347 lines
33 KiB
XML
347 lines
33 KiB
XML
<document ID-DOI="10.5252/zoosystema2023v45a3" ID-GBIF-Dataset="bc090a4d-4bce-42e0-bf6b-fb4bd45ca811" ID-ISSN="1638-9387" ID-Zenodo-Dep="7681295" ID-ZooBank="urn:lsid:zoobank.org:pub:62C2AC78-741C-4675-8C3A-67E359EB369E" checkinTime="1677503147187" checkinUser="felipe" docAuthor="Marchán, Daniel Fernández, Domínguez, Jorge, Hedde, Mickaël & Decaëns, Thibaud" docDate="2023" docId="F03D0809FFCA7452FEA7C4A9A1EEF874" docLanguage="en" docName="Zoosystema.45.3.107-128.pdf" docOrigin="Zoosystema 45 (3)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.3:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="3" docTitle="Scherotheca mausoleana Marchan, Decaens & Dominguez 2023, n. sp." docType="treatment" docVersion="7" lastPageNumber="120" masterDocId="0C047071FFC4745DFFE4C278A344FF98" masterDocTitle="The cradle of giants: insights into the origin of Scherotheca Bouché, 1972 (Lumbricidae, Crassiclitellata) with the descriptions of eight new species from Corsica, France" masterLastPageNumber="128" masterPageNumber="107" pageNumber="119" updateTime="1677510542406" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The cradle of giants: insights into the origin of Scherotheca Bouché, 1972 (Lumbricidae, Crassiclitellata) with the descriptions of eight new species from Corsica, France</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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||
<mods:namePart>Marchán, Daniel Fernández</mods:namePart>
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<mods:affiliation>CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) danifermch @ gmail. com</mods:affiliation>
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<mods:nameIdentifier type="email">danifermch@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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||
<mods:role>
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||
<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Domínguez, Jorge</mods:namePart>
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<mods:affiliation>Universidade de Vigo, Grupo de Ecoloxía Animal (GEA), E- 36310 Vigo (Spain)</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hedde, Mickaël</mods:namePart>
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<mods:affiliation>Eco & Sols, INRAE, IRD, CIRAD, SupAgro Montpellier, Campus SupAgro, Bâtiment 12, 2 place Viala, F- 34060 Montpellier cedex 2 (France) mickael. hedde @ inrae. fr</mods:affiliation>
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<mods:nameIdentifier type="email">hedde@inrae.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Decaëns, Thibaud</mods:namePart>
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<mods:affiliation>CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) thibaud. decaens @ cefe. cnrs. fr</mods:affiliation>
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<mods:nameIdentifier type="email">decaens@cefe.cnrs.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-08</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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||
<mods:number>45</mods:number>
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||
</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>107</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5252/zoosystema2023v45a3</mods:identifier>
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<mods:identifier type="GBIF-Dataset">bc090a4d-4bce-42e0-bf6b-fb4bd45ca811</mods:identifier>
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<mods:identifier type="ISSN">1638-9387</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7681295</mods:identifier>
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<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:62C2AC78-741C-4675-8C3A-67E359EB369E</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7681898" ID-Zenodo-Dep="7681898" LSID="urn:lsid:plazi:treatment:F03D0809FFCA7452FEA7C4A9A1EEF874" httpUri="http://treatment.plazi.org/id/F03D0809FFCA7452FEA7C4A9A1EEF874" lastPageId="15" lastPageNumber="120" pageId="14" pageNumber="119">
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<heading centered="true" fontSize="11" level="2" pageId="14" pageNumber="119" reason="3">
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<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="14" pageNumber="119" phylum="Annelida" rank="species" species="mausoleana" status="n. sp.">
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<emphasis bold="true" box="[323,584,1745,1772]" italics="true" pageId="14" pageNumber="119">Scherotheca mausoleana</emphasis>
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Marchán, Decäens & Domínguez
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</taxonomicName>
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||
<taxonomicNameLabel box="[611,670,1778,1804]" pageId="14" pageNumber="119" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="119" type="description">
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<paragraph blockId="14.[238,670,1745,1836]" box="[402,505,1810,1836]" pageId="14" pageNumber="119">
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(
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<figureCitation box="[410,496,1810,1836]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens & Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens & Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens & Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens & Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens & Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens & Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens & Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens & Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="14" pageNumber="119">Fig. 3H</figureCitation>
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)
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</paragraph>
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<paragraph blockId="14.[157,750,1870,1888]" box="[157,750,1870,1888]" pageId="14" pageNumber="119">
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<uri box="[157,750,1870,1888]" pageId="14" pageNumber="119">
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urn:lsid:zoobank.org:act:
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="119" type="materials_examined">
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<paragraph blockId="14.[132,776,1921,2024]" pageId="14" pageNumber="119">
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||
<materialsCitation collectingDate="2021-03-28" collectionCode="TYPE MATERIAL, MNHN" collectorName="T. Decaens & D. Fernandez Marchan" country="France" county="Tartagine" elevation="779" latitude="42.49389" location="Mausoleo, Tartagine" longLatPrecision="19" longitude="8.995833" municipality="Olmi- Cappella" pageId="14" pageNumber="119" specimenCode="DFM-0327" specimenCount="1" stateProvince="Haute-Corse" typeStatus="holotype">
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<collectionCode box="[132,277,1921,1944]" pageId="14" pageNumber="119">TYPE MATERIAL</collectionCode>
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. —
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||
<emphasis bold="true" box="[318,496,1921,1946]" pageId="14" pageNumber="119">
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||
<typeStatus box="[318,413,1921,1945]" pageId="14" pageNumber="119">Holotype</typeStatus>
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||
.
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||
<collectingCountry box="[422,496,1922,1946]" name="France" pageId="14" pageNumber="119">Corsica</collectingCountry>
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||
</emphasis>
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||
•
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||
<specimenCount box="[515,572,1921,1945]" pageId="14" pageNumber="119" type="adult">Adult</specimenCount>
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||
;
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||
<collectingRegion box="[580,707,1921,1945]" pageId="14" pageNumber="119">Haute-Corse</collectingRegion>
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||
,
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||
<collectingMunicipality pageId="14" pageNumber="119">Olmi-Cappella</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:F03D0809FFCA7452FEA7C4A9A1EEF874:7D4BEFC4FFCA7453FF0FC5E3A2FDF82B" box="[235,441,1947,1971]" country="France" county="Tartagine" latitude="42.49389" longLatPrecision="19" longitude="8.995833" municipality="Olmi- Cappella" name="Mausoleo, Tartagine" pageId="14" pageNumber="119" stateProvince="Haute-Corse">Mausoleo, Tartagine</location>
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;
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,
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<geoCoordinate box="[590,693,1947,1971]" degrees="8" direction="east" minutes="59" orientation="longitude" pageId="14" pageNumber="119" precision="15" seconds="45" value="8.995833">8°59’45”E</geoCoordinate>
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;
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<quantity box="[708,775,1948,1972]" metricMagnitude="2" metricUnit="m" metricValue="7.79" pageId="14" pageNumber="119" unit="m" value="779.0">779 m</quantity>
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a.s.l.
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</elevation>
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;
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<date box="[189,306,1974,1998]" pageId="14" pageNumber="119" value="2021-03-28">
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<collectingDate box="[189,306,1974,1998]" pageId="14" pageNumber="119" value="2021-03-28">28.III.2021</collectingDate>
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</date>
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;
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<collectorName box="[314,423,1974,1997]" pageId="14" pageNumber="119">T. Decaëns</collectorName>
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||
,
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||
<collectorName box="[432,657,1974,1998]" pageId="14" pageNumber="119">D. Fernández Marchán</collectorName>
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leg.; BOLD Sample ID:
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<specimenCode box="[251,369,2000,2024]" pageId="14" pageNumber="119">DFM-0327</specimenCode>
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||
;
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<collectionCode box="[379,465,2001,2024]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="119">MNHN</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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<materialsCitation collectingDate="2021-03-28" collectionCode="T, BOLD, MNHN" country="France" elevation="635" latitude="42.503887" location="Haute-Corse" longLatPrecision="19" longitude="9.003889" municipality="Mausoleo" pageId="14" pageNumber="119" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
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<emphasis bold="true" box="[813,909,1743,1767]" pageId="14" pageNumber="119">
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<typeStatus box="[813,909,1743,1767]" pageId="14" pageNumber="119">Paratypes</typeStatus>
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</emphasis>
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.
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<emphasis bold="true" box="[920,995,1744,1768]" pageId="14" pageNumber="119">
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||
<collectingCountry box="[920,995,1744,1768]" name="France" pageId="14" pageNumber="119">Corsica</collectingCountry>
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||
</emphasis>
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||
•
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||
<specimenCount box="[1015,1109,1743,1767]" pageId="14" pageNumber="119" type="juvenile">1 juvenile</specimenCount>
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specimen;
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,
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||
<collectingMunicipality box="[1354,1452,1743,1767]" pageId="14" pageNumber="119">Mausoleo</collectingMunicipality>
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;
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,
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<geoCoordinate box="[946,1035,1769,1793]" degrees="9" direction="east" minutes="0" orientation="longitude" pageId="14" pageNumber="119" precision="15" seconds="14" value="9.003889">9°0’14”E</geoCoordinate>
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;
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<quantity box="[1049,1115,1769,1793]" metricMagnitude="2" metricUnit="m" metricValue="6.35" pageId="14" pageNumber="119" unit="m" value="635.0">635 m</quantity>
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a.s.l.
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</elevation>
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;
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<date box="[1179,1296,1769,1793]" pageId="14" pageNumber="119" value="2021-03-28">
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<collectingDate box="[1179,1296,1769,1793]" pageId="14" pageNumber="119" value="2021-03-28">28.III.2021</collectingDate>
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</date>
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;
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<collectionCode box="[1306,1323,1770,1793]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="119">T</collectionCode>
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. Decaëns, D. Fernández Marchán leg.;
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||
<collectionCode box="[1056,1125,1796,1820]" pageId="14" pageNumber="119">BOLD</collectionCode>
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Sample ID: DFM-0516;
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<collectionCode box="[1372,1455,1796,1819]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="119">MNHN</collectionCode>
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</materialsCitation>
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•
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||
<materialsCitation collectingDate="2021-03-28" collectionCode="T, BOLD" country="France" county="Tartagine" elevation="779" latitude="42.49389" location="Haute-Corse" longLatPrecision="19" longitude="8.995833" municipality="Mausoleo" pageId="14" pageNumber="119" specimenCount="2" specimenCount-adult="2" typeStatus="paratype">
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specimens;
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,
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<location LSID="urn:lsid:plazi:treatment:F03D0809FFCA7452FEA7C4A9A1EEF874:7D4BEFC4FFCA7453FB78C566A67DF8AE" box="[1180,1337,1822,1846]" country="France" county="Tartagine" latitude="42.49389" longLatPrecision="19" longitude="8.995833" municipality="Mausoleo" name="Olmi-Cappella" pageId="14" pageNumber="119">Olmi-Cappella</location>
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,
|
||
<collectingMunicipality box="[1349,1451,1822,1846]" pageId="14" pageNumber="119">Mausoleo</collectingMunicipality>
|
||
,
|
||
<collectingCounty box="[815,911,1848,1872]" pageId="14" pageNumber="119">Tartagine</collectingCounty>
|
||
;
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<geoCoordinate box="[920,1043,1849,1873]" degrees="42" direction="north" minutes="29" orientation="latitude" pageId="14" pageNumber="119" precision="15" seconds="38" value="42.49389">42°29’38”N</geoCoordinate>
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,
|
||
<geoCoordinate box="[1055,1158,1848,1872]" degrees="8" direction="east" minutes="59" orientation="longitude" pageId="14" pageNumber="119" precision="15" seconds="45" value="8.995833">8°59’45”E</geoCoordinate>
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;
|
||
<elevation box="[1173,1293,1848,1873]" metricMagnitude="2" metricUnit="m" metricValue="7.79" pageId="14" pageNumber="119" unit="m" value="779.0">
|
||
<quantity box="[1173,1239,1849,1873]" metricMagnitude="2" metricUnit="m" metricValue="7.79" pageId="14" pageNumber="119" unit="m" value="779.0">779 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date box="[1306,1424,1849,1873]" pageId="14" pageNumber="119" value="2021-03-28">
|
||
<collectingDate box="[1306,1424,1849,1873]" pageId="14" pageNumber="119" value="2021-03-28">28.III.2021</collectingDate>
|
||
</date>
|
||
;
|
||
<collectionCode box="[1435,1452,1849,1872]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="119">T</collectionCode>
|
||
. Decaëns, D. Fernández Marchán leg.;
|
||
<collectionCode box="[1194,1263,1875,1899]" pageId="14" pageNumber="119">BOLD</collectionCode>
|
||
Sample ID: DFM-0326, DFM-0328; Eco&Sols
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation collectingDate="2021-03-28" collectionCode="BOLD, MNHN" country="France" county="Tartagine" elevation="779" latitude="42.49389" location="Haute-Corse" longLatPrecision="19" longitude="8.995833" municipality="Mausoleo" pageId="14" pageNumber="119" specimenCount="1" specimenCount-adult="1" typeStatus="paratype">
|
||
<specimenCount box="[1122,1191,1901,1925]" pageId="14" pageNumber="119" type="adult">1 adult</specimenCount>
|
||
specimen; same data as for preceding;
|
||
<collectionCode box="[920,989,1928,1952]" pageId="14" pageNumber="119">BOLD</collectionCode>
|
||
Sample ID: DFM-0329;
|
||
<collectionCode box="[1243,1329,1928,1951]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="119">MNHN</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="119" type="etymology">
|
||
<paragraph blockId="14.[813,1455,1975,2026]" pageId="14" pageNumber="119">
|
||
ETYMOLOGY. — The species name is derived from the municipality of Mausoléo where the
|
||
<typeStatus box="[1073,1115,2002,2026]" pageId="14" pageNumber="119">type</typeStatus>
|
||
population has been discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="15" pageNumber="120" type="diagnosis">
|
||
<paragraph blockId="15.[131,777,215,636]" pageId="15" pageNumber="120">
|
||
DIAGNOSIS. —
|
||
<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="mausoleana" status="n. sp.">
|
||
<emphasis box="[291,536,215,239]" italics="true" pageId="15" pageNumber="120">Scherotheca mausoleana</emphasis>
|
||
Marchán, Decäens & Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[263,318,242,265]" pageId="15" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
|
||
can be distinguished from non-Corsican species of
|
||
<taxonomicName authorityName="Bouche" authorityYear="1972" box="[203,312,268,292]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="genus">
|
||
<emphasis box="[203,312,268,292]" italics="true" pageId="15" pageNumber="120">Scherotheca</emphasis>
|
||
</taxonomicName>
|
||
by the shorter and more anterior clitellum. It resembles
|
||
<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" box="[236,662,295,319]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="qiui" status="n. sp.">
|
||
<emphasis box="[236,310,295,319]" italics="true" pageId="15" pageNumber="120">Sc. qiui</emphasis>
|
||
Marchán, Decäens & Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[674,730,295,318]" pageId="15" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" box="[134,574,321,345]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="litoralis" status="n. sp.">
|
||
<emphasis box="[134,237,321,345]" italics="true" pageId="15" pageNumber="120">Sc. litoralis</emphasis>
|
||
Marchán, Decäens & Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[583,636,322,345]" pageId="15" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
|
||
It differs from
|
||
<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" box="[132,537,348,372]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="qiui" status="n. sp.">
|
||
<emphasis box="[132,202,348,371]" italics="true" pageId="15" pageNumber="120">Sc. qiui</emphasis>
|
||
Marchán, Decäens & Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[546,599,348,371]" pageId="15" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
|
||
by the presence of body pigmentation, the number of seminal vesicles (4 pairs instead of 2 pairs) and by the longer
|
||
<emphasis box="[403,586,401,425]" italics="true" pageId="15" pageNumber="120">tubercula pubertatis</emphasis>
|
||
(30-34(35) instead 30-33(1/
|
||
<subScript attach="left" box="[218,227,438,454]" fontSize="7" pageId="15" pageNumber="120">n</subScript>
|
||
34)). It can further be distinguished from
|
||
<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="litoralis" status="n. sp.">
|
||
<emphasis box="[668,775,427,451]" italics="true" pageId="15" pageNumber="120">Sc. litoralis</emphasis>
|
||
Marchán, Decäens & Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[474,527,454,477]" pageId="15" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
|
||
by the position of genital papillae and the longer clitellum ((26)27-36(37) instead of 27-36) (
|
||
<tableCitation box="[139,221,506,531]" captionStart="TABLE" captionStartId="5.[131,142,219,237]" captionTargetPageId="5" captionText="TABLE 1. — Morphological characters of the newly described species of Corsican Scherotheca Bouché, 1972 and their closest relatives. Length in millimetres; weight in grams; 1ST D.P., position of first dorsal pore. Symbols: *, according to Pop (1947): composite between the different populations included in the original description; **, according to Bouché (1972)." httpUri="http://table.plazi.org/id/2CEBE997FFC17458FF67C2A3A2B6FE82" pageId="15" pageNumber="120" tableUuid="2CEBE997FFC17458FF67C2A3A2B6FE82">Tables 1</tableCitation>
|
||
;
|
||
<tableCitation box="[234,248,507,531]" captionStart="TABLE" captionStartId="7.[132,143,219,237]" captionTargetPageId="7" captionText="TABLE 2. — Reproduction-related morphological characters of the newly described species of Corsican Scherotheca Bouché, 1972 and their closest relatives. Abbreviations: Sperm. pores, position of spermathecal pores; S.V., position of seminal vesicles; T.P., position of tubercula pubertatis. Symbols: *, according to Pop (1947): composite between the different populations included in the original description; **, according to Bouché (1972)." httpUri="http://table.plazi.org/id/2CEBE997FFC3745AFF60C2A3A7D6FE82" pageId="15" pageNumber="120" tableUuid="2CEBE997FFC3745AFF60C2A3A7D6FE82">2</tableCitation>
|
||
;
|
||
<figureCitation box="[259,338,506,531]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens & Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens & Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens & Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens & Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens & Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens & Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens & Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens & Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="15" pageNumber="120">Fig. 3H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="15.[131,777,215,636]" pageId="15" pageNumber="120">
|
||
COI uncorrected average pairwise distances and topology of multilocus molecular phylogenetic trees support the status of
|
||
<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="mausoleana" status="n. sp.">
|
||
<emphasis italics="true" pageId="15" pageNumber="120">Sc. mausoleana</emphasis>
|
||
Marchán, Decäens & Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[555,609,586,609]" pageId="15" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
|
||
as separate from other morphologically similar species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="15" pageNumber="120" type="distribution">
|
||
<paragraph blockId="15.[130,777,669,799]" pageId="15" pageNumber="120">
|
||
DISTRIBUTION AND ECOLOGY. —
|
||
<taxonomicName authority="Marchan, Decaens & Dominguez" authorityName="Marchan, Decaens & Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="mausoleana" status="n. sp.">
|
||
<emphasis box="[440,574,669,693]" italics="true" pageId="15" pageNumber="120">Sc. mausoleana</emphasis>
|
||
Marchán, Decäens & Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[264,321,696,719]" pageId="15" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus box="[579,622,696,720]" pageId="15" pageNumber="120">type</typeStatus>
|
||
locality in the municipality of Mausoleo, Haute-Corse (
|
||
<figureCitation box="[555,635,722,746]" captionStart="FIG" captionStartId="14.[132,143,1613,1631]" captionTargetBox="[220,1354,216,1568]" captionTargetId="figure-223@14.[220,1354,216,1568]" captionTargetPageId="14" captionText="FIG. 4. — Distribution of the genus Scherotheca Bouché, 1972 in Corsica: A, localities documented in this work of the newly described species and other Scherotheca species sampled during this work; B, distribution of Corsican species of Scherotheca and Eumenescolex Qiu & Bouché, 1998 according to Bouché (1972), Qiu & Bouché (1998) and Szederjesi et al. (2021)." figureDoi="http://doi.org/10.5281/zenodo.7681307" httpUri="https://zenodo.org/record/7681307/files/figure.png" pageId="15" pageNumber="120">Fig. 4A</figureCitation>
|
||
), where it has been found in
|
||
<taxonomicName box="[280,399,748,772]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="120" phylum="Tracheophyta" rank="species" species="laricio">
|
||
<emphasis box="[280,399,748,772]" italics="true" pageId="15" pageNumber="120">Pinus laricio</emphasis>
|
||
</taxonomicName>
|
||
forest and Mediterranean chaparral at moderate elevation (
|
||
<quantity box="[331,445,775,799]" metricMagnitude="2" metricUnit="m" metricValue="7.07" metricValueMax="7.79" metricValueMin="6.35" pageId="15" pageNumber="120" unit="m" value="707.0" valueMax="779.0" valueMin="635.0">635-779 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="15" pageNumber="120" type="description">
|
||
<paragraph blockId="15.[130,777,854,1422]" box="[132,277,854,880]" pageId="15" pageNumber="120">DESCRIPTION</paragraph>
|
||
<paragraph blockId="15.[130,777,854,1422]" box="[132,346,885,911]" pageId="15" pageNumber="120">
|
||
<emphasis box="[132,346,885,911]" italics="true" pageId="15" pageNumber="120">External morphology</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="15.[130,777,854,1422]" pageId="15" pageNumber="120">
|
||
Body pigmentation brown. Beige with antero-posterior brownish gradients and white-beige clitellum in fixed specimens (
|
||
<figureCitation box="[209,304,980,1007]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens & Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens & Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens & Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens & Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens & Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens & Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens & Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens & Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="15" pageNumber="120">Fig. 3H</figureCitation>
|
||
). Average length unknown (no complete specimens available) body cylindrical in cross-section; average number of segments unknown (no complete specimens). Average weight (fixed specimens) unknown (no complete specimens). Prostomium epilobous, closed. Longitudinal furrows in segments 1 and 2. First dorsal pore unknown. Nephridial pores “en solfège” (irregularly distributed). Spermathecal pores in intersegmental furrows 12/13 and 13/
|
||
<quantity box="[172,230,1236,1262]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="15" pageNumber="120" unit="in" value="14.0">14 in</quantity>
|
||
setal line
|
||
<emphasis box="[335,345,1236,1261]" italics="true" pageId="15" pageNumber="120">c</emphasis>
|
||
. Male pores in segment 15, surrounded by a moderately-developed porophore. Female pores in segment 14. Clitellum saddle-shaped in segments (26)27- 36(37).
|
||
<emphasis box="[222,440,1331,1357]" italics="true" pageId="15" pageNumber="120">Tubercula pubertatis</emphasis>
|
||
in segments 30-34(35).
|
||
<emphasis italics="true" pageId="15" pageNumber="120">Chaetae</emphasis>
|
||
small and closely paired. Genital papillae in segments 11-14,16,28-30,35,36.
|
||
</paragraph>
|
||
<paragraph blockId="15.[130,777,1459,1900]" box="[132,312,1459,1485]" pageId="15" pageNumber="120">
|
||
<emphasis box="[132,312,1459,1485]" italics="true" pageId="15" pageNumber="120">Internal anatomy</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="15.[130,777,1459,1900]" pageId="15" pageNumber="120">
|
||
<collectingRegion box="[132,193,1491,1517]" country="Spain" name="Ceuta" pageId="15" pageNumber="120">Septa</collectingRegion>
|
||
5/6-10/11 thickened and muscular. Hearts in segments 6-11, oesophageal. Calciferous glands in segments 10-14, with diverticula in segment 10. Crop in segments 15-16, gizzard in segments 17-19. Typhlosole pinnate.Male sexual system holandric, testes and funnels (not enclosed in testes sacs, but with sperm present) located ventrally in segments 10 and 11. Four pairs of reniform seminal vesicles in segments 9, 10, 11 and 12, with the latter two pairs being larger. Ovaries and female funnels in segment 13, ovarian receptacles (ovisacs) in segment 14. Two pairs of small globular spermathecae in segments 12 and 13 (intersegments 12/13, 13/14). Nephridial bladders U-shaped, reclinate in segment 30.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="15" pageNumber="120" type="discussion">
|
||
<paragraph blockId="15.[132,775,1938,2028]" box="[132,222,1938,1964]" pageId="15" pageNumber="120">REMARK</paragraph>
|
||
<paragraph blockId="15.[132,775,1938,2028]" pageId="15" pageNumber="120">
|
||
This species corresponds to
|
||
<taxonomicName authorityName="* (Pop" authorityYear="1947" box="[434,565,1969,1995]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="15" pageNumber="120" phylum="Annelida" rank="species" species="corsicana">
|
||
<emphasis box="[434,565,1969,1995]" italics="true" pageId="15" pageNumber="120">Sc. corsicana</emphasis>
|
||
</taxonomicName>
|
||
L
|
||
<quantity box="[590,632,1969,1996]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="15" pageNumber="120" unit="in" value="5.0">5 in</quantity>
|
||
the checklist of Corsican earthworms of
|
||
<bibRefCitation author="MARCHAN D. F. & GERARD S. & HEDDE M. & ROUGERIE R. & DECAENS T." box="[427,676,2001,2028]" pageId="15" pageNumber="120" pagination="439 - 461" refId="ref16506" refString="MARCHAN D. F., GERARD S., HEDDE M., ROUGERIE R. & DECAENS T. 2022 a. - An updated checklist and a DNA barcode library for the earthworms (Crassiclitellata, Oligochaeta) of Corsica, France. Zoosystema 44 (17): 439 - 461. https: // doi. org / 10.5252 / zoosystema 2022 v 44 a 17. http: // zoosystema. com / 44 / 17" type="journal article" year="2022">
|
||
Marchán
|
||
<emphasis box="[531,583,2001,2027]" italics="true" pageId="15" pageNumber="120">et al.</emphasis>
|
||
(2022a)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |