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<document id="E42AFB27A923A9FEE366AA9B3E3D7E1A" ID-CLB-Dataset="27991" ID-DOI="10.11646/zootaxa.4510.1.1" ID-GBIF-Dataset="f3128a32-5f0b-413e-a755-c859bdd7cac4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10072806" ID-ZooBank="EAA35595-0972-4CF8-A128-16267A59112B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553597889117" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2018" docId="5359945697C0FF16FF75FF53FBCCB95D" docLanguage="en" docName="zootaxa.4510.1.1.pdf.imf" docOrigin="Zootaxa 4510 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Capnogryllacris (Capnogryllacris) elongata" docType="treatment" docVersion="13" lastPageNumber="152" masterDocId="AF60EC2E9757FF81FFE2FF8DFFFEBF2F" masterDocTitle="New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera" masterLastPageNumber="278" masterPageNumber="1" pageNumber="152" updateTime="1699662454306" updateUser="ExternalLinkService">
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<mods:title id="C418303A189C5ABDC5EA2CA7FEEDE324">New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera</mods:title>
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<mods:name id="8AD77CCD3DF8215E39D973F0B18A7F39" type="personal">
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<mods:namePart id="72ED4053765317B25282CA105792E6AD">Ingrisch, Sigfrid</mods:namePart>
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<mods:title id="039977B3735AA28E97502F56ECF166A6">Zootaxa</mods:title>
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<mods:date id="87A30F1D30A67E45D602719E1BB88D04">2018</mods:date>
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<mods:number id="AC1F03DD5732F8F207EF2AE8CC8D2544">4510</mods:number>
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<mods:identifier id="336763FD78C8D7CEC705B324CCE566BE" type="DOI">10.11646/zootaxa.4510.1.1</mods:identifier>
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<treatment id="5359945697C0FF16FF75FF53FBCCB95D" ID-DOI="http://doi.org/10.5281/zenodo.5987159" ID-GBIF-Taxon="154497769" ID-Zenodo-Dep="5987159" LSID="urn:lsid:plazi:treatment:5359945697C0FF16FF75FF53FBCCB95D" httpUri="http://treatment.plazi.org/id/5359945697C0FF16FF75FF53FBCCB95D" lastPageNumber="152" pageId="151" pageNumber="152">
<subSubSection id="93EA76CB97C0FF16FF75FF53FD52BFD7" box="[151,684,222,248]" pageId="151" pageNumber="152" type="nomenclature">
<paragraph id="DB4F254097C0FF16FF75FF53FD52BFD7" blockId="151.[151,684,222,282]" box="[151,684,222,248]" pageId="151" pageNumber="152">
<heading id="8007922C97C0FF16FF75FF53FD52BFD7" bold="true" box="[151,684,222,248]" fontSize="11" level="1" pageId="151" pageNumber="152" reason="1">
<taxonomicName id="1CF05EC397C0FF16FF75FF53FD52BFD7" ID-CoL="8P7GG" authority="(Fritze, 1908)" baseAuthorityName="Fritze" baseAuthorityYear="1908" box="[151,684,222,248]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="elongata" subGenus="Capnogryllacris">
<emphasis id="E984F95297C0FF16FF75FF53FD52BFD7" bold="true" box="[151,684,222,248]" pageId="151" pageNumber="152">
<emphasis id="E984F95297C0FF16FF75FF53FE00BFD7" bold="true" box="[151,510,222,248]" italics="true" pageId="151" pageNumber="152">Capnogryllacris (C.) elongata</emphasis>
(
<bibRefCitation id="BF6158B197C0FF16FDEEFF53FD5ABFD7" author="Fritze" box="[524,676,222,248]" pageId="151" pageNumber="152" refId="ref184990" refString="Fritze (1908). See Carl (1908)." type="book" year="1908">Fritze, 1908</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB97C0FF16FF75FE8CFB64BE4D" pageId="151" pageNumber="152" type="description">
<paragraph id="DB4F254097C0FF16FF75FE8CFE66BE35" blockId="151.[151,684,222,282]" box="[151,408,257,282]" pageId="151" pageNumber="152">
<figureCitation id="43CB39C597C0FF16FF75FE8CFEFFBE36" box="[151,257,257,282]" captionStart="FIGURE 47" captionStartId="136.[151,250,1872,1894]" captionTargetBox="[170,1429,193,1848]" captionTargetId="figure@136.[163,1429,193,1856]" captionTargetPageId="136" captionText="FIGURE 47. Genus Capnogryllacris, Habitus dorsal view: AB, C. p. phaeocephala Gorochov, 2003 male (A) and female (B); C, C. basaliatrata (Griffini, 1909) male Brunner's type of G. atrata in NHMW; DE, C. khmerica Gorochov, 2003 male (D) and female (E); F, C. elongata (Fritze, 1908) male; G, C. e. erythrocephala Gorochov, 2003 female; H, C. proxima Gorochov, 2003 female; I, C. (D.) f. fasciculata (Pictet &amp; Sauss., 1891) male; J, C. s. sakaerat Dawwrueng et al., 2015 female; KL, C. s. toxica ssp. nov. male holotype (K) and female paratype (L). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10109110" httpUri="https://zenodo.org/record/10109110/files/figure.png" pageId="151" pageNumber="152">Figs. 47F</figureCitation>
,
<figureCitation id="43CB39C597C0FF16FEEFFE8CFEBEBE36" box="[269,320,257,281]" captionStart="FIGURE 48" captionStartId="137.[151,250,1482,1504]" captionTargetBox="[151,1436,193,1459]" captionTargetId="figure@137.[151,1436,193,1460]" captionTargetPageId="137" captionText="FIGURE 48. Genus Capnogryllacris: AC, C. varifrons sp. nov.; D, C. e. fuscifrons ssp. nov.; E, C. humberti (Griffini, 1914); F, C. nigromaculata sp. nov.; G, C. p. phaeocephala Gorochov, 2003; H, C. e. erythrocephala Gorochov, 2003; I, C. e. borealis Gorochov, 2003; J, C. s. sakaerat Dawwrueng et al., 2015; K, C. elongata (Fritze, 1908); L, C. f. fasciculata (Pictet &amp; Sauss., 1891); M, C. proxima Gorochov, 2003; N, C. s. toxica ssp. nov.; O, C. khmerica Gor.—Face female (A, E, G, I, J, N, O) and male (BD, F, H, KM); not to scale." figureDoi="http://doi.org/10.5281/zenodo.10109112" httpUri="https://zenodo.org/record/10109112/files/figure.png" pageId="151" pageNumber="152">48K</figureCitation>
,
<figureCitation id="43CB39C597C0FF16FEA9FE8CFE98BE35" box="[331,358,257,282]" captionStart="FIGURE 50" captionStartId="139.[151,250,1568,1590]" captionTargetBox="[151,1437,193,1542]" captionTargetId="figure@139.[150,1437,193,1546]" captionTargetPageId="139" captionText="FIGURE 50. Genus Capnogryllacris: AB, C. p. phaeocephala Gorochov, 2003; C, C. khmerica Gorochov, 2003; D, C. basaliatrata (Griffini, 1909); E, C. s. toxica ssp. nov.; FG, C. elongata (Fritze, 1908); H, C. f. fasciculata (Pictet &amp; Sauss., 1891).—Male abdominal apex in apical or ventro-apical view to show projections of ninth abdominal tergite (in BH also subgenital plate)." figureDoi="http://doi.org/10.5281/zenodo.10109116" httpUri="https://zenodo.org/record/10109116/files/figure.png" pageId="151" pageNumber="152">50</figureCitation>
FG
</paragraph>
<paragraph id="DB4F254097C0FF16FF75FEC4FB64BE4D" blockId="151.[151,1437,329,1650]" box="[151,1178,329,354]" pageId="151" pageNumber="152">
<materialsCitation id="6B982F1D97C0FF16FF75FEC4FB64BE4D" ID-GBIF-Occurrence="2028147579" box="[151,1178,329,354]" collectionCode="MHNG" pageId="151" pageNumber="152" specimenCount="1" typeStatus="holotype">
<typeStatus id="044B9BE297C0FF16FF75FEC4FF01BE4D" box="[151,255,329,354]" pageId="151" pageNumber="152" type="holotype">Holotype</typeStatus>
(male): Indo-Malaysia: Borneo—Genève (
<collectionCode id="BDE1BD8597C0FF16FD01FEC4FCC0BE4E" box="[739,830,329,353]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="151" pageNumber="152" type="Museum">MHNG</collectionCode>
), seen but not studied in detail.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB97C0FF16FF25FEE0FDCDBAB6" pageId="151" pageNumber="152" type="materials_examined">
<paragraph id="DB4F254097C0FF16FF25FEE0FEBDBE86" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">
<emphasis id="E984F95297C0FF16FF25FEE0FE4FBEA9" bold="true" box="[199,433,365,390]" pageId="151" pageNumber="152">Material examined.</emphasis>
<materialsCitation id="6B982F1D97C0FF16FE5AFEE0FEC1BE86" ID-GBIF-Occurrence="2028147621" collectingDate="1984-07-19" collectingDateMax="1984-07-22" collectingDateMin="1984-07-19" collectionCode="ZFMK" country="Malaysia" location="Taman Negara" pageId="151" pageNumber="152" specimenCount="1" specimenCount-male="1" stateProvince="Pahang">
<collectingCountry id="A3E765D097C0FF16FE5AFEE0FDDDBEA9" box="[440,547,365,390]" name="Malaysia" pageId="151" pageNumber="152">Malaysia</collectingCountry>
:
<collectingRegion id="1934EBA297C0FF16FDCCFEE0FD7DBEA9" box="[558,643,365,390]" country="Malaysia" name="Pahang" pageId="151" pageNumber="152">Pahang</collectingRegion>
,
<location id="DE2F739B97C0FF16FD72FEE3FCC9BEA9" LSID="urn:lsid:plazi:treatment:5359945697C0FF16FF75FF53FBCCB95D:DE2F739B97C0FF16FD72FEE3FCC9BEA9" box="[656,823,366,390]" country="Malaysia" name="Taman Negara" pageId="151" pageNumber="152" stateProvince="Pahang">Taman Negara</location>
, forest trails,
<date id="AF4E038097C0FF16FC33FEE0FB83BEA9" box="[977,1149,365,390]" pageId="151" pageNumber="152" value="1984-07-19" valueMax="1984-07-22" valueMin="1984-07-19">
<collectingDate id="BF0AFA6897C0FF16FC33FEE0FB83BEA9" box="[977,1149,365,390]" pageId="151" pageNumber="152" value="1984-07-19" valueMax="1984-07-22" valueMin="1984-07-19">1922.vii.1984</collectingDate>
</date>
, leg. S. Ingrisch—
<specimenCount id="CDF6EEC997C0FF16FAB4FEE0FA62BEA9" box="[1366,1436,365,390]" pageId="151" pageNumber="152" type="male">1 male</specimenCount>
(Bonn
<collectionCode id="BDE1BD8597C0FF16FF06FE1CFEC9BE86" box="[228,311,401,425]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="151" pageNumber="152" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DB4F254097C0FF16FF25FE38FADFBD39" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">
Additional
<emphasis id="E984F95297C0FF16FEA4FE38FE26BEE1" bold="true" box="[326,472,437,462]" pageId="151" pageNumber="152">Description.</emphasis>
Medium sized to large species. Head: Face ovoid; forehead nearly smooth; fastigium verticis wider than scapus; ocelli distinct; fastigium frontis separated from fastigium verticis by a very fine suture; subocular furrows very shallow (
<figureCitation id="43CB39C597C0FF16FDEAFE70FD8EBD39" box="[520,624,509,534]" captionStart="FIGURE 48" captionStartId="137.[151,250,1482,1504]" captionTargetBox="[151,1436,193,1459]" captionTargetId="figure@137.[151,1436,193,1460]" captionTargetPageId="137" captionText="FIGURE 48. Genus Capnogryllacris: AC, C. varifrons sp. nov.; D, C. e. fuscifrons ssp. nov.; E, C. humberti (Griffini, 1914); F, C. nigromaculata sp. nov.; G, C. p. phaeocephala Gorochov, 2003; H, C. e. erythrocephala Gorochov, 2003; I, C. e. borealis Gorochov, 2003; J, C. s. sakaerat Dawwrueng et al., 2015; K, C. elongata (Fritze, 1908); L, C. f. fasciculata (Pictet &amp; Sauss., 1891); M, C. proxima Gorochov, 2003; N, C. s. toxica ssp. nov.; O, C. khmerica Gor.—Face female (A, E, G, I, J, N, O) and male (BD, F, H, KM); not to scale." figureDoi="http://doi.org/10.5281/zenodo.10109112" httpUri="https://zenodo.org/record/10109112/files/figure.png" pageId="151" pageNumber="152">Fig. 48K</figureCitation>
). Abdominal tergites two and three without stridulatory pegs.
</paragraph>
<paragraph id="DB4F254097C0FF16FF25FDACFE19BC3D" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">
Wings surpassing middle of stretched hind tibiae (
<figureCitation id="43CB39C597C0FF16FD15FDACFCA5BD16" box="[759,859,545,570]" captionStart="FIGURE 47" captionStartId="136.[151,250,1872,1894]" captionTargetBox="[170,1429,193,1848]" captionTargetId="figure@136.[163,1429,193,1856]" captionTargetPageId="136" captionText="FIGURE 47. Genus Capnogryllacris, Habitus dorsal view: AB, C. p. phaeocephala Gorochov, 2003 male (A) and female (B); C, C. basaliatrata (Griffini, 1909) male Brunner's type of G. atrata in NHMW; DE, C. khmerica Gorochov, 2003 male (D) and female (E); F, C. elongata (Fritze, 1908) male; G, C. e. erythrocephala Gorochov, 2003 female; H, C. proxima Gorochov, 2003 female; I, C. (D.) f. fasciculata (Pictet &amp; Sauss., 1891) male; J, C. s. sakaerat Dawwrueng et al., 2015 female; KL, C. s. toxica ssp. nov. male holotype (K) and female paratype (L). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10109110" httpUri="https://zenodo.org/record/10109110/files/figure.png" pageId="151" pageNumber="152">Fig. 47F</figureCitation>
). Tegmen: Radius forked near tip; media anterior at very base free, then it closely approaches or sub-fuses for about
<quantity id="1C0888A597C0FF16FC85FDC8FC56BD72" box="[871,936,581,606]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="151" pageNumber="152" unit="mm" value="5.0">5 mm</quantity>
with radius, after both veins separated again media receives a short oblique connection vein from radius (probably RS), in apical third of tegmen MA separates from the fused MA+RS vein as an oblique posterior branch while RS terminates into three posterior branches; cubitus anterior at base single-branched and distinctly thickened, it divides at end of basal third, the first branch in a curvature and then divides into MP and CuA1; CuA2 remains single branched. Cubitus posterior undivided, free throughout; with 4 anal veins.
</paragraph>
<paragraph id="DB4F254097C0FF16FF25FC93FBACBC8D" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">Legs: Fore coxa with a spine at fore margin; fore and mid femora unarmed; fore and mid tibiae with four pairs of large ventral spines and one pair of smaller ventral spurs; hind femur with about 67 external and about 1214 internal spines on ventral margins getting distinctly larger towards end; hind tibia with spaced spines on both dorsal margins, ventral margins with one pre-apical spine; with 3 apical spurs on both sides.</paragraph>
<paragraph id="DB4F254097C0FF16FF25FC20FCEDBB79" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">Coloration. General color medium brown with indistinct ornament; rim of pronotum red brown. Face yellowish brown; antennal scrobae, a wide vertical band below compound eyes, margins of clypeus and labrum and maxillary palpi dark brown. Tegmen semi-transparent yellowish, towards base darker; veins medium to dark brown; hind wing dark red brown, towards hind margin lighter and cells with yellowish spot in middle; main veins black to brown; cross veins medium brown to yellowish.</paragraph>
<paragraph id="DB4F254097C0FF16FF25FBECFC00BA7D" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">
Male. Eighth abdominal tergite with dorsal area prolonged, not laterally. Ninth abdominal tergite with a strong medial furrow, globular on both sides; semi-globes at ventro-internal angle compressed and prolonged into a large compressed triangular process pointing about mediad; process at base yellowish, afterwards dark brown with black margin (
<figureCitation id="43CB39C597C0FF16FF14FB40FEADBBC9" box="[246,339,1229,1254]" captionStart="FIGURE 50" captionStartId="139.[151,250,1568,1590]" captionTargetBox="[151,1437,193,1542]" captionTargetId="figure@139.[150,1437,193,1546]" captionTargetPageId="139" captionText="FIGURE 50. Genus Capnogryllacris: AB, C. p. phaeocephala Gorochov, 2003; C, C. khmerica Gorochov, 2003; D, C. basaliatrata (Griffini, 1909); E, C. s. toxica ssp. nov.; FG, C. elongata (Fritze, 1908); H, C. f. fasciculata (Pictet &amp; Sauss., 1891).—Male abdominal apex in apical or ventro-apical view to show projections of ninth abdominal tergite (in BH also subgenital plate)." figureDoi="http://doi.org/10.5281/zenodo.10109116" httpUri="https://zenodo.org/record/10109116/files/figure.png" pageId="151" pageNumber="152">Figs. 50</figureCitation>
FG). Epiproct small triangular. Subgenital plate wider than long with a large triangular excision from base, lateral margins convex and converging, apical margin wide roundly excised, with a shallow furrow in midline between apical and basal excisions; cerci little upcurved, longer than subgenital plate, inserted at wide trunks that occupy all of the non-excised apical margin. Phallus membranous.
</paragraph>
<paragraph id="DB4F254097C0FF16FF25FAD3FDCDBAB6" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">
Measurements (
<specimenCount id="CDF6EEC997C0FF16FE9FFAD3FE37BA59" box="[381,457,1373,1398]" pageId="151" pageNumber="152" type="male">1 male</specimenCount>
).—body w/wings: 51; body w/o wings: 29; pronotum: 6.5; tegmen: 42; tegmen width: 12; hind femur: 18; antenna:
<quantity id="1C0888A597C0FF16FE3FFA0CFDD1BAB6" box="[477,559,1409,1434]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="151" pageNumber="152" unit="mm" value="85.0">85 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB97C0FF16FF25FA28FBCCB95D" pageId="151" pageNumber="152" type="discussion">
<paragraph id="DB4F254097C0FF16FF25FA28FBCCB95D" blockId="151.[151,1437,329,1650]" pageId="151" pageNumber="152">
<emphasis id="E984F95297C0FF16FF25FA28FEB2BA91" bold="true" box="[199,332,1445,1470]" pageId="151" pageNumber="152">Discussion.</emphasis>
Habitus and wing venation agree with the type of
<taxonomicName id="1CF05EC397C0FF16FC75FA28FA69BA91" authority="(Fritze, 1908)" baseAuthorityName="Fritze" baseAuthorityYear="1908" box="[919,1431,1445,1470]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="elongata" subGenus="Capnogryllacris">
<emphasis id="E984F95297C0FF16FC75FA28FB0EBA92" box="[919,1264,1445,1469]" italics="true" pageId="151" pageNumber="152">Capnogryllacris (C.) elongata</emphasis>
(
<bibRefCitation id="BF6158B197C0FF16FAE2FA28FA71BA91" author="Fritze" box="[1280,1423,1445,1470]" pageId="151" pageNumber="152" refId="ref184990" refString="Fritze (1908). See Carl (1908)." type="book" year="1908">Fritze, 1908</bibRefCitation>
)
</taxonomicName>
, except that MA is fused with R in subbasal area instead of only leaning on R but not fused as in the type; basic color of hind wings is dark reddish brown instead of more yellowish brown in the type. Also in the
<typeStatus id="044B9BE297C0FF16FB18FA60FAA2B929" box="[1274,1372,1517,1542]" pageId="151" pageNumber="152" type="holotype">holotype</typeStatus>
there is a short oblique connection branch from R to MA (fusion of RS with M) although it is missing in the drawing that came with the original description by Fritze in
<bibRefCitation id="BF6158B197C0FF16FD4FF9B8FCCAB961" author="Carl, J. &amp; Fritze, A." box="[685,820,1589,1614]" pageId="151" pageNumber="152" pagination="299 - 305" refId="ref184582" refString="Carl, J. [Descriptions by Fritze, A.] (1908) Neue Locustodeen von Ceylon und Borneo (Orthopt.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 11, 299 - 305, pl. 7." type="journal article" year="1908">Carl (1908)</bibRefCitation>
. The shape of the male ninth abdominal tergite and its processes agree with the descriptions in Fritze in
<bibRefCitation id="BF6158B197C0FF16FD5DF9D4FCBFB95D" author="Carl, J. &amp; Fritze, A." box="[703,833,1625,1650]" pageId="151" pageNumber="152" pagination="299 - 305" refId="ref184582" refString="Carl, J. [Descriptions by Fritze, A.] (1908) Neue Locustodeen von Ceylon und Borneo (Orthopt.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 11, 299 - 305, pl. 7." type="journal article" year="1908">Carl (1908)</bibRefCitation>
and
<bibRefCitation id="BF6158B197C0FF16FC95F9D4FBD0B95D" author="Griffini, A." box="[887,1070,1625,1650]" pageId="151" pageNumber="152" pagination="1 - 19" refId="ref186169" refString="Griffini, A. (1911 d) Note sopra alcuni Stenopelmatidi e Grillacridi del Museo di Sarawak. Bollettino del Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 26 (636), 1 - 19." type="journal article" year="1911">Griffini (1911d)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>