and white patterns on the carapace, beneath which lie circular granules resembling the pieces of the game. “ - thus ” is a common suffix for species names within the
|
||
<taxonomicName id="A7C8717B67C68271370507A7E371ACD8" authorityName="MacLeay" authorityYear="1838" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="family">Xanthidae</taxonomicName>
|
||
family. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ESGAI" type="material">
|
||
<paragraph id="9A8021B9FBD48F7C7E2E94F5D059F060">
|
||
<heading id="E88D982E4AA1DA69997F6A756C8E3FC5" reason="title">Comparative material.</heading>
|
||
</paragraph>
|
||
<paragraph id="475B4A1AE962DBC9927EBA0522AF4A93">
|
||
<taxonomicName id="252D2CBE02ACC5801A88EAA598332D7F" authority="Mendoza & PKL Ng, 2008" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="anconis">
|
||
<emphasis id="4DDDD16C38BF8386137F1EAFEE290BAE" bold="true">
|
||
<emphasis id="28034A0D01D17C6F4B962F056D5EF57A" bold="true" italics="true">Rizalthus anconis</emphasis>
|
||
<bibRefCitation id="8FF25CD512FC68BA66693400EBA65300" author="Mendoza JCE & Ng PKL" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385 - 404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza & PKL Ng, 2008</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="3DE5839F5D07FCB5F341F5E363010FB0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 A</figureCitation>
|
||
).
|
||
<materialsCitation id="A5C18F4B5590928A14CC84D96CB43DA6" ID-GBIF-Occurrence="4904612333" collectingDate="2022-05-07" collectorName="Ziming Yuan & Yuli Sun & Shaobo Ma" country="China" elevation="8" latitude="9.877275" location="Meiji Reef" longLatPrecision="1" longitude="115.521416" municipality="Nansha Islands" specimenCount="1" specimenCount-female="1">
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||
<collectingCountry id="A9250961622D5C788C8B9E2C7D45D705" name="China">China</collectingCountry>
|
||
•
|
||
<specimenCount id="006C286F5EA610EFFB9498B17C6A0E2F" count="1" type="female">1 female</specimenCount>
|
||
;
|
||
<abbrev id="ABBRID0EKHAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
<quantity id="2C6C8C021D1068CBC753BA2E34275344" metricMagnitude="-3" metricUnit="m" metricValue="4.2" unit="mm" value="4.2">4.2 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EOHAI" xlink_title="median carapace length">CL</abbrev>
|
||
<quantity id="43B1827420EEAA9013B30C58093ABA68" metricMagnitude="-3" metricUnit="m" metricValue="2.7" unit="mm" value="2.7">2.7 mm</quantity>
|
||
;
|
||
<location id="7410E8C2C500E21B6E5533F083414BF3" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:7410E8C2C500E21B6E5533F083414BF3" country="China" latitude="9.877275" longLatPrecision="1" longitude="115.521416" municipality="Nansha Islands" name="Meiji Reef">Meiji Reef</location>
|
||
,
|
||
<collectingMunicipality id="A9F80452586667FEAC7578C0AD4C6662">Nansha Islands</collectingMunicipality>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVHAI" specific-use="{"type":"Point","coordinates":[115.521411,9.877275]}">
|
||
<geoCoordinate id="A49A16EEF14D91330EE1B7B226670ECF" degrees="9" direction="north" minutes="52" orientation="latitude" precision="1" seconds="38.19" value="9.877275">9 ° 52 ' 38.19 " N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B25505A679164A8BD3761F6FB5E4E74F" degrees="115" direction="east" minutes="31" orientation="longitude" precision="1" seconds="17.08" value="115.521416">115 ° 31 ' 17.08 " E</geoCoordinate>
|
||
</named-content>
|
||
;
|
||
<quantity id="C29CA37EA00D030CC592DBBB99113962" metricMagnitude="0" metricUnit="m" metricValue="8.0" unit="m" value="8.0">
|
||
<elevation id="4363E4460F4578BEF888604D5409E0FE" metricMagnitude="0" metricUnit="m" metricValue="8.0" unit="m" value="8.0">8 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="5972B5E3ECDCEE2C3B90317565069D70" value="2022-05-07">
|
||
<collectingDate id="599DA3983B494A5E350191509BD8FFC6" value="2022-05-07">7 May 2022</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="EEC26BE3D7080F41D9683991F7649EF2">Ziming Yuan</collectorName>
|
||
,
|
||
<collectorName id="B0DCF734493A330AEB9063C9D9FD38CC">Yuli Sun</collectorName>
|
||
,
|
||
<collectorName id="D4AD51BC32B7ABBFBCD5CF6025F143B9">Shaobo Ma</collectorName>
|
||
coll.;
|
||
<specimenCode id="4FE22C923097EC1553EE350DAA535D61">NS-MJ- 2022-1457</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="ABB7DD8E8ABFE2139D4454ED5155900C">
|
||
<taxonomicName id="70F427005986580BAE262FF698DF4F2F" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="haanii">
|
||
<emphasis id="A999FB7F83FF8FEBF19A379C6EF970B0" bold="true">
|
||
<emphasis id="8018C62CC48A3397D19C525E5FE6668B" bold="true" italics="true">Hypocolpus haanii</emphasis>
|
||
<bibRefCitation id="1D8E3DAFDCCC1953129B3B402D76106E" author="Rathbun MJ" journalOrPublisher="Proceedings of the Biological Society of Washington" pagination="107-114" refId="B37" refString="Rathbun MJ (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington 22: 107 - 114." title="New crabs from the Gulf of Siam." volume="22" year="1909">Rathbun, 1909</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="ACB7A56BD084480EE16A21705F8711A5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 B</figureCitation>
|
||
).
|
||
<materialsCitation id="95D7403551F3ABB51375949785DEF08C" ID-GBIF-Occurrence="4904612370" collectingDate="2018-08-20" collectionCode="MBM" collectorName="Yunhao Pan" country="China" elevation="23" location="Island" municipality="Lingao Cape" specimenCode="MBM 286755" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
|
||
<collectingCountry id="9A6DDAC0AE1F0B11C4F6BFEAAF138A42" name="China">China</collectingCountry>
|
||
•
|
||
<specimenCount id="0ECF9D07476895171F2E403C52FB9388" count="1" type="male">1 male</specimenCount>
|
||
;
|
||
<abbrev id="ABBRID0EOIAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
<quantity id="E9E2C55B36ADADD9083F9E8D01A2D6DB" metricMagnitude="-2" metricUnit="m" metricValue="4.529999999999999" unit="mm" value="45.3">45.3 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0ESIAI" xlink_title="median carapace length">CL</abbrev>
|
||
<quantity id="0E443630B37368F4A7B46DC29513AD8F" metricMagnitude="-2" metricUnit="m" metricValue="3.42" unit="mm" value="34.2">34.2 mm</quantity>
|
||
;
|
||
<collectingMunicipality id="1F33211C8D088BC74DBCF31C02B54E78">Lingao Cape</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="819A7D8F96EF8B9780A4323768ECC70E" country="China" name="Hainan">Hainan</collectingRegion>
|
||
<location id="F98152DE6AE523B8904E58538292869E" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:F98152DE6AE523B8904E58538292869E" country="China" municipality="Lingao Cape" name="Island" stateProvince="Hainan">Island</location>
|
||
;
|
||
<quantity id="899E91DCFED1153E63EDD8BBA2B0D239" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" unit="m" value="22.5" valueMax="30.0" valueMin="15.0">
|
||
<elevation id="40D7AE1E8EF7D31A948C96088A547F9E" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" unit="m" value="22.5" valueMax="30.0" valueMin="15.0">15–30 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="E59886B737687E77797305E701ACE1D8" value="2018-08-20">
|
||
<collectingDate id="9262E796BD2A13C20461ECA69A096D29" value="2018-08-20">20 Aug. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="A2AC025783B088D65F364897596B466C">Yunhao Pan</collectorName>
|
||
coll.;
|
||
<specimenCode id="79D12B6E3EC1AA71F70A05DE65C01DE8" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 286755</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A234EF0E7A4BBF55F0829405FC511F27">
|
||
<taxonomicName id="75D82F2CF1849AAE29B509E77A095608" authority="(Herbst, 1790)" baseAuthorityName="Herbst" baseAuthorityYear="1790" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="exsculptus">
|
||
<emphasis id="67F2FCFAB880868295B74D9F67D17EDA" bold="true">
|
||
<emphasis id="EE0D677B767B05FE2DB4E57D6636E88F" bold="true" italics="true">Euxanthus exsculptus</emphasis>
|
||
(Herbst, 1790)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="B89E0020088539BCF20F7CE66717A782" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 C</figureCitation>
|
||
).
|
||
<materialsCitation id="8518A649E25C27A23F8314FB3496FD9B" ID-GBIF-Occurrence="4904612337" collectingDate="2018-12-20" collectorName="Yunhao Pan & Xan" country="China" location="Island" municipality="Wenchang" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
|
||
<collectingCountry id="305494C1B94B7BA311D9377246DED5B8" name="China">China</collectingCountry>
|
||
•
|
||
<specimenCount id="45A80B62CB9F9033B405DB29A51271F9" count="1" type="male">1 male</specimenCount>
|
||
;
|
||
<collectingMunicipality id="14A9FF621970AD7E8266CAA2B9F33EEA">Wenchang</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="2E6FE24A9B704F3E9D97652B4295702A" country="China" name="Hainan">Hainan</collectingRegion>
|
||
<location id="D89B9D48A8927FFCEBD3C5F176551534" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:D89B9D48A8927FFCEBD3C5F176551534" country="China" municipality="Wenchang" name="Island" stateProvince="Hainan">Island</location>
|
||
;
|
||
<date id="382CDF52E0C0B74CF2EAC920F999454B" value="2018-12-20">
|
||
<collectingDate id="14964793435E590ABA9421E8DB7F95D6" value="2018-12-20">20 Dec. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="16A37CD008E6C315B233424CB5974743">Yunhao Pan</collectorName>
|
||
coll.; Xan 016
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="12983F67DF0CABA11071E6FBF94629EF" ID-GBIF-Occurrence="4904612336" collectingDate="1957-05-15" collectingDateMax="1957-05-17" collectingDateMin="1957-05-15" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Yongxing Island" specimenCode="MBM 163793" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
|
||
<specimenCount id="E92AB0F11390D4EDBAA614267480959C" count="1" type="male">1 male</specimenCount>
|
||
;
|
||
<collectingMunicipality id="AE2EFB1A5E81BF4359DDC22BBA9CB82B">Yongxing Island</collectingMunicipality>
|
||
,
|
||
<location id="305776A1A1DDF9DCF0EBABE20751F50F" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:305776A1A1DDF9DCF0EBABE20751F50F" country="China" municipality="Yongxing Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
|
||
;
|
||
<date id="21E22FB76A22F1673F9652A74A343D7D" value="1957-05-15" valueMax="1957-05-17" valueMin="1957-05-15">
|
||
<collectingDate id="4AC6A74F53CCA29C673E531D4D442CCD" value="1957-05-15" valueMax="1957-05-17" valueMin="1957-05-15">15–17 May 1957</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="BCBCA1D7AC0DC336DC4505A6DE78D5CA" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163793</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="DF780EDD5C4B5849D36F5DA76BCF67CE" ID-GBIF-Occurrence="4904612311" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Wood Island" specimenCode="MBM 163791" specimenCount="2" specimenCount-male="2" stateProvince="Hainan">
|
||
<specimenCount id="4EA082789A694640DB84E736A4330898" count="2" type="male">2 males</specimenCount>
|
||
;
|
||
<collectingMunicipality id="66FA3837D51B98D7910F6114C766977D">Wood Island</collectingMunicipality>
|
||
,
|
||
<location id="B58C5907FE38A99FC92554F7FA01DFF2" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:B58C5907FE38A99FC92554F7FA01DFF2" country="China" municipality="Wood Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
|
||
; 1957;
|
||
<specimenCode id="533BC253A89C201185DF101A75CB99F8" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163791</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3795DF3D0082632E6A8500D890972E48" ID-GBIF-Occurrence="4904612352" collectingDate="1957-05-15" collectingDateMax="1957-05-17" collectingDateMin="1957-05-15" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Wood Island" specimenCode="MBM 163788" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hainan">
|
||
<specimenCount id="0C1FCBC424DB8254ED5F336111381157" count="1" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="B15B6932815B5D683B79A98D0B41CD44" count="2" type="female">2 females</specimenCount>
|
||
;
|
||
<collectingMunicipality id="A7DB2FCE85B0810F8B43DB3AD3DE2D6B">Wood Island</collectingMunicipality>
|
||
,
|
||
<location id="F0A8CB032C444FA520C41D801857F85A" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:F0A8CB032C444FA520C41D801857F85A" country="China" municipality="Wood Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
|
||
;
|
||
<date id="61C1A7C2FE88DA9E15E4594817E843CA" value="1957-05-15" valueMax="1957-05-17" valueMin="1957-05-15">
|
||
<collectingDate id="F9C4F1912A38D5A8F93F508E9715E103" value="1957-05-15" valueMax="1957-05-17" valueMin="1957-05-15">15–17 May 1957</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="3D7C0CD59ABCE9544BC56E665CB224BB" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163788</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="5F3BC2A6C3CA7AA01BB97E069098FD0E" ID-GBIF-Occurrence="4904612347" collectingDate="1980-05-28" collectingDateMax="1980-05-31" collectingDateMin="1980-05-28" collectionCode="MBM" country="China" location="Xisha Islands" municipality="East Island" specimenCode="MBM 163785" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hainan">
|
||
<specimenCount id="92041B86DD1200C77E6F6C7A5C6F931B" count="3" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="FE93E01E4820635AEFAA1329E8D6B1C4" count="3" type="female">3 females</specimenCount>
|
||
;
|
||
<collectingMunicipality id="222A9E1C5BF257ED35156A3952A09F8F">East Island</collectingMunicipality>
|
||
,
|
||
<location id="F4067815E0D5F4A33B3410EF7CA2FA39" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:F4067815E0D5F4A33B3410EF7CA2FA39" country="China" municipality="East Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
|
||
;
|
||
<date id="7A155FCC7D3824EA2DE4DFB30A2DA577" value="1980-05-28" valueMax="1980-05-31" valueMin="1980-05-28">
|
||
<collectingDate id="FC4ADE9BE3CC5DFEA1E9023633C22545" value="1980-05-28" valueMax="1980-05-31" valueMin="1980-05-28">28–31 May 1980</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="C79EF35F7E31DFF5C87F364B16D7FCC5" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163785</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="62868AE5C91B856261B6BCE977983FCB" ID-GBIF-Occurrence="4904612363" collectingDate="1980-06-11" collectingDateMax="1980-06-13" collectingDateMin="1980-06-11" collectionCode="MBM" country="China" location="Xisha Islands" municipality="Yongxing Island" specimenCode="MBM 163784" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
|
||
<specimenCount id="F48EDCA92A6DAB1A508A0AA5E6CCABE4" count="1" type="female">1 female</specimenCount>
|
||
;
|
||
<collectingMunicipality id="86A11DDE0CF6FA79EAAEEA21049093F6">Yongxing Island</collectingMunicipality>
|
||
,
|
||
<location id="1935B7D381C3C3896A9366F7FC0336A4" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:1935B7D381C3C3896A9366F7FC0336A4" country="China" municipality="Yongxing Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
|
||
;
|
||
<date id="4E49E2E83A38F4949E95E6E12F7A9BC2" value="1980-06-11" valueMax="1980-06-13" valueMin="1980-06-11">
|
||
<collectingDate id="208847363FD0C04935E4364B209EDE22" value="1980-06-11" valueMax="1980-06-13" valueMin="1980-06-11">11–13 Jun. 1980</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="D84C329D5BE16C01D2E851B24403AA4D" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163784</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="152B7670028C7D37B0BE736BDFC1C6AF" ID-GBIF-Occurrence="4904612349" collectingDate="1975-06-12" collectionCode="MBM" collectorName="Xianqiu Ren" country="China" location="Xisha Islands" municipality="East Island" specimenCode="MBM 163792" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
|
||
<specimenCount id="266D01136283AE6E912154FE859E03F3" count="1" type="female">1 female</specimenCount>
|
||
;
|
||
<collectingMunicipality id="37E0BD100627920841E42B52D70617B2">East Island</collectingMunicipality>
|
||
,
|
||
<location id="484DD05E24E0F20625EF959A4084A5C8" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:484DD05E24E0F20625EF959A4084A5C8" country="China" municipality="East Island" name="Xisha Islands" stateProvince="Hainan">Xisha Islands</location>
|
||
;
|
||
<date id="5C66009BB8C7008C52E7416BFC32C994" value="1975-06-12">
|
||
<collectingDate id="CFDD1CF0E3FEB00BB088F89F629CD2C5" value="1975-06-12">12 Jun. 1975</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="135211B13733F04B9B1D95B9E5D86EE0">Xianqiu Ren</collectorName>
|
||
coll.;
|
||
<specimenCode id="A893DC9EFC3CA9FEB63396B0DC27E9DB" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163792</specimenCode>
|
||
.
|
||
<abbrev id="ABBRID0EKJAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
<quantity id="BD83F5A751126ACA92719FB363668A3E" metricMagnitude="-2" metricUnit="m" metricValue="3.3899999999999997" metricValueMax="5.279999999999999" metricValueMin="1.5" unit="mm" value="33.9" valueMax="52.8" valueMin="15.0">15–52.8 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EOJAI" xlink_title="median carapace length">CL</abbrev>
|
||
<quantity id="410FD656B541655C9025349AAB884261" metricMagnitude="-2" metricUnit="m" metricValue="2.145" metricValueMax="3.32" metricValueMin="0.97" unit="mm" value="21.45" valueMax="33.2" valueMin="9.7">9.7–33.2 mm</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EE63A2E13E4CB3953D18C5796127724D">
|
||
<taxonomicName id="E4B4F28CE8DE8CC651EED8022E3F3E09" authority="(Hombron & Jacquinot, 1846)" baseAuthorityName="Hombron & Jacquinot" baseAuthorityYear="1846" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="huonii">
|
||
<emphasis id="D0B953E8C2DBF3023C48D80D6EFE426B" bold="true">
|
||
<emphasis id="F31BA3892247BA2E44F3B62D9E6044B1" bold="true" italics="true">Euxanthus huonii</emphasis>
|
||
(Hombron & Jacquinot, 1846)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="E5A22BC9DCCD9A2FC29377E918965171" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 D</figureCitation>
|
||
).
|
||
<materialsCitation id="6B61DD6531D38F9AA4EB00DFCFEB79CA" ID-GBIF-Occurrence="4904612326" collectingDate="1958-04-11" collectingDateMax="1958-04-17" collectingDateMin="1958-04-11" collectionCode="MBM" country="China" location="Dengqing Island" municipality="Xisha Islands" specimenCode="MBM 163762" specimenCount="1" specimenCount-male="1">
|
||
<collectingCountry id="8DA774D5C3F63C208FE2A9D6E4348AAA" name="China">China</collectingCountry>
|
||
•
|
||
<specimenCount id="12078B0C67AF35B1F0D2C75FA2837888" count="1" type="male">1 male</specimenCount>
|
||
;
|
||
<location id="59103009792DFCB99C4BA6B31A1D1CBA" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:59103009792DFCB99C4BA6B31A1D1CBA" country="China" municipality="Xisha Islands" name="Dengqing Island">Dengqing Island</location>
|
||
,
|
||
<collectingMunicipality id="30AE7F4F694C5EB11789D9ECD71FB486">Xisha Islands</collectingMunicipality>
|
||
;
|
||
<date id="4F0040B555CDF737C1C5843F9F8B0A6C" value="1958-04-11" valueMax="1958-04-17" valueMin="1958-04-11">
|
||
<collectingDate id="B4B04A10DFD4B58E6BE1610BB48DFB48" value="1958-04-11" valueMax="1958-04-17" valueMin="1958-04-11">11–17 Apr. 1958</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="65832BABB4AD106D14DCC7D7026A8A67" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163762</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3DB625462B925E61E99347877AC52484" ID-GBIF-Occurrence="4904612360" collectingDate="1958-05-01" collectionCode="MBM" country="China" location="Tree Island" municipality="Xisha Islands" specimenCode="MBM 163761" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="6BDCE19A4638AC351080F40389D4DDA0" count="1" type="female">1 female</specimenCount>
|
||
;
|
||
<location id="1E933B1D844C5584DDD5E73B929BE61F" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:1E933B1D844C5584DDD5E73B929BE61F" country="China" municipality="Xisha Islands" name="Tree Island">Tree Island</location>
|
||
,
|
||
<collectingMunicipality id="306C23C0A5BBC9DE322F04F8C031CDB5">Xisha Islands</collectingMunicipality>
|
||
;
|
||
<date id="24A65E338D6271F267BDBFF2291C9AB2" value="1958-05-01">
|
||
<collectingDate id="10681EDA5B9CCABDDF250297262419AF" value="1958-05-01">1 May 1958</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="F227F0C17DA7EACE4476C62261839255" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163761</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="E3C310A158605607B9B4A57BBD170DE2" ID-GBIF-Occurrence="4904612307" collectingDate="1980-05-07" collectionCode="MBM" country="China" location="Yongxing Island" municipality="Xisha Islands" specimenCode="MBM 163780" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9453448AED64511E535C3FAE41BFC102" count="1" type="female">1 female</specimenCount>
|
||
;
|
||
<location id="958F2A15163754B40C9CCEDA1687F0FF" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:958F2A15163754B40C9CCEDA1687F0FF" country="China" municipality="Xisha Islands" name="Yongxing Island">Yongxing Island</location>
|
||
,
|
||
<collectingMunicipality id="6731C40EFC07A192EC74EF784DCC115F">Xisha Islands</collectingMunicipality>
|
||
;
|
||
<date id="A356A477FAED1DDCC9A65BBD5D9B9A38" value="1980-05-07">
|
||
<collectingDate id="36A8567107D55D0FC16A8B401C7C2B29" value="1980-05-07">7 May 1980</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="22AE5269659487F42B4F720677057338" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163780</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="40BB91DD37B7A31C08E6D8CB5C998566" ID-GBIF-Occurrence="4904612353" collectionCode="MBM" country="China" location="Drummond Island" municipality="Xisha Islands" specimenCode="MBM 163781" specimenCount="2" specimenCount-female="2">
|
||
<specimenCount id="905A39BA22F912994BB50DFDE356617E" count="2" type="female">2 females</specimenCount>
|
||
;
|
||
<location id="A8519695E21FB24427587CE459721BC2" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:A8519695E21FB24427587CE459721BC2" country="China" municipality="Xisha Islands" name="Drummond Island">Drummond Island</location>
|
||
,
|
||
<collectingMunicipality id="03E9B236B040D470A4B5D35E5295253F">Xisha Islands</collectingMunicipality>
|
||
; 1980;
|
||
<specimenCode id="F3B0D9A74AEEF9EB2AFFE35D2D440628" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" type="">MBM 163781</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="E1766176494E5CDB1BFF115D0DDB78BC" ID-GBIF-Occurrence="4904612331" collectingDate="2022-05-10" collectorName="Ziming Yuan & Yuli Sun & Shaobo Ma" country="China" elevation="10" latitude="9.8919" location="Meiji Reef" longLatPrecision="1" longitude="115.57279" municipality="Nansha Islands" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="FD51F69D8DA705D64DB6CC72A4101B9B" count="1" type="male">1 male</specimenCount>
|
||
;
|
||
<location id="504F510F406520BF5DBB702E3F2E6DA7" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:504F510F406520BF5DBB702E3F2E6DA7" country="China" latitude="9.8919" longLatPrecision="1" longitude="115.57279" municipality="Nansha Islands" name="Meiji Reef">Meiji Reef</location>
|
||
,
|
||
<collectingMunicipality id="9D536DAFE0F5D690D25190F6C90EF09F">Nansha Islands</collectingMunicipality>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJKAI" specific-use="{"type":"Point","coordinates":[115.572792,9.891900]}">
|
||
<geoCoordinate id="E40C7DB446BEAFF1C8ECD52CAD89A897" degrees="9" direction="north" minutes="53" orientation="latitude" precision="1" seconds="30.84" value="9.8919">9 ° 53 ' 30.84 " N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5F67FEA5B16C9BEC28A16FAA02FF5A69" degrees="115" direction="east" minutes="34" orientation="longitude" precision="1" seconds="22.05" value="115.57279">115 ° 34 ' 22.05 " E</geoCoordinate>
|
||
</named-content>
|
||
;
|
||
<quantity id="74F70F7EA3CBDD4BF4AD8D130B7CD78B" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
|
||
<elevation id="A3E52E159A5E4AD5B844080629E76022" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="BAB7543A210C5BA34F0ED73A798ED7BE" value="2022-05-10">
|
||
<collectingDate id="DD6A272357C370FB1505F2BB733077BE" value="2022-05-10">10 May 2022</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="692A4C6210F43E95E171CD84CA6B6E31">Ziming Yuan</collectorName>
|
||
,
|
||
<collectorName id="45BAF238C17698BEBF21D163828F5FFB">Yuli Sun</collectorName>
|
||
,
|
||
<collectorName id="88951FA4E13EB0B9707155F15E4760BD">Shaobo Ma</collectorName>
|
||
coll.;
|
||
<specimenCode id="C8F64C761817DB927273AEC062381CF4">NS-MJ- 2022-1734</specimenCode>
|
||
.
|
||
<abbrev id="ABBRID0EOKAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
<quantity id="4DBCB0F652AA4D4369D8497958DDCDAE" metricMagnitude="-2" metricUnit="m" metricValue="2.8299999999999996" metricValueMax="3.7299999999999995" metricValueMin="1.9300000000000002" unit="mm" value="28.299999999999997" valueMax="37.3" valueMin="19.3">19.3–37.3 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0ESKAI" xlink_title="median carapace length">CL</abbrev>
|
||
<quantity id="58B022062ACA3DF05AC3336C1610418D" metricMagnitude="-2" metricUnit="m" metricValue="1.975" metricValueMax="2.6399999999999997" metricValueMin="1.31" unit="mm" value="19.75" valueMax="26.4" valueMin="13.1">13.1–26.4 mm</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E02B8AD5212CB4FC959C9BA76E389A48">
|
||
<taxonomicName id="C0940D98FF745F9BF7655646A7B9035B" authority="(Dana, 1852)" baseAuthorityName="Dana" baseAuthorityYear="1852" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="cavipes">
|
||
<emphasis id="73176652923B2BE64689E917A800B302" bold="true">
|
||
<emphasis id="77E706CE3BB5BF2BD805301944FD46D3" bold="true" italics="true">Psaumis cavipes</emphasis>
|
||
(Dana, 1852)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="56542DC705F77FAB95E1A920D10CE549" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 E</figureCitation>
|
||
).
|
||
<materialsCitation id="FCE3FD5F07522D703B704E8D36BCA098" ID-GBIF-Occurrence="4904612308" collectingDate="2019-07-10" country="China" location="Jinqing Island" municipality="Xisha Islands" specimenCount="1" specimenCount-female="1">
|
||
<collectingCountry id="0EB3ADCD7370795B7FB7A2A3EA661B61" name="China">China</collectingCountry>
|
||
•
|
||
<specimenCount id="49E80032DFBFA04311F8A1AE161E0E55" count="1" type="female">1 female</specimenCount>
|
||
ovigerous;
|
||
<collectingMunicipality id="8CCFBBB8A45BCADFCDCF2872583F9B00">Xisha Islands</collectingMunicipality>
|
||
,
|
||
<location id="C95ED0C56B81CFF217CF689F49C29DAF" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:C95ED0C56B81CFF217CF689F49C29DAF" country="China" municipality="Xisha Islands" name="Jinqing Island">Jinqing Island</location>
|
||
;
|
||
<date id="5FCC0D8AEE60CD33BA792F63DCCCA7E5" value="2019-07-10">
|
||
<collectingDate id="5D3ADE880AB1E72D3CA900DD17711D19" value="2019-07-10">10 Jul. 2019</collectingDate>
|
||
</date>
|
||
; azp 01
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="73AEDBAF47A6F4A0F8C16BA1FAD8A374" ID-GBIF-Occurrence="4904612315" collectingDate="2021-04-30" collectorName="Zhang Xu" country="China" location="Sanya Station" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hainan">
|
||
<specimenCount id="74B939B6C8B3CDA52F5B0A263312B350" count="1" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="173427044BBFD64C07BEEEFD93D23A14" count="3" type="female">3 females</specimenCount>
|
||
;
|
||
<location id="B676A012055F136D293305CF7E7EAC3C" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:B676A012055F136D293305CF7E7EAC3C" country="China" name="Sanya Station" stateProvince="Hainan">Sanya Station</location>
|
||
Front,
|
||
<collectingRegion id="C89822CE07F43C678B63CA1AEB0E8A74" country="China" name="Hainan">Hainan</collectingRegion>
|
||
;
|
||
<date id="8AEDDFD541C9D43C7452AA7739446433" value="2021-04-30">
|
||
<collectingDate id="CDD747350E159AFD2A7E1D0FDCE91C71" value="2021-04-30">30 Apr. 2021</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="FB17E5F9AA3F932289FDC8656A38F915">Zhang Xu</collectorName>
|
||
coll.; aop 01
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="13924C21014B2512F89455D26CCFC199" ID-GBIF-Occurrence="4904612317" collectingDate="2019-07-10" country="China" location="Yongle" municipality="Xisha Islands" specimenCount="2" specimenCount-male="2" stateProvince="Hainan">
|
||
<specimenCount id="768DF620C569C082DD1D09C14D0D9EF7" count="2" type="male">2 males</specimenCount>
|
||
;
|
||
<collectingMunicipality id="BAF6C6407F33CC9EFBD5A032822DEDD0">Xisha Islands</collectingMunicipality>
|
||
,
|
||
<location id="DD538A28C73DEBCD6815A724D1C3568C" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:DD538A28C73DEBCD6815A724D1C3568C" country="China" municipality="Xisha Islands" name="Yongle" stateProvince="Hainan">Yongle</location>
|
||
blue hole;
|
||
<date id="1A2AF80BC76117293033608F887FBBFA" value="2019-07-10">
|
||
<collectingDate id="182463F99F556082327F455642929064" value="2019-07-10">10 Jul. 2019</collectingDate>
|
||
</date>
|
||
; aop 02
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="E9D6F92F7D4197A777BBB7DE6FEDC16C" ID-GBIF-Occurrence="4904612361" collectingDate="2019-07-10" country="China" location="Yongle" municipality="Xisha Islands" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
|
||
<specimenCount id="8E80884FE254415E48C6B3D74CB995EA" count="1" type="female">1 female</specimenCount>
|
||
ovigerous;
|
||
<collectingMunicipality id="5DE3740EA2C9A46D38568393D4B6B92A">Xisha Islands</collectingMunicipality>
|
||
,
|
||
<location id="2582A5A87E342DB86B8B31B1D90C4910" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:2582A5A87E342DB86B8B31B1D90C4910" country="China" municipality="Xisha Islands" name="Yongle" stateProvince="Hainan">Yongle</location>
|
||
blue hole;
|
||
<date id="8589445982D5591AC60FD2DB342BBF12" value="2019-07-10">
|
||
<collectingDate id="CF9954CFFED8A0F1AD825EF3467F75E9" value="2019-07-10">10 Jul. 2019</collectingDate>
|
||
</date>
|
||
; aop 03
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="C83D11B10BB6A8C8FA1283F1F8FB8CF2" ID-GBIF-Occurrence="4904612344" collectorName="Zhang Xu" country="China" location="Sanya" municipality="Phoenix Island" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
|
||
<specimenCount id="7C61E14F213B3B94B93A079D138F9F75" count="1" type="male">1 male</specimenCount>
|
||
;
|
||
<collectingMunicipality id="E78D08AF432372236CCA03040FBEF418">Phoenix Island</collectingMunicipality>
|
||
,
|
||
<location id="3F33132E90A19651727ED730F5D63D97" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:3F33132E90A19651727ED730F5D63D97" country="China" municipality="Phoenix Island" name="Sanya" stateProvince="Hainan">Sanya</location>
|
||
,
|
||
<collectingRegion id="9F1BC291E0626FFEAA4AE53CFDC9FED4" country="China" name="Hainan">Hainan</collectingRegion>
|
||
;
|
||
<collectorName id="EF4442CB1ADFA52434519329370663FB">Zhang Xu</collectorName>
|
||
coll.; 2022010
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="9F3631984DFCC4B9610CBD77011D065E" ID-GBIF-Occurrence="4904612328" collectingDate="2019-07-09" country="China" location="Yuzhuo Reef" municipality="Xisha Islands" specimenCount="1" specimenCount-juvenile="1" stateProvince="Hainan">
|
||
<specimenCount id="99344CCC0AA9022A620D139006BA618B" count="1" type="juvenile">1 juvenile</specimenCount>
|
||
;
|
||
<collectingMunicipality id="495260874685B9F17734F90D797F74D5">Xisha Islands</collectingMunicipality>
|
||
,
|
||
<location id="D5692ABB4DC99E50CEEE48321D46B745" LSID="urn:lsid:plazi:treatment:F5D4F37E18E35DD2B528EBEE01F48D06:D5692ABB4DC99E50CEEE48321D46B745" country="China" municipality="Xisha Islands" name="Yuzhuo Reef" stateProvince="Hainan">Yuzhuo Reef</location>
|
||
;
|
||
<date id="930EAD36A536D790DA43B8EEE7F487C5" value="2019-07-09">
|
||
<collectingDate id="F9D92FAA600081147EDED6C43765EB48" value="2019-07-09">9 Jul. 2019</collectingDate>
|
||
</date>
|
||
; aop 04
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="66D67E5BCA6E6564049BC31760E447D5" ID-GBIF-Occurrence="4904612332" collectingDate="2016-11-21" collectorName="Xan" country="China" elevation="10" location="Hainan" specimenCount="2" specimenCount-male="2" stateProvince="Hainan">
|
||
<specimenCount id="85BA78FEBB2EBF5BF6D9D2827FB57B4D" count="2" type="male">2 males</specimenCount>
|
||
;
|
||
<collectingRegion id="F4C9A5E9EC426E79F29AA75552663CEC" country="China" name="Hainan">Hainan</collectingRegion>
|
||
, subtidal
|
||
<quantity id="B027E637D50F94631D1759994A740C5B" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" unit="m" value="9.5" valueMax="10.0" valueMin="9.0">
|
||
<elevation id="8D4CE6F6071AAAE4139814CDDB5C129C" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" unit="m" value="9.5" valueMax="10.0" valueMin="9.0">9–10 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="5CE2E27DF9597E855A705FF199DF9EF6" value="2016-11-21">
|
||
<collectingDate id="7750E133F509045982D5BA70D75D6B55" value="2016-11-21">21 Nov. 2016</collectingDate>
|
||
</date>
|
||
; Xan 020-2.
|
||
<abbrev id="ABBRID0EKLAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
<quantity id="53E582233B268195214E2E9A25A5B6C7" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.7399999999999998" metricValueMin="0.51" unit="mm" value="11.25" valueMax="17.4" valueMin="5.1">5.1–17.4 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EOLAI" xlink_title="median carapace length">CL</abbrev>
|
||
<quantity id="E3CEEEF83699AFF5E7433F9C44CC0AB2" metricMagnitude="-3" metricUnit="m" metricValue="6.949999999999999" metricValueMax="10.6" metricValueMin="3.3" unit="mm" value="6.949999999999999" valueMax="10.6" valueMin="3.3">3.3–10.6 mm</quantity>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ESLAI" type="remarks">
|
||
<paragraph id="61873AF8C438F24A361AFD45CB9EDAC0">
|
||
<heading id="9E5A96F7CEE6B90C5F9228417DFAD8B1" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="7A9CB01597ABE14D1FF674DAAE7B8913">
|
||
<taxonomicName id="D6D3A5EDCC9ED4FD78D7529415FF1E8D" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="014FB4D34A60A2C95BA162A618D855C1" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="086F57D36A6D49B36FA36E255A50730D" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
exhibits the closest resemblance to the subfamily
|
||
<taxonomicName id="B9A741E1B2FE7B837042B9878564AA3C" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Euxanthinae">Euxanthinae</taxonomicName>
|
||
, particularly to Eux 1, as defined and morphologically summarized in the molecular systematic study by
|
||
<bibRefCitation id="6C8C7609412F860F410280A791D74A27" DOI="10.1016/j.jcz.2011.07.002" author="Lai JCY & Mendoza JCE & Guinot D & Clark PF & Ng PKL" etAl="et al." firstAuthor="Lai" issue="4" journalOrPublisher="Zoologischer Anzeiger" pagination="407-448" refId="B23" refString="Lai JCY, Mendoza JCE, Guinot D, Clark PF, Ng PKL (2011) Xanthidae MacLeay, 1838 (Decapoda, Brachyura, Xanthoidea) systematics: A multi-gene approach with support from adult and zoeal morphology. Zoologischer Anzeiger 250 (4): 407–448. https://doi.org/10.1016/j.jcz.2011.07.002" title="Xanthidae MacLeay, 1838 (Decapoda, Brachyura, Xanthoidea) systematics: A multi-gene approach with support from adult and zoeal morphology." volume="250" year="2011">Lai et al. (2011)</bibRefCitation>
|
||
., mainly considering its anterolateral margin of the carapace, which does not clearly meet the orbit but instead continues down to the subhepatic region, presenting an ambiguous starting point (Fig.
|
||
<figureCitation id="26E7219F0BF4363BBF3855D1DAB40F43" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 B</figureCitation>
|
||
). Other characteristics justifying its inclusion are chelipeds almost completely symmetrical, which can be coapted against the carapace (Fig.
|
||
<figureCitation id="73F513B074CBA75738AF61387FD318B5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
|
||
); male pleon long, with the telson reaching to the level above the coxo-sternal condyles of pereiopod 1, and base of somite 3 only slightly wider than tip of somite 5 (Figs
|
||
<figureCitation id="C4C73765DBB5B61FB37F4D7E91BA62D6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A – F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 F</figureCitation>
|
||
,
|
||
<figureCitation id="9A6B12BCCA9B8CA8C64B28230F957338" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
|
||
) (see also
|
||
<bibRefCitation id="59E3DA394C5AFB317B384E7515CC5785" author="Serène R" refId="B42" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1 - 349. [figs A - C + 1 - 243, pls 1 - 48]" year="1984">Serène, 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9174E07A01FDE2FD49D18E1D6C69EB48" DOI="10.1016/j.jcz.2011.07.002" author="Lai" etAl="et al." firstAuthor="Lai" issue="4" journalOrPublisher="Zoologischer Anzeiger" pagination="407-448" refId="B23" refString="Lai JCY, Mendoza JCE, Guinot D, Clark PF, Ng PKL (2011) Xanthidae MacLeay, 1838 (Decapoda, Brachyura, Xanthoidea) systematics: A multi-gene approach with support from adult and zoeal morphology. Zoologischer Anzeiger 250 (4): 407–448. https://doi.org/10.1016/j.jcz.2011.07.002" title="Xanthidae MacLeay, 1838 (Decapoda, Brachyura, Xanthoidea) systematics: A multi-gene approach with support from adult and zoeal morphology." volume="250" year="2011">Lai et al. 2011</bibRefCitation>
|
||
). However, its ambulatory legs do not form a similar perfect coapted structure, especially since the corresponding posterolateral margin is nearly non-concave (Fig.
|
||
<figureCitation id="EB2AF9841DB95FD46C6A575BA2880AE0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
). Other features notably distinguishing it from any member of the subfamily
|
||
<taxonomicName id="8AB76AF1C440598C9553D6D427F5664B" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Euxanthinae">Euxanthinae</taxonomicName>
|
||
are its extremely narrow male pleon with a long and broad, overall truncated oval telson (Figs
|
||
<figureCitation id="3AD93D352E44308073A3DE4483C3DF06" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A – F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 F</figureCitation>
|
||
,
|
||
<figureCitation id="554E2A83A2A16FB8A464B7E311A95CFC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
|
||
), the base of which is wider than the width of the end of the sixth pleonite, the terminal end wide and rounded, with the lateral edges barely converging inward but rather forming two opposing arcs, unlike the typically triangular telson common in the
|
||
<taxonomicName id="038064E56E1BE73AE4186F001BC9A751" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Euxanthinae">Euxanthinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="EA511C47153FF9468A21EDD28FEFE633" ID-DOI="10.3897/zse.100.117859.figure1" ID-arpha="9E67FEC8-E7FA-541D-86D6-4486A48F0EAD" httpUri="https://binary.pensoft.net/fig/1088094" startId="F1">
|
||
<paragraph id="38BF5BD8515981060F9AB6BB4F56E850">
|
||
<label id="2159B6A4A1BAD6CBAFC69F82AC6616AA">Figure 1.</label>
|
||
</paragraph>
|
||
<paragraph id="7D7DAC00F934DDA41706AC99477E778D">
|
||
<taxonomicName id="6BF2A6D5AAA4526E18E201C2717A13D3" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
|
||
<emphasis id="B7966F74BC57D78D73A5C34C5E37838D" italics="true">Gothus teemo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6310A935C6F4220000635C21414AE7CF" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="13F0A634D1384ABF6984C095CB690FCC" bold="true">A – C.</emphasis>
|
||
Holotype, male,
|
||
<abbrev id="ABBRID0EHOAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.7 mm,
|
||
<abbrev id="ABBRID0ELOAI" xlink_title="median carapace length">CL</abbrev>
|
||
2.6 mm, MBM 287027;
|
||
<emphasis id="5AF4AB4D4DB1F508956CFE95BF078D60" bold="true">D – F.</emphasis>
|
||
Paratype,
|
||
<abbrev id="ABBRID0EROAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.2 mm,
|
||
<abbrev id="ABBRID0EVOAI" xlink_title="median carapace length">CL</abbrev>
|
||
2.2 mm, MBM 287026;
|
||
<emphasis id="FAB827111C86EDA7A711208C4E1DC6C2" bold="true">A, D.</emphasis>
|
||
Dorsal view;
|
||
<emphasis id="69E998AAD52B07CCA495AD1BCD2A08AD" bold="true">B, E.</emphasis>
|
||
Frontal view;
|
||
<emphasis id="D8F0C18CE5AEEAB8E4E56B22CB1A06C9" bold="true">C, F.</emphasis>
|
||
Carapace. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9EDDCCC12F2B88AF0D7C7DC7D0CE8E20" ID-DOI="10.3897/zse.100.117859.figure2" ID-arpha="29525E97-5F10-5228-A249-5F63493CF28A" httpUri="https://binary.pensoft.net/fig/1088095" startId="F2">
|
||
<paragraph id="9E6E18827AFE9B62CFC2DCC5E6294642">
|
||
<label id="E64F6E31A7E1FE314CC251B4E36BD37E">Figure 2.</label>
|
||
</paragraph>
|
||
<paragraph id="63FBDFC11E83A12032C3E802A740E0AC">
|
||
<taxonomicName id="5149ACF8AF45FE637A219777CD37E6D2" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
|
||
<emphasis id="25C6E501436E1D275EDA9723031E59E8" italics="true">Gothus teemo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22BB6DAC5B084443D22D0D399424E69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="A514CDCBBCD0F68D8693206BB8E69C00" bold="true">A, C, E.</emphasis>
|
||
Holotype, male,
|
||
<abbrev id="ABBRID0EZPAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.7 mm,
|
||
<abbrev id="ABBRID0E4PAI" xlink_title="median carapace length">CL</abbrev>
|
||
2.6 mm, MBM 287027;
|
||
<emphasis id="A15AD73370B7A6A2BC09A2C0A1B1ACCD" bold="true">B, D, F, G.</emphasis>
|
||
Paratype, female,
|
||
<abbrev id="ABBRID0EDQAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.2 mm,
|
||
<abbrev id="ABBRID0EHQAI" xlink_title="median carapace length">CL</abbrev>
|
||
2.2 mm, MBM 287026;
|
||
<emphasis id="61AD176E11B43E052F013AB8CDFCDF33" bold="true">A.</emphasis>
|
||
Male chelipeds;
|
||
<emphasis id="325417333189B0745DA62235D2763DBD" bold="true">B.</emphasis>
|
||
Female chelipeds;
|
||
<emphasis id="AEF23EE3A17C2CE852E22BD14F38D04A" bold="true">C.</emphasis>
|
||
Male thoracic sternites;
|
||
<emphasis id="E086049D417FA209B143419914DAE82B" bold="true">D.</emphasis>
|
||
Female thoracic sternites;
|
||
<emphasis id="C73EF9A33B4CA564543A2B700158EC15" bold="true">E.</emphasis>
|
||
Male pleon;
|
||
<emphasis id="D806107C8CF28A901100E19869942106" bold="true">F.</emphasis>
|
||
Female pleon;
|
||
<emphasis id="A296985E5661E57A39FF7D5DBC647AFE" bold="true">G.</emphasis>
|
||
Female vulva. Scale bar: 1 mm (
|
||
<emphasis id="90D0BD7A08C736561CB7C60CEEBD3FB8" bold="true">A – F</emphasis>
|
||
); 0.2 mm (
|
||
<emphasis id="CBD97601CB020D658F982697AC60E4A6" bold="true">G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="53B68095186F6E900FDD11D93F1456EB" ID-DOI="10.3897/zse.100.117859.figure3" ID-arpha="331937D4-3489-576E-B1E9-262990AB3EA4" httpUri="https://binary.pensoft.net/fig/1088096" startId="F3">
|
||
<paragraph id="7B2C44A6A64994C778DF8BF4A5A5BC57">
|
||
<label id="779808F587CFED1B1D680BF5429B3067">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="4E782F431D64342CA7E3339D535811EE">
|
||
<taxonomicName id="70C7BDC3E2F78A77B9979135AF765B91" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
|
||
<emphasis id="87370278906F872A7E90B5EB5D842AE3" italics="true">Gothus teemo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DFF77A48C85D2EBEFD74987E40FD1654" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="E2BFC22F7940F24736942431DCD3C52E" bold="true">A – C, F, G – L.</emphasis>
|
||
Holotype, male,
|
||
<abbrev id="ABBRID0EXRAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.7 mm,
|
||
<abbrev id="ABBRID0E2RAI" xlink_title="median carapace length">CL</abbrev>
|
||
2.6 mm, MBM 287027;
|
||
<emphasis id="B5F728BBA8FB1ADE7EB95B11C7D3914F" bold="true">D, E.</emphasis>
|
||
Paratype, female,
|
||
<abbrev id="ABBRID0EBSAI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.2 mm,
|
||
<abbrev id="ABBRID0EFSAI" xlink_title="median carapace length">CL</abbrev>
|
||
2.2 mm, MBM 287026;
|
||
<emphasis id="39E86BC63473D31465FB4FCD7FB99FD5" bold="true">A.</emphasis>
|
||
Carapace;
|
||
<emphasis id="7872007EF257097D8D6A595CE76DF61E" bold="true">B.</emphasis>
|
||
Maxilliped 3;
|
||
<emphasis id="B42AE8725A00B3342CF5158C53C72D45" bold="true">C.</emphasis>
|
||
Cheliped;
|
||
<emphasis id="2E5A8C55D686AD2A28B1C35A7F0974C7" bold="true">D.</emphasis>
|
||
Pereopod 5;
|
||
<emphasis id="5E3B77E2D22DEBF5BCEF6AFC2B387CEB" bold="true">E.</emphasis>
|
||
Female pleon;
|
||
<emphasis id="6CA0F94EB95232AEE76B39C5458524ED" bold="true">F.</emphasis>
|
||
Male pleon;
|
||
<emphasis id="60BCD2B3EC8BB0F3C54EE2D69D80F121" bold="true">G.</emphasis>
|
||
right
|
||
<abbrev id="ABBRID0EXSAI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
, ventral view;
|
||
<emphasis id="5F8A30CA5324B7C8A4E6518B06CC3EDA" bold="true">H.</emphasis>
|
||
Same, dorsal view;
|
||
<emphasis id="C7D47A8A11ECBE88DB5BE8602E0CC256" bold="true">I.</emphasis>
|
||
Right
|
||
<abbrev id="ABBRID0E6SAI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
, distal part, ventral view;
|
||
<emphasis id="16E4465EE330A71147C6822A677FD9F1" bold="true">J.</emphasis>
|
||
Same, dorsal view;
|
||
<emphasis id="FAB2EC46FC50A6ADFB533FF02B92317C" bold="true">K.</emphasis>
|
||
Right
|
||
<abbrev id="ABBRID0EHTAI" xlink_title="second gonopod of male">G 2</abbrev>
|
||
, ventral view;
|
||
<emphasis id="CECFEFC14E69720ADCFAF51586041792" bold="true">L.</emphasis>
|
||
Same, dorsal view. Scale bar: 1 mm (
|
||
<emphasis id="A043B3D5BEB3A874BED3FF047F96120D" bold="true">A, C, D</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis id="AFB186892720B39D9510BFCD8FA0B32E" bold="true">B, E, F, G, H</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis id="B8B94644DEA6FEF5E933E8DFD9E3D11A" bold="true">I – L</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="887117F6C5565DBDA9C00BE7E015E369">
|
||
<taxonomicName id="926F2451E73124615CD956F1A8662022" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="67EDE3FD6C036FAA5154FF9337428710" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D614E18D1F07D5D47FEECFD42F5B3FBC" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
shares the closest similarities with the genus
|
||
<taxonomicName id="7F10218EC3B39BA39B1AE62B60542307" authority="Mendoza & PKL Ng, 2008" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="48EF57AB0A99C3A32955A416DA564FB7" italics="true">Rizalthus</emphasis>
|
||
<bibRefCitation id="37E80F52840E3440606A96B559D288AD" author="Mendoza JCE & Ng PKL" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385 - 404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza & PKL Ng, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
, due to both possessing a granule-covered carapace surface, similar carapace outlines and front, developed cheliped carpus, and analogous
|
||
<abbrev id="ABBRID0EIUAI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
structures. However,
|
||
<taxonomicName id="C25D1177CAA386B413B6AFC77AC167C9" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FCE62CE5785859921108EDC11BB961B0" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from
|
||
<taxonomicName id="B25A228E45DA7B91FC78127C72224E8D" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="803380E750B6715A27CD65EC07CECB5A" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
by several key characteristics: its anterolateral margin with four teeth, first tooth flattened, sometimes completely reduced to appear as three teeth (Figs
|
||
<figureCitation id="C18DB90A95CD27140E7C3FC814328C1F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="9A7EB3DD40DB4ED37101BE1C3B7D27D8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (vs. no clearly defined teeth in
|
||
<taxonomicName id="9BC00DE4120E3E413FD9C72A9FC972C5" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="661F735027CDF8B4C9E1FD49FF263518" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="BA66B5C1D072B892F1A09968C2BC54BB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 A</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="659135080A9D2E055DE467E4951A5644" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: fig. 1 A); absence of etched depressions on body (Fig.
|
||
<figureCitation id="685D7AD64F521DD35F4CF4392C3B364C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
|
||
) (vs. distinct etched depressions on thoracic sternum in
|
||
<taxonomicName id="ED5A8624E2199EBA927B02E99703B029" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2D7EC3D9455A5BC92AF2C470B197DB65" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="FC979381FC0142263C2FD3F05E302178" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: fig. 1 C); central part of epistome raised (Fig.
|
||
<figureCitation id="79655251AADCE9091EEC9E498B92F6B6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 B</figureCitation>
|
||
) (vs. central part of epistome not protruding in
|
||
<taxonomicName id="63631981D7F428BD4B7A5197DA4557AB" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3DB42DD0EBD0D03BAEBEE3F2425D8C58" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="F82D7DBFF38EE518BDBBE7158AC7280B" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: fig. 1 B); robust cheliped carpus, sometimes slightly expanded (Fig.
|
||
<figureCitation id="E3BCA450DB35932411607A74BF81B1EF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
|
||
) (vs. strongly expanded and protruding in
|
||
<taxonomicName id="50BFB99110667E5AFA6EE9C5F1BBDF27" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9FA97E55D68E460697013151E44E0C80" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="9752B584CE02E8C7ECFEE8B8A53C120E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Gothus teemo sp. nov., paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026, artistic illustration, displaying live coloration. Drawn by Fei Gao." figureDoi="10.3897/zse.100.117859.figure5" httpUri="https://binary.pensoft.net/fig/1088098">5 A</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="C0333D1E78662C5DF2C5E85BEC1D8E80" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: fig. 1 A); male pleon with a long, broad, truncated oval telson (Figs
|
||
<figureCitation id="A66CE92E0A2C849D0060A0BAF271677B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A – F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 F</figureCitation>
|
||
,
|
||
<figureCitation id="FD505C7028BD2A3F6693CA6104DBEA43" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
|
||
) (vs. a smaller, triangular telson in
|
||
<taxonomicName id="1D217CCEBCBF7BBA586D76B9A99D34B7" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="36B74979DB12AED0374AB6D74F7248F6" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="39D946DC8601732C19E06D369BCCDFD0" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: fig. 2 C);
|
||
<abbrev id="ABBRID0EVXAI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
distal lobe curved inwards (Fig.
|
||
<figureCitation id="3FDE88EDC7F0DC01E8044D77F61A510A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 G – J</figureCitation>
|
||
) (vs. nearly straight, not curved inwards in
|
||
<taxonomicName id="F447F559718E062546B2B81AFD1881AA" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E639C6EFB83E17E6E8905EA64D795CE0" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="E260218CEBA85AAACB350FA0D14990ED" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: fig. 2 F – H) and
|
||
<abbrev id="ABBRID0EIYAI" xlink_title="second gonopod of male">G 2</abbrev>
|
||
with a longer, straighter distal lobe (Fig.
|
||
<figureCitation id="0AC794D979445AED3D84CAE0C5BF72B0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 K, L</figureCitation>
|
||
) (vs. shorter and curved in
|
||
<taxonomicName id="A80851922B0C1B35A4F1996B91C20615" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FB1D53A527D23B189D7381EC4F33F1AD" italics="true">Rizalthus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="B8988AE4DE1BB675491A2115FE6516C9" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: fig. 2 I).
|
||
</paragraph>
|
||
<paragraph id="8D1EB922A8A99A4CE1686656276314EC">
|
||
Due to its similar carapace outline, particularly the less concave posterolateral margins,
|
||
<taxonomicName id="4CAD0688B9220BD554779FC89818EF68" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="11E0EBD60764F96EBF50D6066F6AB20D" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
also resembles
|
||
<taxonomicName id="270D90727643B872ADDEFE6644C9E8E0" authority="Mendoza & Ng, 2008" authorityName="Mendoza & Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FB27B5400D93CCC282A06235B665F258" italics="true">Visayax</emphasis>
|
||
<bibRefCitation id="E82312DC0EDEC18197A2B2FBA7A5B6D0" author="Mendoza JCE & Ng PKL" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385 - 404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza & Ng, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, it can be easily differentiated by the following characteristics:
|
||
<taxonomicName id="9A04A4B3F9F0D69FCFC88B578A1AE895" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0C2344331AB099848269EC46B712FB4D" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
lacks erosive depressions across body (Fig.
|
||
<figureCitation id="309FF7BA358E4A45C7549984FF4C617C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
|
||
) (vs. chelipeds, ambulatory legs, carapace, and thoracic sternum with erosive depressions in
|
||
<taxonomicName id="26E04CCC606B9F0F4789B9FF1466D871" authorityName="Mendoza & Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C2AAFD46792A35FE0157D05E652A7BF" italics="true">Visayax</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="E3AFA0626ABED684F9292D0C2CB59E2A" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: figs 3–6); carapace regions more flattened (Fig.
|
||
<figureCitation id="E188443856AE4A994D84ACC2A36AA3BC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
) (vs. carapace regions more pronounced in
|
||
<taxonomicName id="A01E2A2BB814B25A200B407598E15081" authorityName="Mendoza & Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="19FAF0A0C130D83C7520AF00FE439602" italics="true">Visayax</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="C0E9ADAD38E9C6E8E60B0DD705A4B364" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: figs 3 A, C, 5 A, C); posterior three teeth on anterolateral margin of carapace well-developed (Fig.
|
||
<figureCitation id="66939BACA3D95401D11A1E4B8A3C684E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="98740A71FE53D58AFA407D9C9CE05602" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (vs. absence of developed teeth on anterolateral margin in
|
||
<taxonomicName id="4FF9352C936F2AB86019627E7AC3C99E" authorityName="Mendoza & Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="88D7913D305AAA05C698F17DDE21EDDF" italics="true">Visayax</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="D7C8A1028625BECF25648FA56A20FE9C" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: figs 3 A, 5 A); male abdominal telson larger, truncated oval (Figs
|
||
<figureCitation id="B70DAECE1503ED242C88A5E5899F5888" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A – F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 F</figureCitation>
|
||
,
|
||
<figureCitation id="C3646B977B4BE3A3C538BA289ED71238" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
|
||
) (vs. smaller, semi-circular in
|
||
<taxonomicName id="A039218B5647707F1ACA601F5225CE61" authorityName="Mendoza & Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4589E6E9E73B39774D39F3DD9B03086E" italics="true">Visayax</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="F6AAD68461F91F97C3B075BD11150CA2" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: figs 4 E, 6 D);
|
||
<abbrev id="ABBRID0EW2AI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
more slender (Fig.
|
||
<figureCitation id="5D974C0A7538D14252DAD1272F8E1CCA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 G – J</figureCitation>
|
||
) (vs.
|
||
<abbrev id="ABBRID0E52AI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
more robust in
|
||
<taxonomicName id="52FC8F7AB3A7DED04C84B15D7DAC788D" authorityName="Mendoza & Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5399E20738B497FCB0EF0B272FBF9CC0" italics="true">Visayax</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="2F918079606EF6B452776DDC9D39BF93" author="Mendoza and Ng" firstAuthor="Mendoza" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385–404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza and Ng 2008</bibRefCitation>
|
||
: figs 4 F, G, 6 F, G).
|
||
</paragraph>
|
||
<caption id="74F0540A2B6677D782EE91A6C0E33EB1" ID-DOI="10.3897/zse.100.117859.figure4" ID-arpha="333B44A7-E891-56F3-8A56-127D1879DDEB" httpUri="https://binary.pensoft.net/fig/1088097" startId="F4">
|
||
<paragraph id="E3711CA4B22FA2C4A4E69DC291376CAC">
|
||
<label id="8D047A23A659EFCD174ABBC8C835C594">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="3EFFE6C8DF7534E14BD849816046C490">
|
||
<taxonomicName id="6CF45AFC57138DFB1609F689B80018E7" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
|
||
<emphasis id="A65479BB42CCFE069251F41A426372C5" italics="true">Gothus teemo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="43677F2C83F1917C29807E3C15EFCB19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="84CA39F43FBF737C7C63DEF7C91CD457" bold="true">A, B.</emphasis>
|
||
Holotype, male,
|
||
<abbrev id="ABBRID0EC4AI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.7 mm,
|
||
<abbrev id="ABBRID0EG4AI" xlink_title="median carapace length">CL</abbrev>
|
||
2.6 mm, MBM 287027;
|
||
<emphasis id="F6AC86B2B9CE427D6F5EDB0BB3F578A2" bold="true">C – E.</emphasis>
|
||
Paratypes, 2 juvenile,
|
||
<abbrev id="ABBRID0EM4AI" xlink_title="maximum carapace width">CW</abbrev>
|
||
1.8–2.2 mm,
|
||
<abbrev id="ABBRID0EQ4AI" xlink_title="median carapace length">CL</abbrev>
|
||
1.3–1.5 mm, MBM 287023;
|
||
<emphasis id="361D96E89994A09770DCBFFAE8DFF43A" bold="true">A – D.</emphasis>
|
||
Live coloration;
|
||
<emphasis id="49D887A887293DC5CC3DA71D0FEDE9C3" bold="true">E.</emphasis>
|
||
Habitat and substrate conditions. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BD88A26C1545864FDC6CF0B0DFA187D0" ID-DOI="10.3897/zse.100.117859.figure5" ID-arpha="FAF93A52-F96A-5F99-B6E7-27CDA1C3E9FC" httpUri="https://binary.pensoft.net/fig/1088098" startId="F5">
|
||
<paragraph id="2B331E807DB4904133E6D01AC53D799A">
|
||
<label id="8788704349C1CBB550A11A1CB37783A9">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="D92540A994FCAA9B0B3E7DB492042B8B">
|
||
<taxonomicName id="B8B8BAB6AAF1F29E1B7F3CFEFC21416A" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="teemo" status="sp. nov.">
|
||
<emphasis id="55DCFD28BD4463DA569D0EAA6117FADE" italics="true">Gothus teemo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FF6E26BA320234ABC59E0D336F99DE22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, female,
|
||
<abbrev id="ABBRID0EQ5AI" xlink_title="maximum carapace width">CW</abbrev>
|
||
3.2 mm,
|
||
<abbrev id="ABBRID0EU5AI" xlink_title="median carapace length">CL</abbrev>
|
||
2.2 mm, MBM 287026, artistic illustration, displaying live coloration. Drawn by Fei Gao.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E425057BB5E51CD40F1A1F079B5D8474" ID-DOI="10.3897/zse.100.117859.figure6" ID-arpha="E49065C0-063E-5E34-8A15-2ACC674275C2" httpUri="https://binary.pensoft.net/fig/1088099" startId="F6">
|
||
<paragraph id="70B0ADD62A1100932069C3DB8AC06777">
|
||
<label id="ED2FB4DB5FCC9E13BBBFE3FEF01370D0">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="F108CFDDA6FA7416E57C973AA23813BD">
|
||
<taxonomicName id="614DB4F0727DCD20CA0467D738440676" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Euxanthinae">Euxanthinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="C0E151413A7BA197D1CB4249584FB934">
|
||
<emphasis id="16CF3AA89BE74F2705D012062859C9F1" italics="true">Actaeodes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in comparative material.
|
||
<emphasis id="2885E6B068BEBD9D25AB8E8CE6F0C8C9" bold="true">A.</emphasis>
|
||
<taxonomicName id="A93373A5B3F23A979EF7BACD34A1B332" authority="Mendoza & PKL Ng, 2008" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="anconis">
|
||
<emphasis id="E56098960C1E0B2D40A7D6D9AB1A41B1" italics="true">Rizalthus anconis</emphasis>
|
||
<bibRefCitation id="74D36376D74A475C98D6C8FC35C6F7B1" author="Mendoza JCE & Ng PKL" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="385-404" refId="B25" refString="Mendoza JCE, Ng PKL (2008) New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines. The Raffles Bulletin of Zoology 56: 385 - 404." title="New genera and species of Euxanthine crabs (Crustacea, Decapoda, Brachyura, Xanthidae) from the Bohol Sea, the Philippines." volume="56" year="2008">Mendoza & PKL Ng, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
, female;
|
||
<abbrev id="ABBRID0E56AI" xlink_title="maximum carapace width">CW</abbrev>
|
||
4.2 mm,
|
||
<abbrev id="ABBRID0ECABI" xlink_title="median carapace length">CL</abbrev>
|
||
2.7 mm,
|
||
<specimenCode id="321A4C9F4F65B199A1F4202DC7281C11">NS-MJ- 2022-1457</specimenCode>
|
||
;
|
||
<emphasis id="5812C85F98692308DD7928E0E0192A02" bold="true">B.</emphasis>
|
||
<taxonomicName id="AC85A0195E16A99A4C7B549857F03708" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="haanii">
|
||
<emphasis id="DF376BC8E7831A648FC2FCCC5256B218" italics="true">Hypocolpus haanii</emphasis>
|
||
<bibRefCitation id="B26E919AA7B124718F126EEEB09D2AE2" author="Rathbun MJ" journalOrPublisher="Proceedings of the Biological Society of Washington" pagination="107-114" refId="B37" refString="Rathbun MJ (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington 22: 107 - 114." title="New crabs from the Gulf of Siam." volume="22" year="1909">Rathbun, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
, 1 male,
|
||
<abbrev id="ABBRID0ETABI" xlink_title="maximum carapace width">CW</abbrev>
|
||
45.3 mm,
|
||
<abbrev id="ABBRID0EXABI" xlink_title="median carapace length">CL</abbrev>
|
||
34.2 mm, MBM 286755;
|
||
<emphasis id="5089538F55C360038E93766B7C34FAD2" bold="true">C.</emphasis>
|
||
<taxonomicName id="A863788734BC391791416611039E337D" authority="(Herbst, 1790)" baseAuthorityName="Herbst" baseAuthorityYear="1790" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="exsculptus">
|
||
<emphasis id="D485E1E8D8209F709319603652BB95D0" italics="true">Euxanthus exsculptus</emphasis>
|
||
(Herbst, 1790)
|
||
</taxonomicName>
|
||
, 1 male,
|
||
<abbrev id="ABBRID0EIBBI" xlink_title="maximum carapace width">CW</abbrev>
|
||
52.6,
|
||
<abbrev id="ABBRID0EMBBI" xlink_title="median carapace length">CL</abbrev>
|
||
33.2 mm, MBM 163793;
|
||
<emphasis id="C330F30741C7D4F6681DB3D3DCBD2F7E" bold="true">D.</emphasis>
|
||
<taxonomicName id="4EF5F24E57023E3821AD739BA8CB20EA" authority="(Hombron & Jacquinot, 1846)" baseAuthorityName="Hombron & Jacquinot" baseAuthorityYear="1846" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="huonii">
|
||
<emphasis id="5F4FE345DC008F2CE5EDD81723FBA58D" italics="true">Euxanthus huonii</emphasis>
|
||
(Hombron & Jacquinot, 1846)
|
||
</taxonomicName>
|
||
,
|
||
<abbrev id="ABBRID0E4BBI" xlink_title="maximum carapace width">CW</abbrev>
|
||
34.0 mm,
|
||
<abbrev id="ABBRID0EBCBI" xlink_title="median carapace length">CL</abbrev>
|
||
24.0 mm,
|
||
<specimenCode id="D221E06A4560BBB8E43839AC53F94E65">NS-MJ- 2022-1734</specimenCode>
|
||
;
|
||
<emphasis id="1B43B5E22396BB70386C0FA462DE982E" bold="true">E.</emphasis>
|
||
<taxonomicName id="8D948F141EDC2AE13090E8F2EF5A4D86" authority="(Dana, 1852)" baseAuthorityName="Dana" baseAuthorityYear="1852" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="cavipes">
|
||
<emphasis id="05C60B640871D98AE71768A83C4C315D" italics="true">Psaumis cavipes</emphasis>
|
||
(Dana, 1852)
|
||
</taxonomicName>
|
||
, 1 male,
|
||
<abbrev id="ABBRID0ESCBI" xlink_title="maximum carapace width">CW</abbrev>
|
||
13.8 mm,
|
||
<abbrev id="ABBRID0EWCBI" xlink_title="median carapace length">CL</abbrev>
|
||
8.6 mm, aop 01;
|
||
<emphasis id="740DCA157D1F0C03DA989A2CFC755422" bold="true">F.</emphasis>
|
||
<taxonomicName id="CE815624970714BADF1285982249C0A3" authority="Guinot, 1976" authorityName="Guinot" authorityYear="1976" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="mutatus">
|
||
<emphasis id="747592ED87261050197183B8793510AB" italics="true">Actaeodes mutatus</emphasis>
|
||
<bibRefCitation id="96434FA60EAB6B42632E7FB41FFAAE61" author="Guinot D" refId="B16" refString="Guinot D (1976) Constitution de quelques groupes naturels chez les Crustaces Decapodes Brachyoures. I. La Superfamille des Bellioidea Dana et Trois Sous-Familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Memoires du Museum National d'Histoire Naturelle, Paris: A, 97, 1 - 308. [figs 1 - 47, pls 1 - 19]" year="1976">Guinot, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
, 1 female,
|
||
<abbrev id="ABBRID0EHDBI" xlink_title="maximum carapace width">CW</abbrev>
|
||
20.8 mm,
|
||
<abbrev id="ABBRID0ELDBI" xlink_title="median carapace length">CL</abbrev>
|
||
12.5 mm, BF 02;
|
||
<emphasis id="7A8B9D281A630BF66882BB7DC8F70372" bold="true">G.</emphasis>
|
||
<taxonomicName id="DD25DD063110A402E1AF6F720EDAAA4F" authority="(Ruppell, 1830)" baseAuthorityName="Ruppell" baseAuthorityYear="1830" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="hirsutissimus">
|
||
<emphasis id="686B1B386E4B7046A02CF1B39C8086E0" italics="true">Actaeodes hirsutissimus</emphasis>
|
||
(
|
||
<bibRefCitation id="52691B709FC44A45AA2322B1D986E051" author="Rüppell E" journalOrPublisher="Brönner, Frankfurt am Main" refId="B39" refString="Ruppell E (1830) Beschreibung und abbildung von 24 arten kurzschwanzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L. Bronner, Frankfurt am Main, 28 pp. [6 pls]" title="Beschreibung und abbildung von 24 arten kurzschwänzigen krabben, als beitrag zur naturgeschichte des rothen meeres. H. L." year="1830">Rüppell, 1830</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, 1 male,
|
||
<abbrev id="ABBRID0E3DBI" xlink_title="maximum carapace width">CW</abbrev>
|
||
31.9 mm,
|
||
<abbrev id="ABBRID0EAEBI" xlink_title="median carapace length">CL</abbrev>
|
||
21.0 mm, MBM 164298;
|
||
<emphasis id="7A1B499A78D3C1959CAF1826AC563B2C" bold="true">H.</emphasis>
|
||
<taxonomicName id="272243FEC67CEFC89277FE029CDA9180" baseAuthorityName="H. Milne-Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Xanthidae" genus="Actaeodes" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="species" species="tomentosus">
|
||
<emphasis id="D13B3DC5F7F33504BDAD20BF3F5EE92F" italics="true">Actaeodes tomentosus</emphasis>
|
||
</taxonomicName>
|
||
(H. Milne Edwards, 1834), 1 male,
|
||
<abbrev id="ABBRID0EREBI" xlink_title="maximum carapace width">CW</abbrev>
|
||
35.8 mm,
|
||
<abbrev id="ABBRID0EVEBI" xlink_title="median carapace length">CL</abbrev>
|
||
22.5 mm, Xan 041. Scale bar: 1 mm (
|
||
<emphasis id="F681C77617E8C99A1450A560D26FC88A" bold="true">A</emphasis>
|
||
); 5 mm (
|
||
<emphasis id="3E02D2799CA7670965D1E5064B7B1C83" bold="true">E, F</emphasis>
|
||
); 10 mm (
|
||
<emphasis id="60CE192F9FBAD01C316A6A2F1342646B" bold="true">B – D, G, F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0FDA7573D272B3C7CDFE05729A859B07">
|
||
The new genus exhibits a general morphological similarity to typical
|
||
<taxonomicName id="418961768593A5C6FA700901CCC7E406" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Euxanthinae">Euxanthinae</taxonomicName>
|
||
members such as
|
||
<taxonomicName id="82A3BD82E7778CE10A580D2E5F263708" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D92863BE9EF350157E666C788FAACD24" italics="true">Euxanthus</emphasis>
|
||
<bibRefCitation id="CF9E3D5403526F3D5A86D89DF86B9BB7" author="Dana JD" refId="B4" refString="Dana JD (1851) On the Classification of the Cancroidea. The American Journal of Science and Arts 2 12 (34): 121 - 131." year="1851">Dana, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3DAADEC49D63C153798CF026E47CE9A3" authority="Rathbun, 1897" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DE587CB3E7C872060AF4CDFA462464A4" italics="true">Hypocolpus</emphasis>
|
||
<bibRefCitation id="E51ACDB528AFB908162B4D79C22DCD16" author="Rathbun MJ" journalOrPublisher="Proceedings of the Biological Society of Washington" pagination="153-167" refId="B36" refString="Rathbun MJ (1897) A revision of the nomenclature of the Brachyura. Proceedings of the Biological Society of Washington 11: 153 - 167." title="A revision of the nomenclature of the Brachyura." volume="11" year="1897">Rathbun, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
. In addition to the existing comparative specimens,
|
||
<bibRefCitation id="2958A4CA9CC6768621904F8367A313BC" author="Guinot-Dumortier D." firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier (1960)</bibRefCitation>
|
||
provided excellent descriptions and photographs of species from the above two genera. Subsequently published species also have relatively clear morphological descriptions and images available for comparison (cf.
|
||
<bibRefCitation id="5D5A0507B12E8501A39E4E6D1EC3DE0A" author="Guinot D" journalOrPublisher="Cahiers du Pacifique:" pagination="19-21" refId="B15" refString="Guinot D (1971 b) Un nouvel Euxanthus de Nouvelle-Calédonie; E. ruali sp. nov. Cahiers du Pacifique: 15: 19–21. [pl. 1–2]" title="Un nouvel Euxanthus de Nouvelle-Calédonie; E. ruali sp. nov." volume="15">Guinot 1971 b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DAA5AE2E8BFE89F2298663CAD6BC443E" DOI="10.1080/03946975.1990.10539447" author="Galil and Vannini" firstAuthor="Galil" issue="1" journalOrPublisher="Tropical Zoology" pagination="21-56" refId="B9" refString="Galil BS, Vannini M (1990) Research on the coast of Somalia. Xanthidae, Trapeziidae, Carpiliidae, Menippidae (Crustacea, Brachyura). Tropical Zoology 3 (1): 21–56. https://doi.org/10.1080/03946975.1990.10539447" title="Research on the coast of Somalia. Xanthidae, Trapeziidae, Carpiliidae, Menippidae (Crustacea, Brachyura)." volume="3" year="1990">Galil and Vannini 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F666B988E7A1B76A34F91FF5C1DEFDD5" DOI="10.5962/p.290344" author="Crosnier" firstAuthor="Crosnier" issue="3-4" journalOrPublisher="A," pagination="557-564" refId="B3" refString="Crosnier A (1996) Hypocolpus pararugosus, espèce nouvelle de l’Indo-Ouest Pacifique (Crustacea, Decapoda, Brachyura, Xanthidae). Bulletin du Muséum National d’Histoire Naturelle 4, sec. A, 18 (3–4): 557–564. https://doi.org/10.5962/p.290344" title="Hypocolpus pararugosus, espèce nouvelle de l’Indo-Ouest Pacifique (Crustacea, Decapoda, Brachyura, Xanthidae). Bulletin du Muséum National d’Histoire Naturelle 4, sec." volume="18" year="1996">Crosnier 1996</bibRefCitation>
|
||
). The new genus can be easily distinguished from
|
||
<emphasis id="A8E3DF04359A46D7662F79C578DC202C" italics="true">Euxanthu</emphasis>
|
||
s by the following features: entire body covered with granules and short pubescence (Fig.
|
||
<figureCitation id="01888EF84FF62066564807259370E7B4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
|
||
) (vs. relatively smooth in
|
||
<taxonomicName id="2A036C4A88C131377CBFD11495855BF5" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="57198F62460E4639CBEE88809CEFCED1" italics="true">Euxanthus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="80ED58BEEBFA89C8BD9D631768FD19E3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 C, D</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="A5927F2292208295B4132A9BB170D3C8" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VIII, figs 42, 44, 46, pl. IX, figs 48–52); carapace anterolateral margin with four teeth, first tooth flattened, sometimes completely reduced to appear as three teeth (Figs
|
||
<figureCitation id="05E4F8194AF35C74DB299479C7A9E8DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="A39E047CDFB8012C7132E5AE6AC4D00D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (vs. 4–6 teeth on anterolateral margin in
|
||
<taxonomicName id="8682077693A764710996DC43F05E4938" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="994C7B16DC3D4E633ECEADFE46F10A5A" italics="true">Euxanthus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="CD86DBCFC9C1FFD54E6E26AE72109870" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 C, D</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="20DFD9E091F849E04CC06058C3B7AC87" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VIII, figs 42, 44, 46, pl. IX, figs 48–52); front not prominent, divided by a V-shaped notch (Figs
|
||
<figureCitation id="C515500E9D10AA69726D659A16694F68" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 C</figureCitation>
|
||
,
|
||
<figureCitation id="1BDFD2AA50055593F674521AD89A4CAD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (vs. more protruding, divided by a narrow fissure in
|
||
<taxonomicName id="2D7A3077E896A66CC13CBE59B5B9A1B1" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C2EC5622AEAD3514BE4C121F9844A108" italics="true">Euxanthus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="D2E69B61A33C85623B79EAED2B6C4739" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 C, D</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="6E84B7FB3A934E365D288B2AEC2BDFF1" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VIII, figs 42, 44, 46, pl. IX, figs 48–52); male abdominal telson large and truncated oval (Figs
|
||
<figureCitation id="8AE45CD722FEE5D431EEEC3ACB130B85" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A – F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 F</figureCitation>
|
||
,
|
||
<figureCitation id="6BF1EC048F542EA9BE21DFA5D3FF2EE7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
|
||
) (vs. small and triangular in
|
||
<taxonomicName id="E129FFD52CDAA995C5055950AC74AF15" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A9C06CB1860546A7008B68615260B936" italics="true">Euxanthus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="B855462103D7ECDC079E362271AF53E4" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VIII, fig. 47);
|
||
<abbrev id="ABBRID0EEJBI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
with a prominent, spoon-shaped distal lobe, and long setae on inner subdistal part (Fig.
|
||
<figureCitation id="954DF2624C4AE5A7A724BAD33A86DBCE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 G – J</figureCitation>
|
||
) (vs.
|
||
<abbrev id="ABBRID0EMJBI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
with a short, non-protruding distal lobe, inwardly curved and encircling, with short setae on inner subdistal part in
|
||
<taxonomicName id="38138B0821611ADB9E29C82E252C63BD" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9DD70BAFA63C1414F1F9A63B374A8E29" italics="true">Euxanthus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="21AEF24F12C50F3775869D7992676D2E" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VI, figs 36–39). Similarly, the new genus is easily distinguishable from
|
||
<taxonomicName id="778A2BB91529AFBB3ABC10557930DDCD" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3BC5FD139DAAF2224F77E793DDF77C33" italics="true">Hypocolpus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of a developed subhepatic cavity (Fig.
|
||
<figureCitation id="23E7C48430CC5D27F6CEEF6639C84574" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 B</figureCitation>
|
||
) (vs. a developed subhepatic cavity in
|
||
<taxonomicName id="ED86C5EE5017A54BE3A80579EC285CAD" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="45E8F7210FB088D8FF0F18670E854C1F" italics="true">Hypocolpus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="8960515B3966B98224E3492744F14609" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. II, figs 40–41); posterior three teeth on anterolateral margin well-developed (Figs
|
||
<figureCitation id="401ACA2CCEC070BF9769096A8FDDF866" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 C</figureCitation>
|
||
,
|
||
<figureCitation id="142742F72D7B23DAD764497009A35A22" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (vs. underdeveloped teeth in
|
||
<taxonomicName id="0CC400502472421151AB0B3A56D37F77" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C46F3B2AE72D55488EEEB7997997B0EF" italics="true">Hypocolpus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="29D3D5BADA6A3653B7398EF0B686AE6F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 B</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="8213E932C60F5B9588C8DD04489EE0C9" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VII, figs 40–41); front not prominent, divided by a V-shaped notch (Figs
|
||
<figureCitation id="C9E5E128940824456C1FE1EDB6402277" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 C</figureCitation>
|
||
,
|
||
<figureCitation id="2282C4B347318E3D2E9CC900F35BDBE8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (vs. more protruding, divided by a narrow fissure in
|
||
<taxonomicName id="F5E116BDCD7E40DA599F150FE65DF1A1" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6E546CF335AC4F4CA3CB2D9C861D3989" italics="true">Hypocolpus</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="9B70D3C1BECB3E428996BE75770FCC2C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 B</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="FD7E5852092EA1DC1CE694F2B48F294D" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VII, figs 40–41); male abdominal telson large and truncated oval (Figs
|
||
<figureCitation id="03472CE6BD8FAC5955FBEA692831685A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Gothus teemo sp. nov. A, C, E. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; B, D, F, G. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Male chelipeds; B. Female chelipeds; C. Male thoracic sternites; D. Female thoracic sternites; E. Male pleon; F. Female pleon; G. Female vulva. Scale bar: 1 mm (A – F); 0.2 mm (G)." figureDoi="10.3897/zse.100.117859.figure2" httpUri="https://binary.pensoft.net/fig/1088095">2 F</figureCitation>
|
||
,
|
||
<figureCitation id="6222379E1F3457CC87524375B03683B1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 F</figureCitation>
|
||
) (vs. small and triangular in
|
||
<taxonomicName id="71966950A6C922517B44E0F2CD2B0840" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0F02AC2BC216920119773AB1D26467AF" italics="true">Hypocolpus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="7C90B9C76CE9786F1646E81E6E145388" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. IX, fig. 53, pl. X, fig. 55);
|
||
<abbrev id="ABBRID0ESMBI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
with a prominent, spoon-shaped distal lobe (Fig.
|
||
<figureCitation id="7FA26F8805203000F2774CF0C97402AD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 G – J</figureCitation>
|
||
) (vs.
|
||
<abbrev id="ABBRID0E1MBI" xlink_title="first gonopod of male">G 1</abbrev>
|
||
with a short, non-protruding distal lobe, inwardly curved and encircling in
|
||
<taxonomicName id="F8792F7F66E4BF04DD093EDC362439F8" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAA30C3FA2414D8AA5293631AAB273E" italics="true">Hypocolpus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="F8E5E36530C659FF31100FD210B70879" author="Guinot-Dumortier" firstAuthor="Guinot-Dumortier" issue="2" journalOrPublisher="Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie" pagination="153-218" refId="B17" refString="Guinot-Dumortier D. (1960) Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura). Memoirs du Museum national d’Histoire naturelle, A (Zoology), nouvelle sèrie 20 (2): 153–218." title="Révision des genres Euxanthus Dana et Hypocolpus Rathbun (Crustacea, Decapoda, Brachyura)." volume="20" year="1960">Guinot-Dumortier 1960</bibRefCitation>
|
||
: pl. VI, figs 32–35).
|
||
</paragraph>
|
||
<paragraph id="8984042887A1F80BBAE7D5FD2517AC35">
|
||
The new genus slightly resembles
|
||
<taxonomicName id="153488E81B9F54F7465330BAFF7B0C1F" authority="Kossmann, 1877" authorityName="Kossmann" authorityYear="1877" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2ECE1B416A17757BEF9E80EFAEF50A6F" italics="true">Psaumis</emphasis>
|
||
<bibRefCitation id="982747E8C06396BDB7A98279E69342C5" DOI="10.5962/bhl.title.12140" author="Kossmann R" journalOrPublisher="Wilhelm Engelmann, Leipzig" refId="B22" refString="Kossmann R (1877) Malacostraca (I. Theil, Brachyura). Zoologische ergebnisse einer im auftrage der koniglichen academie der wissenschaften zu Berlin ausgefuhrten reise in die kustengebiete des Rothen Meeres. Wilhelm Engelmann, Leipzig, 66 pp. [3 pls] https: // doi. org / 10.5962 / bhl. title. 12140" title="Malacostraca (I. Theil, Brachyura). Zoologische ergebnisse einer im auftrage der königlichen academie der wissenschaften zu Berlin ausgeführten reise in die küstengebiete des Rothen Meeres." year="1877">Kossmann, 1877</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="59398E4719555D29324C5CAE5A41FD50" authority="Serene, 1984" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Paractaeopsis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AF2DBDA0EF512F505D56F98DB0C5D1E0" italics="true">Paractaeopsis</emphasis>
|
||
<bibRefCitation id="0B1B90D9C1C96B140776F3B7BF2FBDEB" author="Serène R" refId="B42" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1 - 349. [figs A - C + 1 - 243, pls 1 - 48]" year="1984">Serène, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
, but can be readily distinguished.
|
||
<taxonomicName id="6CA504AAB7407B90C9819ED9F1BB3B24" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8CD641100F204F10534DC9F339094FAE" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from
|
||
<taxonomicName id="ADB3E49DB088C17700C2F09EDE169264" authorityName="Kossmann" authorityYear="1877" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6268A3B23FF952A5C6E23195EF3993F5" italics="true">Psaumis</emphasis>
|
||
</taxonomicName>
|
||
by lack of erosive depressions across body (Fig.
|
||
<figureCitation id="8DBE64E751A61CFC1E6B1DF80E6DC5BC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
|
||
) (vs. chelipeds, ambulatory legs, carapace with strong erosive depressions in
|
||
<taxonomicName id="A24A1CAFC090D91681B923A313D37465" authorityName="Kossmann" authorityYear="1877" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EA3B37FA9920EEA896AF1C99A2E89D09" italics="true">Psaumis</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="18ED19158AFFD943F0248F8452CD0F23" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 E</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="4887F822E8DCEDCADFA15EACDA296B13" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de l’Océan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1–349. [figs A – C + 1–243, pls 1–48]" year="1984">Serène 1984</bibRefCitation>
|
||
: pl. XVIII, fig. E); front divided by a V-shaped notch (Figs
|
||
<figureCitation id="4C0115C38748B190D5FEF2F03B1B964F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 C</figureCitation>
|
||
,
|
||
<figureCitation id="2689BAA8C562E0619C7BCCE1EA92C6C4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (front divided by a narrow fissure in
|
||
<taxonomicName id="66B016DA01913883131ABC754EE84602" authorityName="Kossmann" authorityYear="1877" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C41AFFC84ADCD20C8F23AFB77FF61F9C" italics="true">Psaumis</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="8F6072A68380820450DCEF9C47BA5FE0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 E</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="3788F0C5CA501904E4F0730FBD58A5C1" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de l’Océan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1–349. [figs A – C + 1–243, pls 1–48]" year="1984">Serène 1984</bibRefCitation>
|
||
: pl. XVIII, fig. E); anterolateral margin with four teeth, first tooth flattened, sometimes completely reduced to appear as three teeth (Figs
|
||
<figureCitation id="53F77FE914C57D3DBCB093FA762A2D9A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="1D3EFBA5AA358F478E316E843C5E8804" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (anterolateral margin with very flat teeth, except for fourth tooth at junction of anterior and posterior lateral margins, which is more prominent in
|
||
<taxonomicName id="4AF3260D46FF68DE37CCAD338A3C3D9B" authorityName="Kossmann" authorityYear="1877" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BE56F6F1283BBC160D6F6083233ACF40" italics="true">Psaumis</emphasis>
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="AEE726A56D6C52ECB371220F75D25822" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Euxanthinae and Actaeodes species in comparative material. A. Rizalthus anconis Mendoza & PKL Ng, 2008, female; CW 4.2 mm, CL 2.7 mm, NS-MJ- 2022-1457; B. Hypocolpus haanii Rathbun, 1909, 1 male, CW 45.3 mm, CL 34.2 mm, MBM 286755; C. Euxanthus exsculptus (Herbst, 1790), 1 male, CW 52.6, CL 33.2 mm, MBM 163793; D. Euxanthus huonii (Hombron & Jacquinot, 1846), CW 34.0 mm, CL 24.0 mm, NS-MJ- 2022-1734; E. Psaumis cavipes (Dana, 1852), 1 male, CW 13.8 mm, CL 8.6 mm, aop 01; F. Actaeodes mutatus Guinot, 1976, 1 female, CW 20.8 mm, CL 12.5 mm, BF 02; G. Actaeodes hirsutissimus (Rüppell, 1830), 1 male, CW 31.9 mm, CL 21.0 mm, MBM 164298; H. Actaeodes tomentosus (H. Milne Edwards, 1834), 1 male, CW 35.8 mm, CL 22.5 mm, Xan 041. Scale bar: 1 mm (A); 5 mm (E, F); 10 mm (B – D, G, F)." figureDoi="10.3897/zse.100.117859.figure6" httpUri="https://binary.pensoft.net/fig/1088099">6 E</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="1EDC9AB214DF569E970003D108A42871" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de l’Océan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1–349. [figs A – C + 1–243, pls 1–48]" year="1984">Serène 1984</bibRefCitation>
|
||
: pl. XVIII, fig. E). It can be distinguished from
|
||
<taxonomicName id="4A9C870648B3CBB8311566F36A9017F3" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Paractaeopsis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="04DFC087B4DBF916F337D4BAD08B47E3" italics="true">Paractaeopsis</emphasis>
|
||
</taxonomicName>
|
||
by anterolateral margin with four teeth, first tooth flattened, sometimes completely reduced to appear as three teeth (Figs
|
||
<figureCitation id="8E941E61C235B9AA2B5F38AAAB5820DF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="C61A4E1689F011E1C8250BD9039C6E59" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (anterolateral margin with four well development teeth in
|
||
<taxonomicName id="2EDF2BCB56093AC8DC49A34F04FA07B3" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Paractaeopsis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74F4A36170B8CCF18FD57CC5F12D4FD9" italics="true">Paractaeopsis</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="C108AAC97D2F7C85B044E2B9DC3FCF20" author="Takeda and Miyake" firstAuthor="Takeda" issue="9" journalOrPublisher="OHMU, Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University" pagination="183-189" refId="B45" refString="Takeda M, Miyake S (1968) Two new xanthid crabs inhabiting coral reefs of the Ryukyu Islands. OHMU, Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University 1 (9): 183–189. [pl. 8]" title="Two new xanthid crabs inhabiting coral reefs of the Ryukyu Islands." volume="1" year="1968">Takeda and Miyake 1968</bibRefCitation>
|
||
: fig. 1 a); carapace broad, approximately 1.5 times as wide as long, with a relatively flat dorsal surface (Figs
|
||
<figureCitation id="FCCCE97E549CD5EEEEAE0AC9BD0E887C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="A70A7B51A560D98E84906F26B7F22C5F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (carapace narrower, with a width not exceeding 1.4 times the length, and dorsal surface convex both anteroposteriorly and laterally in
|
||
<taxonomicName id="26E4B969D78BFB6DFE9EAE853831737C" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Paractaeopsis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F803947C58E167C94CF020D41B347D0D" italics="true">Paractaeopsis</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="218216BC0FD10B3C278352AF059335FE" author="Serene" firstAuthor="Serene" refId="B42" refString="Serène R (1984) Crustacés Décapodes Brachyoures de l’Océan Indien Occidental et de la Mer Rouge, Xanthoidea, Xanthidae et Trapeziidae. Faune Tropicale, no. XXIV, 1–349. [figs A – C + 1–243, pls 1–48]" year="1984">Serène 1984</bibRefCitation>
|
||
: pl. XVII, fig. E); carapace 2 M region divided into two lobes (Figs
|
||
<figureCitation id="31ECCAFEA4B1B0F3C3377EB4F71375B1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="50247BD71B9847EF9EE8D4D907304862" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 A</figureCitation>
|
||
) (carapace 2 M region divided into four lobes in
|
||
<taxonomicName id="829768593DBC4EA295F0167A2F724B62" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Paractaeopsis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DBEDD4875177A231C77BE9DD9F2E6F05" italics="true">Paractaeopsis</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="222978B408BBAB46E3FDE431FA1B29CE" author="Takeda and Miyake" firstAuthor="Takeda" issue="9" journalOrPublisher="OHMU, Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University" pagination="183-189" refId="B45" refString="Takeda M, Miyake S (1968) Two new xanthid crabs inhabiting coral reefs of the Ryukyu Islands. OHMU, Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University 1 (9): 183–189. [pl. 8]" title="Two new xanthid crabs inhabiting coral reefs of the Ryukyu Islands." volume="1" year="1968">Takeda and Miyake 1968</bibRefCitation>
|
||
: fig. 1 a); ambulatory legs comparatively slender (Figs
|
||
<figureCitation id="4F5E1294A8E39D1BC4968C0A43E52A2E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Gothus teemo sp. nov. A – C. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D – F. Paratype, CW 3.2 mm, CL 2.2 mm, MBM 287026; A, D. Dorsal view; B, E. Frontal view; C, F. Carapace. Scale bar: 1 mm." figureDoi="10.3897/zse.100.117859.figure1" httpUri="https://binary.pensoft.net/fig/1088094">1 A</figureCitation>
|
||
,
|
||
<figureCitation id="7C174611E853F7330E068458178D624E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gothus teemo sp. nov. A – C, F, G – L. Holotype, male, CW 3.7 mm, CL 2.6 mm, MBM 287027; D, E. Paratype, female, CW 3.2 mm, CL 2.2 mm, MBM 287026; A. Carapace; B. Maxilliped 3; C. Cheliped; D. Pereopod 5; E. Female pleon; F. Male pleon; G. right G 1, ventral view; H. Same, dorsal view; I. Right G 1, distal part, ventral view; J. Same, dorsal view; K. Right G 2, ventral view; L. Same, dorsal view. Scale bar: 1 mm (A, C, D); 0.5 mm (B, E, F, G, H); 0.1 mm (I – L)." figureDoi="10.3897/zse.100.117859.figure3" httpUri="https://binary.pensoft.net/fig/1088096">3 D</figureCitation>
|
||
) (ambulatory legs very short and stout in
|
||
<taxonomicName id="0F7576AFB1BA4F2EF7F3798C0919A9B5" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Paractaeopsis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D5FFAFE02F36E1B805D593057857A701" italics="true">Paractaeopsis</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="84C57A5E3E7D1AFFA776FDD1A2B68096" author="Takeda and Miyake" firstAuthor="Takeda" issue="9" journalOrPublisher="OHMU, Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University" pagination="183-189" refId="B45" refString="Takeda M, Miyake S (1968) Two new xanthid crabs inhabiting coral reefs of the Ryukyu Islands. OHMU, Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University 1 (9): 183–189. [pl. 8]" title="Two new xanthid crabs inhabiting coral reefs of the Ryukyu Islands." volume="1" year="1968">Takeda and Miyake 1968</bibRefCitation>
|
||
: fig. 1 c).
|
||
</paragraph>
|
||
<paragraph id="4CF73810B59D51A2057C7C7791D7FE7B">
|
||
Given the above comparisons, the current species cannot be placed within any known genera, necessitating the establishment of a new genus. The main morphological characteristics comparing
|
||
<taxonomicName id="E8F7B36F41B6C2389D1E1190C720FF89" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="0B9E96DDC60EB3C1CCC83C9CB118D219" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EC46514EA22A13793FA4CD7A3837DB3E" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
with closely related genera are listed in Table
|
||
<tableCitation id="1F2F14373BB3075A545103C63FDAAE24" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of the characters of Gothus gen. nov. and six related genera included in the subfamily Euxanthinae." httpUri="http://table.plazi.org/id/610B39C88C5F69F61DF06E8AC3EE2C92" tableUuid="610B39C88C5F69F61DF06E8AC3EE2C92">2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="610B39C88C5F69F61DF06E8AC3EE2C92" ID-Table-UUID="610B39C88C5F69F61DF06E8AC3EE2C92" httpUri="http://table.plazi.org/id/610B39C88C5F69F61DF06E8AC3EE2C92" startId="T2">
|
||
<paragraph id="519E8CFA2054762C31A07E84138DE07D">
|
||
<label id="E73042B43E37EE134CB55136AC1D2883">Table 2.</label>
|
||
</paragraph>
|
||
<paragraph id="AD053962564A89BCC5A6C3EF4E967F95">
|
||
Comparison of the characters of
|
||
<taxonomicName id="E6F5867BFED067003ED2C3A8FD67F6E4" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="427DA2F8E1DD618C116624A9A4A3E7ED" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D28E18FA13AE9D19BCCBFD026B562163" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
and six related genera included in the subfamily
|
||
<taxonomicName id="D661AE91F4B2A54C873C8FAFB7862F47" authorityName="Alcock" authorityYear="1898" class="Malacostraca" family="Xanthidae" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="subFamily" subFamily="Euxanthinae">Euxanthinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="14FE645A28675D51C9A5537398B64550">
|
||
<table id="TID0EEMCI" rules="all">
|
||
<tbody id="5C22CA682CC637F2A0B925F9A6089DED">
|
||
<tr id="24056743BE84059E297B8795C7CAF626">
|
||
<th id="7EB3EBA671FC3DC49BCD609EE4A427EA" colspan="1" rowspan="1">Character</th>
|
||
<th id="B931541620F1F549B69CB40113402A79" colspan="1" rowspan="1">
|
||
<taxonomicName id="72F47F95105A201887792939700BDCA4" authorityName="Yuan & Jiang & Sha" authorityYear="2024" class="Malacostraca" family="Xanthidae" genus="Gothus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="54843B961B8CD671002D5969CCE65F6B" italics="true">Gothus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DE4BBF5C925D1FAD6E10B620030B59FC" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<emphasis id="28583CD04458A44301C932402BB53E1B" italics="true">
|
||
<th id="6B721DEDAA12EDC322F95AEBFEBC93F1" colspan="1" rowspan="1">
|
||
<taxonomicName id="CCD033339376780E923220F82957350C" authorityName="Mendoza & PKL Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">Rizalthus</taxonomicName>
|
||
</th>
|
||
<th id="49223C94E486D70622812932486D6D0D" colspan="1" rowspan="1">
|
||
<taxonomicName id="737BF6C959B90AE6D0267E2365D1A423" authorityName="Mendoza & Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">Visayax</taxonomicName>
|
||
</th>
|
||
<th id="2AAE460265950B485EB5A5F4C75ADF7D" colspan="1" rowspan="1">
|
||
<taxonomicName id="2EB220375D87A751F393502A4EDCBAD6" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">Euxanthus</taxonomicName>
|
||
</th>
|
||
<th id="34C0B34A3B40DAFD6326F9ECACD3579C" colspan="1" rowspan="1">
|
||
<taxonomicName id="133742C3B762E1B31B2038F194CEDF1A" authorityName="Rathbun" authorityYear="1897" class="Malacostraca" family="Xanthidae" genus="Hypocolpus" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">Hypocolpus</taxonomicName>
|
||
</th>
|
||
<th id="56ED19073C8D66561A193A59A76BFCB7" colspan="1" rowspan="1">
|
||
<taxonomicName id="861321AA3B5C8871AA8B99F98AB96A7A" authorityName="Kossmann" authorityYear="1877" class="Malacostraca" family="Xanthidae" genus="Psaumis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">Psaumis</taxonomicName>
|
||
</th>
|
||
<th id="7A62B41C5B0AF33ED1E8E5A00CCA4789" colspan="1" rowspan="1">
|
||
<taxonomicName id="6B5EAE8534CC71790283A6644B5C96ED" authorityName="Serene" authorityYear="1984" class="Malacostraca" family="Xanthidae" genus="Paractaeopsis" kingdom="Animalia" order="Decapoda" phylum="Arthropoda" rank="genus">Paractaeopsis</taxonomicName>
|
||
</th>
|
||
</emphasis>
|
||
</tr>
|
||
<tr id="349F216F5CA95FFE4244381E3435569E">
|
||
<td id="944364ACA2D671F24D5C5A3F66065FED" colspan="1" rowspan="1">carapace dorsal surface</td>
|
||
<td id="0CD56403A8B3222533066AD374597800" colspan="1" rowspan="1">with round granules, with clustered long setae or scattered short setae</td>
|
||
<td id="BD76EDD79C703E312F66A4408B984942" colspan="1" rowspan="1">with large granules, surrounded by short setae basally</td>
|
||
<td id="17CDA3731841297BAAE789A712375519" colspan="1" rowspan="1">with distinct or faint reticulate pattern of fused granules</td>
|
||
<td id="4AC1AAB76066827BCFC41B1A159A992A" colspan="1" rowspan="1">relatively smooth, without well-developed granules.</td>
|
||
<td id="A729384BF8E2233EAD87D3D428E406EA" colspan="1" rowspan="1">with granules and setae</td>
|
||
<td id="87440D2A1A0B20972234FCAA59A5B29F" colspan="1" rowspan="1">densely covered with granules</td>
|
||
<td id="1294BBBB97023BE0534A7D811FFE3318" colspan="1" rowspan="1">with pearly granules, scattered long tubular setae</td>
|
||
</tr>
|
||
<tr id="0A17B60CF510F0F259606E1C718568EC">
|
||
<td id="43A53651D6E659BA13E64F812A89F2F6" colspan="1" rowspan="1">carapace anterolateral margin</td>
|
||
<td id="3D8779B356CFC7111C339F284E0888CC" colspan="1" rowspan="1">with four anterolateral teeth, first tooth flattened, posterior three teeth well-developed</td>
|
||
<td id="CBF8C40A80335BC644C9ED403FA1A376" colspan="1" rowspan="1">without clearly defined anterolateral teeth</td>
|
||
<td id="5BE5970B1040BBED604FC209EC5CFD43" colspan="1" rowspan="1">absence of developed anterolateral teeth</td>
|
||
<td id="2D32B993A45848453B7D830C825E510D" colspan="1" rowspan="1">with four to six anterolateral teeth</td>
|
||
<td id="D19DCEA0286C258AEA4CBBF57B03F5F7" colspan="1" rowspan="1">anterolateral teeth underdeveloped</td>
|
||
<td id="2517A1D0C13E077B0C57CE193EDAAC6F" colspan="1" rowspan="1">with very flat anterolateral teeth, only fourth tooth prominent</td>
|
||
<td id="1930B2BB7D3B803B98CA553D59B65708" colspan="1" rowspan="1">with four developed anterolateral teeth</td>
|
||
</tr>
|
||
<tr id="EE702BDEAFB121B389C02C8C392E3A39">
|
||
<td id="A0AD1687AEF3D363C4597F19285D91DB" colspan="1" rowspan="1">carapace posterolateral margin</td>
|
||
<td id="47EA9990CE65FBF55907F43C9C06CD2F" colspan="1" rowspan="1">non-concave</td>
|
||
<td id="E123284AF258E459300A47B68703AF2A" colspan="1" rowspan="1">concave</td>
|
||
<td id="294FC5F6F46F8D139592EF5C31032FB0" colspan="1" rowspan="1">concave</td>
|
||
<td id="1107E24189EA4C703B313E7DCBB1EBC9" colspan="1" rowspan="1">concave</td>
|
||
<td id="7D68F3E7EF09B1DEB0200FAFB3404B45" colspan="1" rowspan="1">concave</td>
|
||
<td id="8092B11C7949AF5E0CDBD6C232EA747D" colspan="1" rowspan="1">concave</td>
|
||
<td id="8E180323BE63F1495C83A94E24CB910D" colspan="1" rowspan="1">non-concave</td>
|
||
</tr>
|
||
<tr id="8020FD0AAC96F54FB0B8577304889E29">
|
||
<td id="191F01067309BC86059C695C7790E4C0" colspan="1" rowspan="1">front</td>
|
||
<td id="D9E123D27AD40DAC179AB0A805EF5C32" colspan="1" rowspan="1">not protruding, divided by a V-shaped notch</td>
|
||
<td id="18770F010447403E1A15558FDE20FBA2" colspan="1" rowspan="1">not protruding, divided by a V-shaped notch</td>
|
||
<td id="CC0104E6F19FF471CC7ECC998A19E64C" colspan="1" rowspan="1">not protruding, divided by a V-shaped notch</td>
|
||
<td id="4853F48DCD5ECE7DB89A2E1A8516BF76" colspan="1" rowspan="1">protruding, divided by a narrow fissure</td>
|
||
<td id="38C0C79F5153BE3864DEEA63C43D0EB4" colspan="1" rowspan="1">protruding, divided by a narrow fissure</td>
|
||
<td id="C0AEAACA3137EDDC35B9F0431EE488B8" colspan="1" rowspan="1">not protruding, divided by a narrow fissure</td>
|
||
<td id="927DC203F10D19DEC4311F62BFDE505C" colspan="1" rowspan="1">protruding, divided by a V-shaped notch</td>
|
||
</tr>
|
||
<tr id="D8A12762349DD89DD25E161BEC7CDD92">
|
||
<td id="D249F35D38605C91EED25981B0390A5F" colspan="1" rowspan="1">epistome</td>
|
||
<td id="BCF7314724BCB4D4EB95022C28F9BBA9" colspan="1" rowspan="1">central part protruding</td>
|
||
<td id="22F1EF9938A406A3C2DC8F8EBCEC4D20" colspan="1" rowspan="1">central part not protruding</td>
|
||
<td id="0BBC5CA1E45596342793AE0F80F224D9" colspan="1" rowspan="1">central part not protruding or slightly protruding</td>
|
||
<td id="F0FE5569BE4A1A47755DE1190E3DBC1E" colspan="1" rowspan="1">central part protruding</td>
|
||
<td id="9E8B9EEB43CE97C9C44671F6D36361D0" colspan="1" rowspan="1">central part protruding</td>
|
||
<td id="749ADD13D6839C7A4549C08D6860D2B5" colspan="1" rowspan="1">central part protruding</td>
|
||
<td id="A46EA6B86C8453C5230911EAB9B9FE8E" colspan="1" rowspan="1">unknown</td>
|
||
</tr>
|
||
<tr id="CD0014BBFA54C0FD6FA85A9530B90E03">
|
||
<td id="F921505C4DF73F47330CF0E6B70FD72E" colspan="1" rowspan="1">subhepatic cavity</td>
|
||
<td id="74933E0470692A49BF39A129B659D135" colspan="1" rowspan="1">absent</td>
|
||
<td id="9548096BF5F49CE3E31166ACA6C81B59" colspan="1" rowspan="1">absent</td>
|
||
<td id="48A860EE6A78349758E052E4E6A7A986" colspan="1" rowspan="1">absent</td>
|
||
<td id="9568B4BF5F82A811CF1EB2343B419FAC" colspan="1" rowspan="1">absent</td>
|
||
<td id="D44FA34DFF4B70408FD62800A5FBCA13" colspan="1" rowspan="1">present</td>
|
||
<td id="C5CBAAFE694ACCEF4E3F288EC56CD5C8" colspan="1" rowspan="1">absent</td>
|
||
<td id="C5169556106632EAFA8297577D8202DF" colspan="1" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="42D7C758FA69A515BD76038761994F2D">
|
||
<td id="246A2DCF8452C7B7D14AD1B7D7BAF628" colspan="1" rowspan="1">etched depressions</td>
|
||
<td id="877843A84852F3B9DC9D3A03ECBE767F" colspan="1" rowspan="1">absent</td>
|
||
<td id="0D8A68B50A8A7B69123C9A5D8731C055" colspan="1" rowspan="1">present on thoracic sternum</td>
|
||
<td id="284344CFD4F3766D5914EB2226D85E08" colspan="1" rowspan="1">present on chelipeds, ambulatory legs, carapace, and thoracic sternum</td>
|
||
<td id="BE49037F14F445DCB43D152A2302EBE4" colspan="1" rowspan="1">absent</td>
|
||
<td id="8B4096854CB629BA0593FE1F88B38EC3" colspan="1" rowspan="1">present on thoracic sternum</td>
|
||
<td id="CBAEC6695C4125AEE899068A8CBD5BF4" colspan="1" rowspan="1">present on chelipeds, ambulatory legs and carapace</td>
|
||
<td id="0A270FD694E7483FB90E1329F97B26D2" colspan="1" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="6582F5B5BBA3FF544B1F3393F457D61D">
|
||
<td id="E7C67707388218A119279A095DD8E721" colspan="1" rowspan="1">male telson</td>
|
||
<td id="270936DE0FF8F016E3A3FC6B04FEEB73" colspan="1" rowspan="1">broad, truncated oval</td>
|
||
<td id="781875CF6D8751BDD19751D7D5B24B46" colspan="1" rowspan="1">smaller, triangular</td>
|
||
<td id="FEF79FBD14FC89A4E0EFC1599951145B" colspan="1" rowspan="1">smaller, semi-circular</td>
|
||
<td id="3C2294B53786614FD90CA3AA9987B624" colspan="1" rowspan="1">smaller, triangular</td>
|
||
<td id="4F9F889B01480B47551D24C7376F4B71" colspan="1" rowspan="1">smaller, triangular</td>
|
||
<td id="659D5DA6F266FEE26DE15287527DA181" colspan="1" rowspan="1">smaller, semi-circular</td>
|
||
<td id="7AEE274C71F6EBE4ECAD1038E52F8F1C" colspan="1" rowspan="1">unknown</td>
|
||
</tr>
|
||
<tr id="FFCFF3CC93115268AABAD8CEAC30521E">
|
||
<td id="8C056B605367DE13549CE100ABB22865" colspan="1" rowspan="1">cheliped carpus</td>
|
||
<td id="744D5860AFA2B8114087258675B6B883" colspan="1" rowspan="1">robust, sometimes slightly expanded</td>
|
||
<td id="5BAA7AE424350A6BE02F296959EFC8D9" colspan="1" rowspan="1">strongly expanded and protruding</td>
|
||
<td id="F2D3601613DAAA3621FB9995DDB058E9" colspan="1" rowspan="1">robust</td>
|
||
<td id="9669688F05BFED5A5149910B61F63337" colspan="1" rowspan="1">robust</td>
|
||
<td id="5A984589898FDFD410D274DC5330FCA2" colspan="1" rowspan="1">robust</td>
|
||
<td id="FCB97177407FE62D3FE0772BEA8A0077" colspan="1" rowspan="1">slightly robust</td>
|
||
<td id="BB6BC8B1A6BED143AF0A6902C97BF251" colspan="1" rowspan="1">robust</td>
|
||
</tr>
|
||
<tr id="F494A5A97285116CC4FC559D7D7A7207">
|
||
<td id="DB00F31F8195E5108AFEAF108170C8A8" colspan="1" rowspan="1">ambulatory leg</td>
|
||
<td id="9A0840B4F57933574583995BFC6B95F3" colspan="1" rowspan="1">comparatively slender</td>
|
||
<td id="1E6F6608005AAA273C96DADA2DCDC64B" colspan="1" rowspan="1">comparatively slender</td>
|
||
<td id="E13BF53DA28037B8855CD0ACC419451F" colspan="1" rowspan="1">comparatively slender</td>
|
||
<td id="F9C2365C8AF033F99F78039368A7EC24" colspan="1" rowspan="1">comparatively slender</td>
|
||
<td id="AE2CC38E101184633C0FD8AAB2362F46" colspan="1" rowspan="1">comparatively slender</td>
|
||
<td id="A727517A44D3F47C8409BFC2777F132B" colspan="1" rowspan="1">comparatively slender</td>
|
||
<td id="0CA484841F0C387D0AADEFCBB810155B" colspan="1" rowspan="1">very short and stout</td>
|
||
</tr>
|
||
<tr id="C47CDA1BEBFB5877592BD7BD290D3027">
|
||
<td id="20800937E7DAB83D0080B29538DD9FC6" colspan="1" rowspan="1">male first gonopod</td>
|
||
<td id="532A8D2E5D48B6FE1D14371DA7587937" colspan="1" rowspan="1">slender, distal lobe prominent, spoon-shaped, curved inwards, with long setae on inner subdistal part</td>
|
||
<td id="906424D90A7B1E4268D4CA4C1D103EE7" colspan="1" rowspan="1">slender, distal lobe prominent, spoon-shaped, nearly straight, with long setae on inner subdistal part</td>
|
||
<td id="BF40BBBAC3E86D8E65588D7779D80AEB" colspan="1" rowspan="1">robust, distal lobe prominent, nearly straight, with long setae on inner subdistal part</td>
|
||
<td id="43B21F83545F8719ABCEF358BB1B3353" colspan="1" rowspan="1">slender, distal lobe non-protruding, inwardly curved and encircling, with short setae on inner subdistal part</td>
|
||
<td id="8EAB70ACC370F559DD862C15689F6B29" colspan="1" rowspan="1">slender, distal lobe non-protruding, inwardly curved and encircling, with long setae on inner subdistal part</td>
|
||
<td id="BDA531082ED3D11FFD4062FBBE88BFEB" colspan="1" rowspan="1">slender, distal lobe prominent, spoon-shaped, nearly straight, with long setae on inner subdistal part</td>
|
||
<td id="2F624DABE98CE715BFCC7325949F96FE" colspan="1" rowspan="1">robust, distal lobe prominent, nearly straight, with long setae on inner subdistal part</td>
|
||
</tr>
|
||
</tbody>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |