406 lines
58 KiB
XML
406 lines
58 KiB
XML
<document id="7B7AEAFE4DF790DE7107B6EC2F59B8DD" ID-CLB-Dataset="55780" ID-DOI="10.11646/zootaxa.5129.4.3" ID-GBIF-Dataset="6d75f66c-89f6-45b4-b9c3-e8e376c4dbb6" ID-ISSN="1175-5326" ID-Zenodo-Dep="6504000" ID-ZooBank="B01CE641-CB32-4A28-B7D3-84D534BFE3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651218226472" checkinUser="plazi" docAuthor="Denisenko, Nina V." docDate="2022" docId="93797F2BFFB55E0678E04A7AD125B3D6" docLanguage="en" docName="zootaxa.5129.4.3.pdf" docOrigin="Zootaxa 5129 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Reteporella obscura Denisenko 2022, n. sp." docType="treatment" docVersion="8" lastPageNumber="535" masterDocId="6F400753FFB75E0378774F6BD550B32C" masterDocTitle="New species and new records of Reteporella (Bryozoa: Cheilostomatida) from Greenland waters" masterLastPageNumber="542" masterPageNumber="530" pageNumber="532" updateTime="1699329445803" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F869003FE7D08E1230E85D25FBE1C093">New species and new records of Reteporella (Bryozoa: Cheilostomatida) from Greenland waters</mods:title>
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<mods:name id="F47FBF5AC1C1B52F6C8FA2FB2B216744" type="personal">
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<mods:namePart id="9AA20CB4AD1BEA952D614C84008B7CF8">Denisenko, Nina V.</mods:namePart>
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<mods:title id="1496A577FFC8F7A58A85F9BC13A17766">Zootaxa</mods:title>
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<mods:part id="F085BEA4A4BE395935D0B9ACB9248BCD">
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<mods:date id="56AF04E34CE8536E3CFAE846A63B65B6">2022</mods:date>
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<mods:number id="6EC8F4C00B16D584DDA92A209243CB47">2022-04-29</mods:number>
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<mods:number id="B9B7A5A810BF024F7EE0F8F345439F98">5129</mods:number>
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<mods:identifier id="F651175C10889799594A130911303208" type="DOI">10.11646/zootaxa.5129.4.3</mods:identifier>
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<treatment id="93797F2BFFB55E0678E04A7AD125B3D6" ID-DOI="http://doi.org/10.5281/zenodo.6504110" ID-GBIF-Taxon="194877490" ID-Zenodo-Dep="6504110" LSID="urn:lsid:plazi:treatment:93797F2BFFB55E0678E04A7AD125B3D6" httpUri="http://treatment.plazi.org/id/93797F2BFFB55E0678E04A7AD125B3D6" lastPageId="5" lastPageNumber="535" pageId="2" pageNumber="532">
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<subSubSection id="53CA9DB6FFB55E0178E04A7AD483B607" box="[151,467,1297,1324]" pageId="2" pageNumber="532" type="nomenclature">
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<paragraph id="1B6FCE3DFFB55E0178E04A7AD483B607" blockId="2.[151,467,1297,1359]" box="[151,467,1297,1324]" pageId="2" pageNumber="532">
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<heading id="40277951FFB55E0178E04A7AD483B607" bold="true" box="[151,467,1297,1324]" fontSize="11" level="1" pageId="2" pageNumber="532" reason="1">
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<emphasis id="29A4122FFFB55E0178E04A7AD483B607" bold="true" box="[151,467,1297,1324]" pageId="2" pageNumber="532">
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<taxonomicName id="DCD0B5BEFFB55E0178E04A7AD4D9B600" authority="Denisenko, 2022" authorityName="Denisenko" authorityYear="2022" box="[151,393,1297,1324]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="532" phylum="Bryozoa" rank="species" species="obscura" status="sp. nov.">
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<emphasis id="29A4122FFFB55E0178E04A7AD4D9B600" bold="true" box="[151,393,1297,1324]" italics="true" pageId="2" pageNumber="532">Reteporella obscura</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3297AF54FFB55E0179E64A7AD483B607" box="[401,467,1297,1323]" pageId="2" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="53CA9DB6FFB55E0178E04A5ED41CB663" box="[151,332,1333,1359]" pageId="2" pageNumber="532" type="description">
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<paragraph id="1B6FCE3DFFB55E0178E04A5ED41CB663" blockId="2.[151,467,1297,1359]" box="[151,332,1333,1359]" pageId="2" pageNumber="532">
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(
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<figureCitation id="83EBD2B8FFB55E0178D74A5ED5B5B663" box="[160,229,1333,1359]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="2" pageNumber="532">Fig. 2</figureCitation>
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;
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<tableCitation id="5652FB86FFB55E0178874A5ED411B663" box="[240,321,1333,1359]" captionStart="TABLE 1" captionStartId="3.[151,239,548,572]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in mm) of Reteporella obscura n. sp. (holotype). SD, standard deviation; N, number of measurements." httpUri="http://table.plazi.org/id/4FAF9EB5FFB45E0078E04D4FD467B14C" pageId="2" pageNumber="532" tableUuid="4FAF9EB5FFB45E0078E04D4FD467B14C">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="53CA9DB6FFB55E0178E04A12D01DB6D7" pageId="2" pageNumber="532" type="materials_examined">
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<paragraph id="1B6FCE3DFFB55E0178E04A12D0D9B69F" blockId="2.[151,1437,1401,2036]" pageId="2" pageNumber="532">
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<emphasis id="29A4122FFFB55E0178E04A12D42CB6BF" bold="true" box="[151,380,1401,1427]" pageId="2" pageNumber="532">Material examined.</emphasis>
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<materialsCitation id="ABB8C460FFB55E0179F44A12D0D9B69F" ID-GBIF-Occurrence="3757952306" collectingDate="2016-07" collectionCode="MT" elevation="277" latitude="64.52787" longLatPrecision="1" longitude="-54.7777" pageId="2" pageNumber="532" specimenCode="ZIN1, T 3.934" specimenCount="1" typeStatus="holotype">
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<typeStatus id="C46B709FFFB55E0179F44A12D4BDB6BF" box="[387,493,1401,1427]" pageId="2" pageNumber="532" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="4B766646FFB55E0179834A11D77EB6B8" box="[500,558,1401,1428]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="2" pageNumber="532" type="Museum">ZIN1</specimenCode>
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/50742, one colony;Stn 2016_PA_2_151 (
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, ˗
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<geoCoordinate id="7EE4A8FAFFB55E017CCB4A12D5E3B698" degrees="54.7776999791463" direction="west" orientation="longitude" pageId="2" pageNumber="532" precision="1" value="-54.7777">54.7776999791463° W</geoCoordinate>
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;
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<quantity id="DC2863D8FFB55E0178C84AF2D457B698" box="[191,263,1433,1460]" metricMagnitude="2" metricUnit="m" metricValue="2.77" pageId="2" pageNumber="532" unit="m" value="277.0">
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<elevation id="90FD290EFFB55E0178C84AF2D457B698" box="[191,263,1433,1460]" metricMagnitude="2" metricUnit="m" metricValue="2.77" pageId="2" pageNumber="532" unit="m" value="277.0">277 m</elevation>
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</quantity>
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;
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<specimenCode id="4B766646FFB55E0179654AF1D43DB69F" box="[274,365,1433,1460]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="532">T 3.934</specimenCode>
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°C); collected by BEAMTRAWL,
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<collectionCode id="7DC156F8FFB55E017A944AF1D65CB698" box="[739,780,1434,1460]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="2" pageNumber="532">MT</collectionCode>
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<emphasis id="29A4122FFFB55E017B644AF1D620B69F" box="[787,880,1434,1459]" italics="true" pageId="2" pageNumber="532">Paamiut</emphasis>
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, fish-shrimp trawl assessment survey.
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<date id="6F6EE8FDFFB55E017D614AF2D0D5B69F" box="[1302,1413,1433,1459]" pageId="2" pageNumber="532" value="2016-07">
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<collectingDate id="7F2A1115FFB55E017D614AF2D0D5B69F" box="[1302,1413,1433,1459]" pageId="2" pageNumber="532" value="2016-07">July 2016</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="1B6FCE3DFFB55E0178B04AD5D01DB6D7" blockId="2.[151,1437,1401,2036]" pageId="2" pageNumber="532">
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<materialsCitation id="ABB8C460FFB55E0178B04AD5D42BB6F4" ID-GBIF-Occurrence="3757952309" box="[199,379,1469,1496]" collectionCode="ZIN" pageId="2" pageNumber="532" specimenCode="ZIN 2" specimenCount="1" typeStatus="paratype">
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<typeStatus id="C46B709FFFB55E0178B04AD5D47DB6F4" box="[199,301,1470,1496]" pageId="2" pageNumber="532" type="paratype">Paratype</typeStatus>
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:
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<specimenCode id="4B766646FFB55E01794F4AD5D42BB6F4" box="[312,379,1469,1496]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="2" pageNumber="532" type="Museum">ZIN 2</specimenCode>
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</materialsCitation>
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||
/
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<materialsCitation id="ABB8C460FFB55E0179F24AD6D46CB6D0" ID-GBIF-Occurrence="3757952302" collectionCode="ZIN" elevation="165" latitude="64.35197" longLatPrecision="1" longitude="-53.87652" pageId="2" pageNumber="532" specimenCode="T 1.575" specimenCount="1" typeStatus="paratype">
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50747; colony fragment; Stn 2016_PA_1_16 (
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||
<geoCoordinate id="7EE4A8FAFFB55E017BFD4AD6D119B6F4" box="[906,1097,1469,1496]" degrees="64.35196667" direction="north" orientation="latitude" pageId="2" pageNumber="532" precision="1" value="64.35197">64.35196667° N</geoCoordinate>
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; ˗
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<geoCoordinate id="7EE4A8FAFFB55E017C294AD6D037B6F4" box="[1118,1383,1469,1496]" degrees="53.8765166600545" direction="west" orientation="longitude" pageId="2" pageNumber="532" precision="1" value="-53.87652">53.8765166600545° W</geoCoordinate>
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;
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<quantity id="DC2863D8FFB55E017D054AD6D5E0B6D0" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="532" unit="m" value="165.0">
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<elevation id="90FD290EFFB55E017D054AD6D5E0B6D0" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="532" unit="m" value="165.0">165 m</elevation>
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</quantity>
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;
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<specimenCode id="4B766646FFB55E0178CC4A89D447B6D0" box="[187,279,1505,1532]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="532">T 1.575</specimenCode>
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°C)
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</materialsCitation>
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; collected by BEAMTRAWL,
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<materialsCitation id="ABB8C460FFB55E017AE04A89D019B6D7" ID-GBIF-Occurrence="3757952303" box="[663,1353,1505,1532]" collectingDate="2016-07" collectionCode="MT" pageId="2" pageNumber="532" specimenCount="1" typeStatus="paratype">
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<collectionCode id="7DC156F8FFB55E017AE04A89D790B6D0" box="[663,704,1506,1532]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="2" pageNumber="532">MT</collectionCode>
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||
<emphasis id="29A4122FFFB55E017ABF4A89D677B6D7" box="[712,807,1506,1531]" italics="true" pageId="2" pageNumber="532">Paamiut</emphasis>
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||
, fish-shrimp trawl assessment survey.
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<date id="6F6EE8FDFFB55E017CA04A8AD019B6D7" box="[1239,1353,1505,1531]" pageId="2" pageNumber="532" value="2016-07">
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<collectingDate id="7F2A1115FFB55E017CA04A8AD019B6D7" box="[1239,1353,1505,1531]" pageId="2" pageNumber="532" value="2016-07">July 2016</collectingDate>
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||
</date>
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||
</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="53CA9DB6FFB55E0178B0496ED700B4AB" pageId="2" pageNumber="532" type="diagnosis">
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<paragraph id="1B6FCE3DFFB55E0178B0496ED700B4AB" blockId="2.[151,1437,1401,2036]" pageId="2" pageNumber="532">
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||
<emphasis id="29A4122FFFB55E0178B0496ED412B533" bold="true" box="[199,322,1541,1567]" pageId="2" pageNumber="532">Diagnosis.</emphasis>
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Colony reticulate, broadly fan-shaped; fenestrulae large; different levels of granulation on frontal and abfrontal surfaces. Trabeculae of variable length, with 2‒3 series of alternating zooids, doubling at bifurcations. Zooids flat, with 3‒5 large marginal pores; marginal zooids near fenestrulae with 5‒6 marginal pores. Orifice semicircular, approximately as wide as long with rounded condyles and slightly prominent proximal edge; distal rim smooth with a well-developed row of denticles in its lower part. Spines paired, one on each side of orifice at about mid-length; peristome asymmetric with a widely open pseudosinus; larger lobe with a small circular avicularium directed terminally-proximally relative to the zooidal surface; rostrum finely toothed, crossbar complete, columella absent, cystid invisible, mandible edge raised. Frontal avicularia sporadic, oval and relatively small, crossbar complete, columella absent, palate forming a more or less developed shelf, distal uncalcified area semicircular. Abfrontal avicularia rare, circular, distal part raised with fine denticulation. Ovicells longer than wide with long fissure, and relatively narrow labellum.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="53CA9DB6FFB55E0178B048FAD4C6B4FC" pageId="2" pageNumber="532" type="etymology">
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<paragraph id="1B6FCE3DFFB55E0178B048FAD4C6B4FC" blockId="2.[151,1437,1401,2036]" pageId="2" pageNumber="532">
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<emphasis id="29A4122FFFB55E0178B048FAD41EB487" bold="true" box="[199,334,1937,1963]" pageId="2" pageNumber="532">Etymology.</emphasis>
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Latin “
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<taxonomicName id="DCD0B5BEFFB55E0179DB48FAD756B487" authorityName="Denisenko" authorityYear="2022" box="[428,518,1937,1963]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="532" phylum="Bryozoa" rank="species" species="obscura">
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<emphasis id="29A4122FFFB55E0179DB48FAD756B487" box="[428,518,1937,1963]" italics="true" pageId="2" pageNumber="532">obscura</emphasis>
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</taxonomicName>
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” (unclear), alluding to the possibility of misidentification with closely related species of
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<taxonomicName id="DCD0B5BEFFB55E01797A48DED4DFB4E3" authorityName="Busk" authorityYear="1884" box="[269,399,1973,1999]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="532" phylum="Bryozoa" rank="genus">
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<emphasis id="29A4122FFFB55E01797A48DED4DFB4E3" box="[269,399,1973,1999]" italics="true" pageId="2" pageNumber="532">Reteporella</emphasis>
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</taxonomicName>
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||
.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="53CA9DB6FFB55E0078B048B2D5ACB45E" lastPageId="3" lastPageNumber="533" pageId="2" pageNumber="532" type="description">
|
||
<paragraph id="1B6FCE3DFFB55E0078B048B2D7B6B249" blockId="2.[151,1437,1401,2036]" lastBlockId="3.[151,1437,150,501]" lastPageId="3" lastPageNumber="533" pageId="2" pageNumber="532">
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||
<emphasis id="29A4122FFFB55E0178B048B2D409B4DF" bold="true" box="[199,345,2009,2035]" pageId="2" pageNumber="532">Description.</emphasis>
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The colony is
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||
<quantity id="DC2863D8FFB55E01798B48B2D724B4D8" box="[508,628,2009,2036]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="532" unit="mm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 mm</quantity>
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high, broadly fan-shaped. The fenestrulae are mainly oval, twice as long as wide, commonly 1.1 x
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<quantity id="DC2863D8FFB45E0079C34FFDD747B39D" box="[436,535,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="533" unit="mm" value="0.53">0.53 mm</quantity>
|
||
(
|
||
<figureCitation id="83EBD2B8FFB45E007A524FFCD7CDB39D" box="[549,669,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2A, B</figureCitation>
|
||
;
|
||
<tableCitation id="5652FB86FFB45E007ADF4FFCD7A9B39D" box="[680,761,151,177]" captionStart="TABLE 1" captionStartId="3.[151,239,548,572]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in mm) of Reteporella obscura n. sp. (holotype). SD, standard deviation; N, number of measurements." httpUri="http://table.plazi.org/id/4FAF9EB5FFB45E0078E04D4FD467B14C" pageId="3" pageNumber="533" tableUuid="4FAF9EB5FFB45E0078E04D4FD467B14C">Table 1</tableCitation>
|
||
). The trabeculae (0.85 x
|
||
<quantity id="DC2863D8FFB45E007C674FFDD125B39D" box="[1040,1141,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="533" unit="mm" value="1.05">1.05 mm</quantity>
|
||
) consisting of two to three alternating, longitudinal series of autozooids, doubled at points of trabecular bifurcations (
|
||
<figureCitation id="83EBD2B8FFB45E007CE34FD0D1A0B3F9" box="[1172,1264,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2A</figureCitation>
|
||
). The abfrontal surface of the colony is densely and finely granulated, covered by flat kenozooids clearly delimited by vibices, and with rare round avicularia (
|
||
<figureCitation id="83EBD2B8FFB45E007A714E68D731B231" box="[518,609,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2B</figureCitation>
|
||
). One horn-shaped kenozooid (about
|
||
<quantity id="DC2863D8FFB45E007C634E69D13DB231" box="[1044,1133,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="533" unit="mm" value="0.5">0.5 mm</quantity>
|
||
in length) with a rounded avicularium was observed (
|
||
<figureCitation id="83EBD2B8FFB45E0079A54E4CD726B26D" box="[466,630,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2A, C, G</figureCitation>
|
||
). The granulation on the frontal side is comparatively coarser but less dense than that on the abfrontal surface (
|
||
<figureCitation id="83EBD2B8FFB45E007A284E20D788B249" box="[607,728,331,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1B6FCE3DFFB45E0078B04E04D197B2D9" blockId="3.[151,1437,150,501]" pageId="3" pageNumber="533">
|
||
Autozooids are elongate hexagonal, delimited by mainly straight edges, and occasionally by sutures, obscured by later calcification; autozooids size is about 0.49 x
|
||
<quantity id="DC2863D8FFB45E007A9A4EF9D632B281" box="[749,866,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="533" unit="mm" value="0.275">0.275 mm</quantity>
|
||
, but width and length vary considerably especially in areas of bifurcation (
|
||
<figureCitation id="83EBD2B8FFB45E0079ED4EDCD75BB2FD" box="[410,523,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2D, J</figureCitation>
|
||
;
|
||
<tableCitation id="5652FB86FFB45E007A614EDCD738B2FD" box="[534,616,439,465]" captionStart="TABLE 1" captionStartId="3.[151,239,548,572]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in mm) of Reteporella obscura n. sp. (holotype). SD, standard deviation; N, number of measurements." httpUri="http://table.plazi.org/id/4FAF9EB5FFB45E0078E04D4FD467B14C" pageId="3" pageNumber="533" tableUuid="4FAF9EB5FFB45E0078E04D4FD467B14C">Table 1</tableCitation>
|
||
). The autozooids have few (2–3) large, distinct marginal pores but zooids located near fenestrulae have an additional row of 5–6 pores near the external edge (
|
||
<figureCitation id="83EBD2B8FFB45E007C3F4EB0D1E9B2D9" box="[1096,1209,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2D, J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="4FAF9EB5FFB45E0078E04D4FD467B14C" ID-Table-UUID="4FAF9EB5FFB45E0078E04D4FD467B14C" httpUri="http://table.plazi.org/id/4FAF9EB5FFB45E0078E04D4FD467B14C" pageId="3" pageNumber="533" startId="3.[151,239,548,572]" targetBox="[166,1347,625,1293]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="1B6FCE3DFFB45E0078E04D4FD467B14C" blockId="3.[151,1436,548,608]" pageId="3" pageNumber="533">
|
||
<emphasis id="29A4122FFFB45E0078E04D4FD45EB110" bold="true" box="[151,270,548,572]" pageId="3" pageNumber="533">TABLE 1.</emphasis>
|
||
Measurements (in mm) of
|
||
<taxonomicName id="DCD0B5BEFFB45E007A324D4FD672B110" authority="Denisenko, 2022" authorityName="Denisenko" authorityYear="2022" box="[581,802,548,572]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="533" phylum="Bryozoa" rank="species" species="obscura" status="sp. nov.">
|
||
<emphasis id="29A4122FFFB45E007A324D4FD672B110" box="[581,802,548,572]" italics="true" pageId="3" pageNumber="533">Reteporella obscura</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="29A4122FFFB45E007B5B4D4ED63CB111" bold="true" box="[812,876,549,573]" pageId="3" pageNumber="533">
|
||
<taxonomicNameLabel id="3297AF54FFB45E007B5B4D4ED63CB111" box="[812,876,549,573]" pageId="3" pageNumber="533" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype). SD, standard deviation; N, number of measurements.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1B6FCE3DFFB45E0078D14D1AD067B620" pageId="3" pageNumber="533">
|
||
<table id="69D03C9DFFB4A1FC78D14D1AD013B621" box="[166,1347,625,1293]" colsContinueIn="6.[166,1346,337,1005]" gridcols="4" gridrows="17" pageId="3" pageNumber="533">
|
||
<tr id="A5E0CC7FFFB4A1FC78D14D1AD013B1A5" box="[166,1347,625,649]" gridrow="0" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14D1AD741B1A5" box="[166,529,625,649]" gridcol="0" gridrow="0" pageId="3" pageNumber="533">Character measured</th>
|
||
<th id="E631A503FFB4A1FC7A8A4D1AD62EB1A5" box="[765,894,625,649]" gridcol="1" gridrow="0" pageId="3" pageNumber="533">Mean ± SD</th>
|
||
<th id="E631A503FFB4A1FC7C644D1AD1C2B1A5" box="[1043,1170,625,649]" gridcol="2" gridrow="0" pageId="3" pageNumber="533">Range</th>
|
||
<th id="E631A503FFB4A1FC7D5D4D1AD013B1A5" box="[1322,1347,625,649]" gridcol="3" gridrow="0" pageId="3" pageNumber="533">N</th>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14DF2D013B19D" box="[166,1347,665,689]" gridrow="1" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14DF2D741B19D" box="[166,529,665,689]" gridcol="0" gridrow="1" pageId="3" pageNumber="533">No. zooid rows between fenestrulae</th>
|
||
<td id="E631A503FFB4A1FC7A8A4DF2D62EB19D" box="[765,894,665,689]" gridcol="1" gridrow="1" pageId="3" pageNumber="533">‒</td>
|
||
<td id="E631A503FFB4A1FC7C644DF2D1C2B19D" box="[1043,1170,665,689]" gridcol="2" gridrow="1" pageId="3" pageNumber="533">2‒3</td>
|
||
<td id="E631A503FFB4A1FC7D5D4DF2D013B19D" box="[1322,1347,665,689]" gridcol="3" gridrow="1" pageId="3" pageNumber="533">30</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14DAAD013B1F5" box="[166,1347,705,729]" gridrow="2" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14DAAD741B1F5" box="[166,529,705,729]" gridcol="0" gridrow="2" pageId="3" pageNumber="533">Fenestrula length</th>
|
||
<td id="E631A503FFB4A1FC7A8A4DAAD62EB1F5" box="[765,894,705,729]" gridcol="1" gridrow="2" pageId="3" pageNumber="533">1.098±0.146</td>
|
||
<td id="E631A503FFB4A1FC7C644DAAD1C2B1F5" box="[1043,1170,705,729]" gridcol="2" gridrow="2" pageId="3" pageNumber="533">0.803‒1.304</td>
|
||
<td id="E631A503FFB4A1FC7D5D4DAAD013B1F5" box="[1322,1347,705,729]" gridcol="3" gridrow="2" pageId="3" pageNumber="533">15</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14D82D013B02D" box="[166,1347,745,769]" gridrow="3" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14D82D741B02D" box="[166,529,745,769]" gridcol="0" gridrow="3" pageId="3" pageNumber="533">Fenestrula width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4D82D62EB02D" box="[765,894,745,769]" gridcol="1" gridrow="3" pageId="3" pageNumber="533">0.513±0.105</td>
|
||
<td id="E631A503FFB4A1FC7C644D82D1C2B02D" box="[1043,1170,745,769]" gridcol="2" gridrow="3" pageId="3" pageNumber="533">0.305‒0.683</td>
|
||
<td id="E631A503FFB4A1FC7D5D4D82D013B02D" box="[1322,1347,745,769]" gridcol="3" gridrow="3" pageId="3" pageNumber="533">15</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14C7AD013B006" box="[166,1347,785,810]" gridrow="4" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14C7AD741B006" box="[166,529,785,810]" gridcol="0" gridrow="4" pageId="3" pageNumber="533">Trabecula length</th>
|
||
<td id="E631A503FFB4A1FC7A8A4C7AD62EB006" box="[765,894,785,810]" gridcol="1" gridrow="4" pageId="3" pageNumber="533">0.857±0.282</td>
|
||
<td id="E631A503FFB4A1FC7C644C7AD1C2B006" box="[1043,1170,785,810]" gridcol="2" gridrow="4" pageId="3" pageNumber="533">0.491‒1.311</td>
|
||
<td id="E631A503FFB4A1FC7D5D4C7AD013B006" box="[1322,1347,785,810]" gridcol="3" gridrow="4" pageId="3" pageNumber="533">15</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14C51D013B07E" box="[166,1347,826,850]" gridrow="5" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14C51D741B07E" box="[166,529,826,850]" gridcol="0" gridrow="5" pageId="3" pageNumber="533">Trabecula width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4C51D62EB07E" box="[765,894,826,850]" gridcol="1" gridrow="5" pageId="3" pageNumber="533">1.070±0.146</td>
|
||
<td id="E631A503FFB4A1FC7C644C51D1C2B07E" box="[1043,1170,826,850]" gridcol="2" gridrow="5" pageId="3" pageNumber="533">0.743‒1.221</td>
|
||
<td id="E631A503FFB4A1FC7D5D4C51D013B07E" box="[1322,1347,826,850]" gridcol="3" gridrow="5" pageId="3" pageNumber="533">13</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14C09D013B056" box="[166,1347,866,890]" gridrow="6" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14C09D741B056" box="[166,529,866,890]" gridcol="0" gridrow="6" pageId="3" pageNumber="533">Autozooid length</th>
|
||
<td id="E631A503FFB4A1FC7A8A4C09D62EB056" box="[765,894,866,890]" gridcol="1" gridrow="6" pageId="3" pageNumber="533">0.491±0.073</td>
|
||
<td id="E631A503FFB4A1FC7C644C09D1C2B056" box="[1043,1170,866,890]" gridcol="2" gridrow="6" pageId="3" pageNumber="533">0.378‒0.665</td>
|
||
<td id="E631A503FFB4A1FC7D5D4C09D013B056" box="[1322,1347,866,890]" gridcol="3" gridrow="6" pageId="3" pageNumber="533">17</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14CE1D013B08E" box="[166,1347,906,930]" gridrow="7" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14CE1D741B08E" box="[166,529,906,930]" gridcol="0" gridrow="7" pageId="3" pageNumber="533">Autozooid width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4CE1D62EB08E" box="[765,894,906,930]" gridcol="1" gridrow="7" pageId="3" pageNumber="533">0.273±0.045</td>
|
||
<td id="E631A503FFB4A1FC7C644CE1D1C2B08E" box="[1043,1170,906,930]" gridcol="2" gridrow="7" pageId="3" pageNumber="533">0.210‒0.371</td>
|
||
<td id="E631A503FFB4A1FC7D5D4CE1D013B08E" box="[1322,1347,906,930]" gridcol="3" gridrow="7" pageId="3" pageNumber="533">17</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14CD9D013B0E7" box="[166,1347,946,971]" gridrow="8" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14CD9D741B0E7" box="[166,529,946,971]" gridcol="0" gridrow="8" pageId="3" pageNumber="533">Orifice length</th>
|
||
<td id="E631A503FFB4A1FC7A8A4CD9D62EB0E7" box="[765,894,946,971]" gridcol="1" gridrow="8" pageId="3" pageNumber="533">0.094±0.015</td>
|
||
<td id="E631A503FFB4A1FC7C644CD9D1C2B0E7" box="[1043,1170,946,971]" gridcol="2" gridrow="8" pageId="3" pageNumber="533">0.079‒0.109</td>
|
||
<td id="E631A503FFB4A1FC7D5D4CD9D013B0E7" box="[1322,1347,946,971]" gridcol="3" gridrow="8" pageId="3" pageNumber="533">4</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14CB0D013B0DF" box="[166,1347,987,1011]" gridrow="9" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14CB0D741B0DF" box="[166,529,987,1011]" gridcol="0" gridrow="9" pageId="3" pageNumber="533">Orifice width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4CB0D62EB0DF" box="[765,894,987,1011]" gridcol="1" gridrow="9" pageId="3" pageNumber="533">0.100±0.012</td>
|
||
<td id="E631A503FFB4A1FC7C644CB0D1C2B0DF" box="[1043,1170,987,1011]" gridcol="2" gridrow="9" pageId="3" pageNumber="533">0.083‒0.117</td>
|
||
<td id="E631A503FFB4A1FC7D5D4CB0D013B0DF" box="[1322,1347,987,1011]" gridcol="3" gridrow="9" pageId="3" pageNumber="533">6</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14B68D013B737" box="[166,1347,1027,1051]" gridrow="10" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14B68D741B737" box="[166,529,1027,1051]" gridcol="0" gridrow="10" pageId="3" pageNumber="533">Suboral avicularium length</th>
|
||
<td id="E631A503FFB4A1FC7A8A4B68D62EB737" box="[765,894,1027,1051]" gridcol="1" gridrow="10" pageId="3" pageNumber="533">0.041±0.003</td>
|
||
<td id="E631A503FFB4A1FC7C644B68D1C2B737" box="[1043,1170,1027,1051]" gridcol="2" gridrow="10" pageId="3" pageNumber="533">0.038‒0.046</td>
|
||
<td id="E631A503FFB4A1FC7D5D4B68D013B737" box="[1322,1347,1027,1051]" gridcol="3" gridrow="10" pageId="3" pageNumber="533">7</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14B40D013B76F" box="[166,1347,1067,1091]" gridrow="11" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14B40D741B76F" box="[166,529,1067,1091]" gridcol="0" gridrow="11" pageId="3" pageNumber="533">Suboral avicularium width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4B40D62EB76F" box="[765,894,1067,1091]" gridcol="1" gridrow="11" pageId="3" pageNumber="533">0.049±0.004</td>
|
||
<td id="E631A503FFB4A1FC7C644B40D1C2B76F" box="[1043,1170,1067,1091]" gridcol="2" gridrow="11" pageId="3" pageNumber="533">0.041–0.051</td>
|
||
<td id="E631A503FFB4A1FC7D5D4B40D013B76F" box="[1322,1347,1067,1091]" gridcol="3" gridrow="11" pageId="3" pageNumber="533">15</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14B38D013B740" box="[166,1347,1107,1132]" gridrow="12" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14B38D741B740" box="[166,529,1107,1132]" gridcol="0" gridrow="12" pageId="3" pageNumber="533">Frontal avicularium length</th>
|
||
<td id="E631A503FFB4A1FC7A8A4B38D62EB740" box="[765,894,1107,1132]" gridcol="1" gridrow="12" pageId="3" pageNumber="533">0.084±0.014</td>
|
||
<td id="E631A503FFB4A1FC7C644B38D1C2B740" box="[1043,1170,1107,1132]" gridcol="2" gridrow="12" pageId="3" pageNumber="533">0.066‒0.108</td>
|
||
<td id="E631A503FFB4A1FC7D5D4B38D013B740" box="[1322,1347,1107,1132]" gridcol="3" gridrow="12" pageId="3" pageNumber="533">7</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14B17D013B7B8" box="[166,1347,1148,1172]" gridrow="13" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14B17D741B7B8" box="[166,529,1148,1172]" gridcol="0" gridrow="13" pageId="3" pageNumber="533">Frontal avicularium width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4B17D62EB7B8" box="[765,894,1148,1172]" gridcol="1" gridrow="13" pageId="3" pageNumber="533">0.053±0.009</td>
|
||
<td id="E631A503FFB4A1FC7C644B17D1C2B7B8" box="[1043,1170,1148,1172]" gridcol="2" gridrow="13" pageId="3" pageNumber="533">0.043‒0.063</td>
|
||
<td id="E631A503FFB4A1FC7D5D4B17D013B7B8" box="[1322,1347,1148,1172]" gridcol="3" gridrow="13" pageId="3" pageNumber="533">5</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14BCFD013B790" box="[166,1347,1188,1212]" gridrow="14" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14BCFD741B790" box="[166,529,1188,1212]" gridcol="0" gridrow="14" pageId="3" pageNumber="533">Abfrontal avicularium length/width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4BCFD62EB790" box="[765,894,1188,1212]" gridcol="1" gridrow="14" pageId="3" pageNumber="533">0.067±0.002</td>
|
||
<td id="E631A503FFB4A1FC7C644BCFD1C2B790" box="[1043,1170,1188,1212]" gridcol="2" gridrow="14" pageId="3" pageNumber="533">0.066–0.070</td>
|
||
<td id="E631A503FFB4A1FC7D5D4BCFD013B790" box="[1322,1347,1188,1212]" gridcol="3" gridrow="14" pageId="3" pageNumber="533">3</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14BA7D013B7C8" box="[166,1347,1228,1252]" gridrow="15" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14BA7D741B7C8" box="[166,529,1228,1252]" gridcol="0" gridrow="15" pageId="3" pageNumber="533">Ooecium length (without labellum)</th>
|
||
<td id="E631A503FFB4A1FC7A8A4BA7D62EB7C8" box="[765,894,1228,1252]" gridcol="1" gridrow="15" pageId="3" pageNumber="533">0.214±0.020</td>
|
||
<td id="E631A503FFB4A1FC7C644BA7D1C2B7C8" box="[1043,1170,1228,1252]" gridcol="2" gridrow="15" pageId="3" pageNumber="533">0.182‒0.231</td>
|
||
<td id="E631A503FFB4A1FC7D5D4BA7D013B7C8" box="[1322,1347,1228,1252]" gridcol="3" gridrow="15" pageId="3" pageNumber="533">8</td>
|
||
</tr>
|
||
<tr id="A5E0CC7FFFB4A1FC78D14B9FD013B621" box="[166,1347,1268,1293]" gridrow="16" pageId="3" pageNumber="533">
|
||
<th id="E631A503FFB4A1FC78D14B9FD741B621" box="[166,529,1268,1293]" gridcol="0" gridrow="16" pageId="3" pageNumber="533">Ooecium width</th>
|
||
<td id="E631A503FFB4A1FC7A8A4B9FD62EB621" box="[765,894,1268,1293]" gridcol="1" gridrow="16" pageId="3" pageNumber="533">0.201±0.014</td>
|
||
<td id="E631A503FFB4A1FC7C644B9FD1C2B621" box="[1043,1170,1268,1293]" gridcol="2" gridrow="16" pageId="3" pageNumber="533">0.175‒0.220</td>
|
||
<td id="E631A503FFB4A1FC7D5D4B9FD013B621" box="[1322,1347,1268,1293]" gridcol="3" gridrow="16" pageId="3" pageNumber="533">7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="1B6FCE3DFFB45E0078B04A57D742B6EE" blockId="3.[151,1437,1340,1906]" pageId="3" pageNumber="533">The primary orifice is semicircular with slightly prominent proximal edge, slightly wider than long (mean size 0.094 x 0.100 mm); the distal margin of the orifice is smooth along its upper edge and with well pronounced denticles along its lower margin; condyles are large and rounded. A pair of oral spines is present, one on each side of the orifice at about mid-length.</paragraph>
|
||
<paragraph id="1B6FCE3DFFB45E0078B04AA7D71CB502" blockId="3.[151,1437,1340,1906]" pageId="3" pageNumber="533">
|
||
The peristome hides the orifice in frontal view and consists of two unequal lobes forming an open sinus (
|
||
<figureCitation id="83EBD2B8FFB45E007D184AA7D58CB526" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2D–F</figureCitation>
|
||
). A single circular avicularium is present on its rim, mid-proximally,
|
||
<quantity id="DC2863D8FFB45E007BAC4A9BD11CB527" box="[987,1100,1520,1547]" metricMagnitude="-5" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="533" unit="mm" value="0.041">0.041 mm</quantity>
|
||
long, proximally directed and oblique to the frontal surface (
|
||
<figureCitation id="83EBD2B8FFB45E00799D497FD710B502" box="[490,576,1556,1582]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1B6FCE3DFFB45E0078B04953D43AB42A" blockId="3.[151,1437,1340,1906]" pageId="3" pageNumber="533">
|
||
The avicularium is immersed in the peristome, the avicularian cystid is invisible, and the distal rim of the mandible is finely toothed. Frontal avicularia are sporadic, oval (0.084 x
|
||
<quantity id="DC2863D8FFB45E007BA84937D104B55B" box="[991,1108,1628,1655]" metricMagnitude="-5" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="533" unit="mm" value="0.053">0.053 mm</quantity>
|
||
) demarcated by a raised rim (
|
||
<figureCitation id="83EBD2B8FFB45E0078D749EBD440B5B7" box="[160,272,1664,1691]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2D, J</figureCitation>
|
||
); the distal margin of the rostrum is smooth, the mandible is semi-oval, the crossbar is complete without columella, and the distal uncalcified area is semicircular. Abfrontal avicularia are small, nearly circular (diameter
|
||
<quantity id="DC2863D8FFB45E0078E049A3D45EB5CF" box="[151,270,1736,1763]" metricMagnitude="-5" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="533" unit="mm" value="0.042">0.042 mm</quantity>
|
||
); the distal margin of the rostrum is toothed, the mandible is semicircular, and the crossbar is complete without columella.
|
||
</paragraph>
|
||
<paragraph id="1B6FCE3DFFB45E0078B0487BD5ACB45E" blockId="3.[151,1437,1340,1906]" pageId="3" pageNumber="533">
|
||
Ovicells are located above the distal rim of the orifice; ooecia are longer than wide, with an elongate median fissure and a narrow squared labellum; ooecia become partly obscured by secondary calcification in late ontogeny (
|
||
<figureCitation id="83EBD2B8FFB45E0078D74833D5BFB45E" box="[160,239,1880,1906]" captionStart="FIGURE 2" captionStartId="4.[151,250,1434,1458]" captionTargetBox="[170,1417,181,1409]" captionTargetId="figure-15@4.[170,1417,181,1409]" captionTargetPageId="4" captionText="FIGURE 2. Reteporella obscura n. sp., holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G.Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="3" pageNumber="533">Fig. 2I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="4FAF9EB5FFB35E0778E04AF1D069B56E" ID-DOI="http://doi.org/10.5281/zenodo.6504008" ID-Zenodo-Dep="6504008" httpUri="https://zenodo.org/record/6504008/files/figure.png" pageId="4" pageNumber="534" startId="4.[151,250,1434,1458]" targetBox="[170,1417,181,1409]" targetPageId="4">
|
||
<paragraph id="1B6FCE3DFFB35E0778E04AF1D069B56E" blockId="4.[151,1437,1434,1602]" pageId="4" pageNumber="534">
|
||
<emphasis id="29A4122FFFB35E0778E04AF1D444B69E" bold="true" box="[151,276,1434,1458]" pageId="4" pageNumber="534">FIGURE 2.</emphasis>
|
||
<taxonomicName id="DCD0B5BEFFB35E07796C4AF1D4B9B69E" authority="Denisenko, 2022" authorityName="Denisenko" authorityYear="2022" box="[283,489,1434,1458]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="obscura" status="sp. nov.">
|
||
<emphasis id="29A4122FFFB35E07796C4AF1D4B9B69E" box="[283,489,1434,1458]" italics="true" pageId="4" pageNumber="534">Reteporella obscura</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="29A4122FFFB35E0779874AF1D77AB69E" bold="true" box="[496,554,1434,1458]" pageId="4" pageNumber="534">
|
||
<taxonomicNameLabel id="3297AF54FFB35E0779874AF1D77AB69E" box="[496,554,1434,1458]" pageId="4" pageNumber="534" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ZIN 1/50742. A. Frontal part of colony; autozooids organized in 2–3 series between fenestrulae. B. Densely granulated abfrontal surface. C. Horn-shaped kenozooid with small avicularium. D. Group of zooids. E. Rounded condyles and denticulation of primary orifice. F. Distal part of orifice, showing denticulate distal margin and the circular, proximo-laterally directed suboral avicularium. G. Abfrontal avicularium. E. Ovicell with well-developed labellum and elongate median fissure. J. Group of zooids. Scale bars: A, B, 1 mm; C, E, F, G, 50 μm; D, 400 μm; I, J, 200 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="53CA9DB6FFB35E0678B049FFD038B3F9" lastPageId="5" lastPageNumber="535" pageId="4" pageNumber="534" type="discussion">
|
||
<paragraph id="1B6FCE3DFFB35E0678B049FFD038B3F9" blockId="4.[151,1437,1684,2035]" lastBlockId="5.[151,1437,151,250]" lastPageId="5" lastPageNumber="535" pageId="4" pageNumber="534">
|
||
<emphasis id="29A4122FFFB35E0778B049FFD410B582" bold="true" box="[199,320,1684,1710]" pageId="4" pageNumber="534">Remarks.</emphasis>
|
||
<taxonomicName id="DCD0B5BEFFB35E07793049FFD767B582" authority="Denisenko, 2022" authorityName="Denisenko" authorityYear="2022" box="[327,567,1684,1710]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="obscura" status="sp. nov.">
|
||
<emphasis id="29A4122FFFB35E07793049FFD767B582" box="[327,567,1684,1710]" italics="true" pageId="4" pageNumber="534">Reteporella obscura</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="29A4122FFFB35E077A4A49FFD7D0B582" bold="true" box="[573,640,1684,1710]" pageId="4" pageNumber="534">
|
||
<taxonomicNameLabel id="3297AF54FFB35E077A4A49FFD7D0B582" box="[573,640,1684,1710]" pageId="4" pageNumber="534" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was found in the sublittoral zone, at two stations in the Davis Strait of SW
|
||
<collectingCountry id="63C78EADFFB35E07789149D3D435B5FE" box="[230,357,1720,1746]" name="Greenland" pageId="4" pageNumber="534">Greenland</collectingCountry>
|
||
.At first glance, the new species is very similar to
|
||
<taxonomicName id="DCD0B5BEFFB35E077BDF49D2D164B5FE" baseAuthorityName="King" baseAuthorityYear="1846" box="[936,1076,1720,1746]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="beaniana">
|
||
<emphasis id="29A4122FFFB35E077BDF49D2D164B5FE" box="[936,1076,1720,1746]" italics="true" pageId="4" pageNumber="534">R. beaniana</emphasis>
|
||
</taxonomicName>
|
||
, which is widely distributed in the Arctic Atlantic, in the appearance of colonies and zooids, in the morphology of the secondary orifice and peristome with widely open pseudosinus, and in having a pair of orificial spines. However, the new species is distinguishable from
|
||
<taxonomicName id="DCD0B5BEFFB35E0779C2484ED715B412" baseAuthorityName="King" baseAuthorityYear="1846" box="[437,581,1828,1854]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="beaniana">
|
||
<emphasis id="29A4122FFFB35E0779C2484ED715B412" box="[437,581,1828,1854]" italics="true" pageId="4" pageNumber="534">R. beaniana</emphasis>
|
||
</taxonomicName>
|
||
in having a distinct granulation on both surfaces of the colony, and in the shape of the primary orifice which is bell-shaped in
|
||
<taxonomicName id="DCD0B5BEFFB35E077B354822D681B44E" baseAuthorityName="King" baseAuthorityYear="1846" box="[834,977,1864,1890]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="beaniana">
|
||
<emphasis id="29A4122FFFB35E077B354822D681B44E" box="[834,977,1864,1890]" italics="true" pageId="4" pageNumber="534">R. beaniana</emphasis>
|
||
</taxonomicName>
|
||
and semicircular in
|
||
<taxonomicName id="DCD0B5BEFFB35E077CBB4822D01DB44E" authority="Denisenko, 2022" authorityName="Denisenko" authorityYear="2022" box="[1228,1357,1864,1890]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="obscura" status="sp. nov.">
|
||
<emphasis id="29A4122FFFB35E077CBB4822D01DB44E" box="[1228,1357,1864,1890]" italics="true" pageId="4" pageNumber="534">R. obscura</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="29A4122FFFB35E077D2F4823D0CDB44E" bold="true" box="[1368,1437,1864,1890]" pageId="4" pageNumber="534">
|
||
<taxonomicNameLabel id="3297AF54FFB35E077D2F4823D0CDB44E" box="[1368,1437,1864,1890]" pageId="4" pageNumber="534" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Differences are also observed in the shape of the condyles, which are roundish in the new species and rounded triangular in
|
||
<taxonomicName id="DCD0B5BEFFB35E07794148FAD496B486" baseAuthorityName="King" baseAuthorityYear="1846" box="[310,454,1936,1962]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="beaniana">
|
||
<emphasis id="29A4122FFFB35E07794148FAD496B486" box="[310,454,1936,1962]" italics="true" pageId="4" pageNumber="534">R. beaniana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7F41B3CCFFB35E0779AF48FBD65BB486" author="Hayward, P. J. & Ryland, J. S." box="[472,779,1936,1962]" pageId="4" pageNumber="534" pagination="103 - 112" refId="ref7819" refString="Hayward, P. J. & Ryland, J. S. (1996) Some British Phidoloporidae (Bryozoa: Cheilostomatida). Zoological Journal of the Linnean Society, 117, 103 - 112. https: // doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02150. x" type="journal article" year="1996">Hayward & Ryland 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7F41B3CCFFB35E077B6E48FBD609B486" author="Hayward, P. J. & Ryland, J. S." box="[793,857,1936,1962]" pageId="4" pageNumber="534" pagination="1 - 416" refId="ref7876" refString="Hayward, P. J. & Ryland, J. S. (1999) Cheilostomatous Bryozoa. Part 2. Hippothoidea - Celleporoidea. Synopses of the British Fauna, New Series, 14, 1 - 416." type="journal article" year="1999">1999</bibRefCitation>
|
||
). The suboral avicularium of the new species is round with an indistinct cystid, whereas in
|
||
<taxonomicName id="DCD0B5BEFFB35E077ADC48DED66CB4E2" baseAuthorityName="King" baseAuthorityYear="1846" box="[683,828,1972,1998]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="beaniana">
|
||
<emphasis id="29A4122FFFB35E077ADC48DED66CB4E2" box="[683,828,1972,1998]" italics="true" pageId="4" pageNumber="534">R. beaniana</emphasis>
|
||
</taxonomicName>
|
||
it is oval and the cystid is prominently emerging above the frontal shield. In addition,
|
||
<taxonomicName id="DCD0B5BEFFB35E077A2748B3D610B4DE" authority="Denisenko, 2022" authorityName="Denisenko" authorityYear="2022" box="[592,832,2008,2034]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="534" phylum="Bryozoa" rank="species" species="obscura" status="sp. nov.">
|
||
<emphasis id="29A4122FFFB35E077A2748B3D610B4DE" box="[592,832,2008,2034]" italics="true" pageId="4" pageNumber="534">Reteporella obscura</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="29A4122FFFB35E077B3F48B3D6DBB4DE" bold="true" box="[840,907,2008,2034]" pageId="4" pageNumber="534">
|
||
<taxonomicNameLabel id="3297AF54FFB35E077B3F48B3D6DBB4DE" box="[840,907,2008,2034]" pageId="4" pageNumber="534" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is characterized by globular ovicells, longer than wide with a prominent but relatively narrow labellum, whereas
|
||
<taxonomicName id="DCD0B5BEFFB25E067BBC4FFCD10AB39D" baseAuthorityName="King" baseAuthorityYear="1846" box="[971,1114,151,177]" class="Gymnolaemata" family="Phidoloporidae" genus="Reteporella" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="535" phylum="Bryozoa" rank="species" species="beaniana">
|
||
<emphasis id="29A4122FFFB25E067BBC4FFCD10AB39D" box="[971,1114,151,177]" italics="true" pageId="5" pageNumber="535">R. beaniana</emphasis>
|
||
</taxonomicName>
|
||
has ovicells that are wider than long, and with an arched aperture, and a flattened frontal surface (
|
||
<bibRefCitation id="7F41B3CCFFB25E067B944FD0D040B3F9" author="Hayward, P. J. & Ryland, J. S." box="[995,1296,187,213]" pageId="5" pageNumber="535" pagination="103 - 112" refId="ref7819" refString="Hayward, P. J. & Ryland, J. S. (1996) Some British Phidoloporidae (Bryozoa: Cheilostomatida). Zoological Journal of the Linnean Society, 117, 103 - 112. https: // doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02150. x" type="journal article" year="1996">Hayward & Ryland 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7F41B3CCFFB25E067D6B4FD0D00BB3F9" author="Hayward, P. J. & Ryland, J. S." box="[1308,1371,187,213]" pageId="5" pageNumber="535" pagination="1 - 416" refId="ref7876" refString="Hayward, P. J. & Ryland, J. S. (1999) Cheilostomatous Bryozoa. Part 2. Hippothoidea - Celleporoidea. Synopses of the British Fauna, New Series, 14, 1 - 416." type="journal article" year="1999">1999</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="53CA9DB6FFB25E0678B04FB5D125B3D6" box="[199,1141,222,250]" pageId="5" pageNumber="535" type="distribution">
|
||
<paragraph id="1B6FCE3DFFB25E0678B04FB5D125B3D6" blockId="5.[151,1437,151,250]" box="[199,1141,222,250]" pageId="5" pageNumber="535">
|
||
<emphasis id="29A4122FFFB25E0678B04FB5D431B3D4" bold="true" box="[199,353,222,248]" pageId="5" pageNumber="535">Distribution.</emphasis>
|
||
Lower sublittoral zone (
|
||
<quantity id="DC2863D8FFB25E067A034FB5D7A7B3D5" box="[628,759,222,249]" metricMagnitude="2" metricUnit="m" metricValue="2.21" metricValueMax="2.77" metricValueMin="1.65" pageId="5" pageNumber="535" unit="m" value="221.0" valueMax="277.0" valueMin="165.0">165–277 m</quantity>
|
||
), Davis Strait (SW of
|
||
<collectingCountry id="63C78EADFFB25E067B984FB4D139B3D6" box="[1007,1129,223,250]" name="Greenland" pageId="5" pageNumber="535">Greenland</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |