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<document id="EC59221221DE9DB26E5857A75B792D13" ID-CLB-Dataset="27991" ID-DOI="10.11646/zootaxa.4510.1.1" ID-GBIF-Dataset="f3128a32-5f0b-413e-a755-c859bdd7cac4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10072806" ID-ZooBank="EAA35595-0972-4CF8-A128-16267A59112B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553597889117" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2018" docId="535994569781FF56FF75FE3CFE96BD75" docLanguage="en" docName="zootaxa.4510.1.1.pdf.imf" docOrigin="Zootaxa 4510 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Melanogryllacris atrata subsp. atrata atrata (Walker 1869" docType="treatment" docVersion="12" lastPageNumber="216" masterDocId="AF60EC2E9757FF81FFE2FF8DFFFEBF2F" masterDocTitle="New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera" masterLastPageNumber="278" masterPageNumber="1" pageNumber="215" updateTime="1699662454306" updateUser="ExternalLinkService">
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<mods:title id="C80A4ED2B4192F3ED9B969BAA00EEBF3">New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera</mods:title>
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<mods:namePart id="9BAA5AF13D2B0B27ED1F2EF286D9F72B">Ingrisch, Sigfrid</mods:namePart>
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<mods:date id="F9C266184CF7F2D9BB0BF790D9672889">2018</mods:date>
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<treatment id="535994569781FF56FF75FE3CFE96BD75" ID-DOI="http://doi.org/10.5281/zenodo.5987287" ID-GBIF-Taxon="154497824" ID-Zenodo-Dep="5987287" LSID="urn:lsid:plazi:treatment:535994569781FF56FF75FE3CFE96BD75" httpUri="http://treatment.plazi.org/id/535994569781FF56FF75FE3CFE96BD75" lastPageId="215" lastPageNumber="216" pageId="214" pageNumber="215">
<subSubSection id="93EA76CB9781FF57FF75FE3CFD3EBEE4" box="[151,704,433,459]" pageId="214" pageNumber="215" type="nomenclature">
<paragraph id="DB4F25409781FF57FF75FE3CFD3EBEE4" blockId="214.[151,704,433,493]" box="[151,704,433,459]" pageId="214" pageNumber="215">
<heading id="8007922C9781FF57FF75FE3CFD3EBEE4" bold="true" box="[151,704,433,459]" fontSize="11" level="1" pageId="214" pageNumber="215" reason="1">
<taxonomicName id="1CF05EC39781FF57FF75FE3CFD3EBEE4" ID-CoL="5JLCB" authority="atrata (Walker, 1869)" authorityName="atrata (Walker" authorityYear="1869" baseAuthorityName="Walker" baseAuthorityYear="1869" box="[151,704,433,459]" class="Insecta" family="Gryllacrididae" genus="Melanogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="subSpecies" species="atrata" subSpecies="atrata">
<emphasis id="E984F9529781FF57FF75FE3CFD3EBEE4" bold="true" box="[151,704,433,459]" pageId="214" pageNumber="215">
<emphasis id="E984F9529781FF57FF75FE3CFDFDBEE4" bold="true" box="[151,515,433,459]" italics="true" pageId="214" pageNumber="215">Melanogryllacris atrata atrata</emphasis>
(
<bibRefCitation id="BF6158B19781FF57FDF0FE3CFD46BEE4" author="Walker, F." box="[530,696,433,459]" pageId="214" pageNumber="215" refId="ref189674" refString="Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum. Pt 1. [iv +] 1 - 224; London (British Museum of Natural History)." type="book" year="1869">Walker, 1869</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB9781FF57FF75FE59FB86BD1B" pageId="214" pageNumber="215" type="description">
<paragraph id="DB4F25409781FF57FF75FE59FE81BEC2" blockId="214.[151,704,433,493]" box="[151,383,468,493]" pageId="214" pageNumber="215">
<figureCitation id="43CB39C59781FF57FF75FE59FF0CBEC3" box="[151,242,468,493]" captionStart="FIGURE 74" captionStartId="210.[151,250,1874,1896]" captionTargetBox="[151,1436,269,1851]" captionTargetId="figure@210.[151,1436,267,1852]" captionTargetPageId="210" captionText="FIGURE 74. Genus Melaneremus: AD, M. fruhstorferi (Griffini, 1914) male holotype (A), female allotype (B) and male from Hoang Lien (CD), EH, M. sikkimensis sp. nov. holotype; I, M. atrotectus (Brunner, 1888) male holotype.—JK, Melanogryllacris atrata (Walker, 1869) female in NHMW.—Habitus lateral view (AB, IJ), habitus dorsal view (C, E, K), face (D, H), anterior area of body with head, pronotum and thorax (F), lateral view on reduced wings (G)." figureDoi="http://doi.org/10.5281/zenodo.10109164" httpUri="https://zenodo.org/record/10109164/files/figure.png" pageId="214" pageNumber="215">Figs. 74</figureCitation>
JK, 76KN
</paragraph>
<paragraph id="DB4F25409781FF57FF75FD91FB86BD1B" blockId="214.[151,1437,540,2021]" box="[151,1144,540,565]" pageId="214" pageNumber="215">
<materialsCitation id="6B982F1D9781FF57FF75FD91FB86BD1B" ID-GBIF-Occurrence="2028147480" box="[151,1144,540,565]" collectionCode="UMO" country="Malaysia" location="Borneo" pageId="214" pageNumber="215" specimenCount="1" typeStatus="holotype">
<typeStatus id="044B9BE29781FF57FF75FD91FF01BD1A" box="[151,255,540,565]" pageId="214" pageNumber="215" type="holotype">Holotype</typeStatus>
(male, not seen):
<location id="DE2F739B9781FF57FE2AFD90FDE3BD1B" LSID="urn:lsid:plazi:treatment:535994569781FF56FF75FE3CFE96BD75:DE2F739B9781FF57FE2AFD90FDE3BD1B" box="[456,541,541,564]" country="Malaysia" name="Borneo" pageId="214" pageNumber="215">Borneo</location>
,
<collectingCountry id="A3E765D09781FF57FDCBFD91FD73BD1A" box="[553,653,540,565]" name="Malaysia" pageId="214" pageNumber="215">Sarawak</collectingCountry>
, (Oxford
<collectionCode id="BDE1BD859781FF57FD1AFD91FCC7BD1B" box="[760,825,540,564]" country="USA" name="University of Maine" pageId="214" pageNumber="215">UMO</collectionCode>
according to
<bibRefCitation id="BF6158B19781FF57FC31FD91FB95BD1B" author="Griffini, A." box="[979,1131,540,565]" pageId="214" pageNumber="215" pagination="82 - 94" refId="ref186022" refString="Griffini, A. (1910) Revision des types de certaines Gryllacris decrites par F. Walker, existant au Musee d' Oxford (Orth.). Deutsche Entomologische Zeitschrift, 1910, 82 - 94; Berlin." type="journal article" year="1910">Griffini 1910</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB9781FF57FF25FDB3FE1CBDCD" pageId="214" pageNumber="215" type="materials_examined">
<paragraph id="DB4F25409781FF57FF25FDB3FE1CBDCD" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
<emphasis id="E984F9529781FF57FF25FDB3FE4EBD78" bold="true" box="[199,432,574,599]" pageId="214" pageNumber="215">Material examined.</emphasis>
<materialsCitation id="6B982F1D9781FF57FE54FDB3FE50BD55" ID-GBIF-Occurrence="2028147499" collectingDate="1925-10-10" collectionCode="NHMW" collectorName="Wien" country="Indonesia" location="Midden Oost Borneo" pageId="214" pageNumber="215" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
<collectingCountry id="A3E765D09781FF57FE54FDB3FDD9BD78" box="[438,551,574,599]" name="Indonesia" pageId="214" pageNumber="215">Indonesia</collectingCountry>
:
<collectingRegion id="1934EBA29781FF57FDD0FDB3FCFCBD78" box="[562,770,574,599]" country="Indonesia" name="Kalimantan Timur" pageId="214" pageNumber="215">Kalimantan Timur</collectingRegion>
,
<location id="DE2F739B9781FF57FCEFFDB3FC06BD79" LSID="urn:lsid:plazi:treatment:535994569781FF56FF75FE3CFE96BD75:DE2F739B9781FF57FCEFFDB3FC06BD79" box="[781,1016,574,599]" country="Indonesia" name="Midden Oost Borneo" pageId="214" pageNumber="215" stateProvince="Kalimantan Timur">Midden Oost Borneo</location>
155,
<date id="AF4E03809781FF57FBD5FDB3FB57BD78" box="[1079,1193,574,599]" pageId="214" pageNumber="215" value="1925-10-10">
<collectingDate id="BF0AFA689781FF57FBD5FDB3FB57BD78" box="[1079,1193,574,599]" pageId="214" pageNumber="215" value="1925-10-10">10.x.1925</collectingDate>
</date>
, leg. H.C. Siebers—
<specimenCount id="CDF6EEC99781FF57FA76FDB3FF1DBD55" pageId="214" pageNumber="215" type="female">1 female</specimenCount>
(
<collectorName id="760540969781FF57FF1BFDECFEC8BD55" box="[249,310,609,634]" pageId="214" pageNumber="215">Wien</collectorName>
<collectionCode id="BDE1BD859781FF57FEA7FDEFFE58BD55" box="[325,422,610,634]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="214" pageNumber="215" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6B982F1D9781FF57FE22FDEFFE0EBD56" ID-GBIF-Occurrence="2028147513" box="[448,496,610,633]" country="Indonesia" location="East" pageId="214" pageNumber="215" specimenCount="1" stateProvince="Kalimantan Timur">
<location id="DE2F739B9781FF57FE22FDEFFE0EBD56" LSID="urn:lsid:plazi:treatment:535994569781FF56FF75FE3CFE96BD75:DE2F739B9781FF57FE22FDEFFE0EBD56" box="[448,496,610,633]" country="Indonesia" name="East" pageId="214" pageNumber="215" stateProvince="Kalimantan Timur">East</location>
</materialsCitation>
<materialsCitation id="6B982F1D9781FF57FE1DFDECFAE1BDB3" ID-GBIF-Occurrence="2028147514" collectingDate="1997-02-24" collectionCode="ZFMK" collectorName="A. Floren &amp; Fog &amp; Bonn" country="Malaysia" county="Mt. Kinabalu" elevation="600" latitude="6.0833335" location="Topou" longLatPrecision="1305" longitude="116.55" municipality="Poring" pageId="214" pageNumber="215" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<collectingCountry id="A3E765D09781FF57FE1DFDECFD94BD55" box="[511,618,609,634]" name="Malaysia" pageId="214" pageNumber="215">Malaysia</collectingCountry>
:
<collectingRegion id="1934EBA29781FF57FD99FDECFD3CBD55" box="[635,706,609,634]" country="Malaysia" name="Sabah" pageId="214" pageNumber="215">Sabah</collectingRegion>
,
<collectingCounty id="322E5DCC9781FF57FD37FDEFFC8DBD55" box="[725,883,609,634]" pageId="214" pageNumber="215">Mt. Kinabalu</collectingCounty>
NP,
<collectingMunicipality id="3B2BBF3A9781FF57FC55FDECFBFABD55" box="[951,1028,609,634]" pageId="214" pageNumber="215">Poring</collectingMunicipality>
,
<location id="DE2F739B9781FF57FBF5FDEFFB9FBD56" LSID="urn:lsid:plazi:treatment:535994569781FF56FF75FE3CFE96BD75:DE2F739B9781FF57FBF5FDEFFB9FBD56" box="[1047,1121,610,633]" country="Malaysia" county="Mt. Kinabalu" latitude="6.0833335" longLatPrecision="1305" longitude="116.55" municipality="Poring" name="Topou" pageId="214" pageNumber="215" stateProvince="Sabah">Topou</location>
, elev.
<quantity id="1C0888A59781FF57FB54FDECFAC3BD56" box="[1206,1341,609,634]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="214" pageNumber="215" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">
<elevation id="50DDC2739781FF57FB54FDECFAC3BD56" box="[1206,1341,609,634]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="214" pageNumber="215" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500700 m</elevation>
</quantity>
(
<geoCoordinate id="BEC443879781FF57FAB6FDECFA69BD55" box="[1364,1431,609,634]" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="925" value="6.0833335">6°5'N</geoCoordinate>
,
<geoCoordinate id="BEC443879781FF57FF75FD09FEFEBDB2" box="[151,256,644,669]" direction="east" orientation="longitude" pageId="214" pageNumber="215" precision="925" value="116.55">116°33'E</geoCoordinate>
),
<date id="AF4E03809781FF57FEFBFD09FE72BDB2" box="[281,396,644,669]" pageId="214" pageNumber="215" value="1997-02-24">
<collectingDate id="BF0AFA689781FF57FEFBFD09FE72BDB2" box="[281,396,644,669]" pageId="214" pageNumber="215" value="1997-02-24">24.ii.1997</collectingDate>
</date>
, coll.
<collectorName id="760540969781FF57FE3BFD09FDB7BDB2" box="[473,585,644,669]" pageId="214" pageNumber="215">A. Floren</collectorName>
(SeWaI 5B5 NL 7B5 NL 7
<collectorName id="760540969781FF57FC41FD09FC2DBDB3" box="[931,979,644,668]" pageId="214" pageNumber="215">Fog</collectorName>
3)—
<specimenCount id="CDF6EEC99781FF57FBE5FD09FB99BDB2" box="[1031,1127,644,669]" pageId="214" pageNumber="215" type="female">1 female</specimenCount>
(
<collectorName id="760540969781FF57FB98FD09FB46BDB3" box="[1146,1208,644,668]" pageId="214" pageNumber="215">Bonn</collectorName>
<collectionCode id="BDE1BD859781FF57FB26FD09FAE9BDB3" box="[1220,1303,644,668]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="214" pageNumber="215" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6B982F1D9781FF57FACCFD09FE20BDCD" ID-GBIF-Occurrence="2028147515" collectingDate="1915-01-01" collectingDateMax="1915-01-31" collectingDateMin="1915-01-01" collectionCode="BMNH" collectorName="London" country="Malaysia" elevation="610" location="Selangor-Pahang" pageId="214" pageNumber="215" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="A3E765D09781FF57FACCFD09FA67BDB2" box="[1326,1433,644,669]" name="Malaysia" pageId="214" pageNumber="215">Malaysia</collectingCountry>
:
<location id="DE2F739B9781FF57FF75FD2BFEA7BD90" LSID="urn:lsid:plazi:treatment:535994569781FF56FF75FE3CFE96BD75:DE2F739B9781FF57FF75FD2BFEA7BD90" box="[151,345,678,703]" country="Malaysia" name="Selangor-Pahang" pageId="214" pageNumber="215">Selangor-Pahang</location>
, [Ile Gep] should be “Semangko Pass“,
<quantity id="1C0888A59781FF57FCB3FD2BFC5CBD91" box="[849,930,678,703]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="214" pageNumber="215" unit="m" value="610.0">
<elevation id="50DDC2739781FF57FCB3FD2BFC5CBD91" box="[849,930,678,703]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="214" pageNumber="215" unit="m" value="610.0">610 m</elevation>
</quantity>
,
<date id="AF4E03809781FF57FC57FD2BFBC4BD90" box="[949,1082,678,703]" pageId="214" pageNumber="215" value="1915-01-01" valueMax="1915-01-31" valueMin="1915-01-01">
<collectingDate id="BF0AFA689781FF57FC57FD2BFBC4BD90" box="[949,1082,678,703]" pageId="214" pageNumber="215" value="1915-01-01" valueMax="1915-01-31" valueMin="1915-01-01">131.i.1915</collectingDate>
</date>
<specimenCount id="CDF6EEC99781FF57FBBEFD2BFB54BD90" box="[1116,1194,678,703]" pageId="214" pageNumber="215" type="male">1 male</specimenCount>
(
<typeStatus id="044B9BE29781FF57FB23FD2BFADBBD90" box="[1217,1317,678,703]" pageId="214" pageNumber="215" type="holotype">holotype</typeStatus>
of
<emphasis id="E984F9529781FF57FAB8FD2AFEFBBDCE" italics="true" pageId="214" pageNumber="215">M. a. conjuncta</emphasis>
) (
<collectorName id="760540969781FF57FEFFFD44FE88BDCD" box="[285,374,713,738]" pageId="214" pageNumber="215">London</collectorName>
<collectionCode id="BDE1BD859781FF57FE9FFD47FE28BDCD" box="[381,470,714,738]" pageId="214" pageNumber="215">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB9781FF57FF25FD61FC01BC65" pageId="214" pageNumber="215" type="diagnosis">
<paragraph id="DB4F25409781FF57FF25FD61FC01BC65" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
<emphasis id="E984F9529781FF57FF25FD61FEBFBC2A" bold="true" box="[199,321,748,773]" pageId="214" pageNumber="215">Diagnosis.</emphasis>
<taxonomicName id="1CF05EC39781FF57FEABFD60FE4ABC2B" baseAuthorityName="Walker" baseAuthorityYear="1869" box="[329,436,749,772]" class="Insecta" family="Gryllacrididae" genus="Melanogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="E984F9529781FF57FEABFD60FE4ABC2B" box="[329,436,749,772]" italics="true" pageId="214" pageNumber="215">M. atrata</emphasis>
</taxonomicName>
is characterized by the dark coloration. Tegmen and hind wings are black while most of the cross-veins of tegmen are white. In the female, the short ovipositor with widened base but otherwise slightly down-curved valves is also characteristic for the species and genus (
<figureCitation id="43CB39C59781FF57FC71FCBCFC0DBC65" box="[915,1011,817,842]" captionStart="FIGURE 74" captionStartId="210.[151,250,1874,1896]" captionTargetBox="[151,1436,269,1851]" captionTargetId="figure@210.[151,1436,267,1852]" captionTargetPageId="210" captionText="FIGURE 74. Genus Melaneremus: AD, M. fruhstorferi (Griffini, 1914) male holotype (A), female allotype (B) and male from Hoang Lien (CD), EH, M. sikkimensis sp. nov. holotype; I, M. atrotectus (Brunner, 1888) male holotype.—JK, Melanogryllacris atrata (Walker, 1869) female in NHMW.—Habitus lateral view (AB, IJ), habitus dorsal view (C, E, K), face (D, H), anterior area of body with head, pronotum and thorax (F), lateral view on reduced wings (G)." figureDoi="http://doi.org/10.5281/zenodo.10109164" httpUri="https://zenodo.org/record/10109164/files/figure.png" pageId="214" pageNumber="215">Fig. 74J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB9781FF57FF25FCD9FA87B8CA" pageId="214" pageNumber="215" type="description">
<paragraph id="DB4F25409781FF57FF25FCD9FD0FBCFA" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
<emphasis id="E984F9529781FF57FF25FCD9FEA7BC42" bold="true" box="[199,345,852,877]" pageId="214" pageNumber="215">Description.</emphasis>
Medium sized species. Head: Face ovoid; forehead nearly smooth with faint transverse riffles in lower area; fastigium verticis wider than scapus; ocelli indistinctly separated from fastigium frontis by a curved suture; ocelli little distinct; subocular furrow distinct (
<figureCitation id="43CB39C59781FF57FD1EFC14FC98BC9D" box="[764,870,921,946]" captionStart="FIGURE 76" captionStartId="216.[151,250,1871,1893]" captionTargetBox="[165,1428,193,1847]" captionTargetId="figure@216.[164,1428,193,1849]" captionTargetPageId="216" captionText="FIGURE 76. AJ, Genus Metriogryllacris: AE, M. (M.) amitarum (Griffini, 1914) female syntype (AC), male syntype (D), and male from Tam Dao (E); FG, M. (M.) permodesta (Griffini, 1914) female holotype; HJ, M. (Xiphilarnaca) xiphiura (Karny, 1928) syntype.—KN, Melanogryllacris atrata (Walker, 1869) female from Kinabalu.—Face (A, F, H, N), habitus dorsal view (C, E, I, K), habitus lateral view (B, D, G, J), stridulatory pegs on second and third abdominal tergites, front is left (L), female abdominal apex with ovipositor and subgenital plate (M)." figureDoi="http://doi.org/10.5281/zenodo.10109168" httpUri="https://zenodo.org/record/10109168/files/figure.png" pageId="214" pageNumber="215">Fig. 76N</figureCitation>
). Abdominal tergites two and three each with two rows of stridulatory pegs (2, 6; 7, 19; n = 1;
<figureCitation id="43CB39C59781FF57FD64FC31FD1ABCFA" box="[646,740,956,981]" captionStart="FIGURE 76" captionStartId="216.[151,250,1871,1893]" captionTargetBox="[165,1428,193,1847]" captionTargetId="figure@216.[164,1428,193,1849]" captionTargetPageId="216" captionText="FIGURE 76. AJ, Genus Metriogryllacris: AE, M. (M.) amitarum (Griffini, 1914) female syntype (AC), male syntype (D), and male from Tam Dao (E); FG, M. (M.) permodesta (Griffini, 1914) female holotype; HJ, M. (Xiphilarnaca) xiphiura (Karny, 1928) syntype.—KN, Melanogryllacris atrata (Walker, 1869) female from Kinabalu.—Face (A, F, H, N), habitus dorsal view (C, E, I, K), habitus lateral view (B, D, G, J), stridulatory pegs on second and third abdominal tergites, front is left (L), female abdominal apex with ovipositor and subgenital plate (M)." figureDoi="http://doi.org/10.5281/zenodo.10109168" httpUri="https://zenodo.org/record/10109168/files/figure.png" pageId="214" pageNumber="215">Fig 76L</figureCitation>
).
</paragraph>
<paragraph id="DB4F25409781FF57FF25FC53FE78BA22" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
Wings surpassing abdomen and reaching about midst of stretched hind tibia (
<figureCitation id="43CB39C59781FF57FBC0FC53FB6ABCD8" box="[1058,1172,990,1015]" captionStart="FIGURE 74" captionStartId="210.[151,250,1874,1896]" captionTargetBox="[151,1436,269,1851]" captionTargetId="figure@210.[151,1436,267,1852]" captionTargetPageId="210" captionText="FIGURE 74. Genus Melaneremus: AD, M. fruhstorferi (Griffini, 1914) male holotype (A), female allotype (B) and male from Hoang Lien (CD), EH, M. sikkimensis sp. nov. holotype; I, M. atrotectus (Brunner, 1888) male holotype.—JK, Melanogryllacris atrata (Walker, 1869) female in NHMW.—Habitus lateral view (AB, IJ), habitus dorsal view (C, E, K), face (D, H), anterior area of body with head, pronotum and thorax (F), lateral view on reduced wings (G)." figureDoi="http://doi.org/10.5281/zenodo.10109164" httpUri="https://zenodo.org/record/10109164/files/figure.png" pageId="214" pageNumber="215">Figs. 74K</figureCitation>
,
<figureCitation id="43CB39C59781FF57FB7DFC53FB2ABCD8" box="[1183,1236,990,1015]" captionStart="FIGURE 76" captionStartId="216.[151,250,1871,1893]" captionTargetBox="[165,1428,193,1847]" captionTargetId="figure@216.[164,1428,193,1849]" captionTargetPageId="216" captionText="FIGURE 76. AJ, Genus Metriogryllacris: AE, M. (M.) amitarum (Griffini, 1914) female syntype (AC), male syntype (D), and male from Tam Dao (E); FG, M. (M.) permodesta (Griffini, 1914) female holotype; HJ, M. (Xiphilarnaca) xiphiura (Karny, 1928) syntype.—KN, Melanogryllacris atrata (Walker, 1869) female from Kinabalu.—Face (A, F, H, N), habitus dorsal view (C, E, I, K), habitus lateral view (B, D, G, J), stridulatory pegs on second and third abdominal tergites, front is left (L), female abdominal apex with ovipositor and subgenital plate (M)." figureDoi="http://doi.org/10.5281/zenodo.10109168" httpUri="https://zenodo.org/record/10109168/files/figure.png" pageId="214" pageNumber="215">76K</figureCitation>
). Tegmen: Radius with two branches, both forked near tip; in the male
<typeStatus id="044B9BE29781FF57FD3FFB8FFCF3BB35" box="[733,781,1026,1050]" pageId="214" pageNumber="215">type</typeStatus>
of
<emphasis id="E984F9529781FF57FCD3FB8FFC21BB36" box="[817,991,1026,1049]" italics="true" pageId="214" pageNumber="215">M. a. conjuncta</emphasis>
and in the female from
<collectingRegion id="1934EBA29781FF57FB08FB8CFA8EBB35" box="[1258,1392,1025,1050]" country="Indonesia" name="Kalimantan Utara" pageId="214" pageNumber="215">Kalimantan</collectingRegion>
, RS just after arising from R fuses with MA for about two cell lengths and then separates again, in the female from Kinabalu there is no connection between both veins; media anterior free from base, single branched or fused in about mid-length for about two cells lengths with RS; MP and CuA with common stem, dividing behind basal third, and the anterior branch divides again about at mid-length resulting in three branches, MP, CuA1 and CuA2; cubitus posterior undivided, free throughout; with 4 anal veins. Legs: Hind femur with rather strong spines on both ventral margins, the last internal spine markedly larger than the preceding. Hind tibia with a pre-apical spine on both ventral margins.
</paragraph>
<paragraph id="DB4F25409781FF57FF25FA9BFB5BB90D" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
Coloration. General color black; face, central areas of vertex and pronotum, thoracic sternites, coxae and basal two thirds of all femora brown or of light color. Face yellowish brown, marbled with lighter and darker spots and flecks. Tegmen blackish brown; veins and veinlets in about basal half light greyish, in apical half veins dark brown, veinlets of lighter color; hind wing blackish brown with dark veins and veinlets, partly with light cross-veins. The female from Mt. Kinabalu that had been originally conserved in alcohol is of lighter color: the basic color is medium to dark brown, the pronotum has only the margin black, and the black color of the genicular area of all legs is less extended. The tegmen is medium brown but almost transparent along fore and hind margins. The hind wings have a light brown infumation, dark brown veins and medium brown cross-veins (
<figureCitation id="43CB39C59781FF57FBCDF984FB66B90D" box="[1071,1176,1545,1570]" captionStart="FIGURE 76" captionStartId="216.[151,250,1871,1893]" captionTargetBox="[165,1428,193,1847]" captionTargetId="figure@216.[164,1428,193,1849]" captionTargetPageId="216" captionText="FIGURE 76. AJ, Genus Metriogryllacris: AE, M. (M.) amitarum (Griffini, 1914) female syntype (AC), male syntype (D), and male from Tam Dao (E); FG, M. (M.) permodesta (Griffini, 1914) female holotype; HJ, M. (Xiphilarnaca) xiphiura (Karny, 1928) syntype.—KN, Melanogryllacris atrata (Walker, 1869) female from Kinabalu.—Face (A, F, H, N), habitus dorsal view (C, E, I, K), habitus lateral view (B, D, G, J), stridulatory pegs on second and third abdominal tergites, front is left (L), female abdominal apex with ovipositor and subgenital plate (M)." figureDoi="http://doi.org/10.5281/zenodo.10109168" httpUri="https://zenodo.org/record/10109168/files/figure.png" pageId="214" pageNumber="215">Fig. 76K</figureCitation>
).
</paragraph>
<paragraph id="DB4F25409781FF57FF25F9A1FCC5B982" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
Male. Subgenital plate (
<typeStatus id="044B9BE29781FF57FE3CF9A1FDEEB96B" box="[478,528,1580,1604]" pageId="214" pageNumber="215">type</typeStatus>
of
<emphasis id="E984F9529781FF57FDDFF9A0FD0BB96B" box="[573,757,1581,1604]" italics="true" pageId="214" pageNumber="215">M. a. conjuncta</emphasis>
) wide at base, strongly constricted before mid-length and projecting central area with moderately converging lateral margins; apical margin subtruncte; styli small and thin, inserted at medial constriction of plate, their tips not reaching half the length of the median process. See also extensive descriptions in
<bibRefCitation id="BF6158B19781FF57FE51F919FDA7B983" author="Griffini, A." box="[435,601,1684,1709]" pageId="214" pageNumber="215" pagination="82 - 94" refId="ref186022" refString="Griffini, A. (1910) Revision des types de certaines Gryllacris decrites par F. Walker, existant au Musee d' Oxford (Orth.). Deutsche Entomologische Zeitschrift, 1910, 82 - 94; Berlin." type="journal article" year="1910">Griffini (1910)</bibRefCitation>
and
<bibRefCitation id="BF6158B19781FF57FD72F918FCC9B982" author="Karny, H. H." box="[656,823,1684,1709]" pageId="214" pageNumber="215" pagination="1 - 67" refId="ref187404" refString="Karny, H. H. (1926 a) On Malaysian cricket-locusts (Gryllacridae). Journal of the Federated Malay States Museum, Kuala Lumpur, 13 [1925], 1 - 67, pls 1 - 2." type="journal article" year="1926">Karny (1926a)</bibRefCitation>
.
</paragraph>
<paragraph id="DB4F25409781FF57FF25F93BFDC7B8B0" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
Female. Seventh abdominal sternite larger than preceding sternites, black; in middle of hind margin with a small yellowish modification consisting of a pair of lateral pits with swollen margins and a short medial furrow with swollen lateral margins, open anteriorly and terminating posteriorly in a small elevation. Subgenital plate black, with medial furrow and converging lateral margins, base little excised to fit the modification of the seventh sternite, apex slightly bilobate (
<figureCitation id="43CB39C59781FF57FDE3F8CCFD8FB875" box="[513,625,1857,1882]" captionStart="FIGURE 76" captionStartId="216.[151,250,1871,1893]" captionTargetBox="[165,1428,193,1847]" captionTargetId="figure@216.[164,1428,193,1849]" captionTargetPageId="216" captionText="FIGURE 76. AJ, Genus Metriogryllacris: AE, M. (M.) amitarum (Griffini, 1914) female syntype (AC), male syntype (D), and male from Tam Dao (E); FG, M. (M.) permodesta (Griffini, 1914) female holotype; HJ, M. (Xiphilarnaca) xiphiura (Karny, 1928) syntype.—KN, Melanogryllacris atrata (Walker, 1869) female from Kinabalu.—Face (A, F, H, N), habitus dorsal view (C, E, I, K), habitus lateral view (B, D, G, J), stridulatory pegs on second and third abdominal tergites, front is left (L), female abdominal apex with ovipositor and subgenital plate (M)." figureDoi="http://doi.org/10.5281/zenodo.10109168" httpUri="https://zenodo.org/record/10109168/files/figure.png" pageId="214" pageNumber="215">Fig. 76M</figureCitation>
). Base of ovipositor with oval swelling at very base; basal to subbasal area stout and thick, black. Ovipositor behind subbasal area narrowed, yellowish brown and with concave ventral margin; tip subacute (
<figureCitation id="43CB39C59781FF57FE68F80BFE16B8B0" box="[394,488,1926,1951]" captionStart="FIGURE 74" captionStartId="210.[151,250,1874,1896]" captionTargetBox="[151,1436,269,1851]" captionTargetId="figure@210.[151,1436,267,1852]" captionTargetPageId="210" captionText="FIGURE 74. Genus Melaneremus: AD, M. fruhstorferi (Griffini, 1914) male holotype (A), female allotype (B) and male from Hoang Lien (CD), EH, M. sikkimensis sp. nov. holotype; I, M. atrotectus (Brunner, 1888) male holotype.—JK, Melanogryllacris atrata (Walker, 1869) female in NHMW.—Habitus lateral view (AB, IJ), habitus dorsal view (C, E, K), face (D, H), anterior area of body with head, pronotum and thorax (F), lateral view on reduced wings (G)." figureDoi="http://doi.org/10.5281/zenodo.10109164" httpUri="https://zenodo.org/record/10109164/files/figure.png" pageId="214" pageNumber="215">Fig. 74J</figureCitation>
,
<figureCitation id="43CB39C59781FF57FE16F80BFDD3B8B0" box="[500,557,1926,1951]" captionStart="FIGURE 76" captionStartId="216.[151,250,1871,1893]" captionTargetBox="[165,1428,193,1847]" captionTargetId="figure@216.[164,1428,193,1849]" captionTargetPageId="216" captionText="FIGURE 76. AJ, Genus Metriogryllacris: AE, M. (M.) amitarum (Griffini, 1914) female syntype (AC), male syntype (D), and male from Tam Dao (E); FG, M. (M.) permodesta (Griffini, 1914) female holotype; HJ, M. (Xiphilarnaca) xiphiura (Karny, 1928) syntype.—KN, Melanogryllacris atrata (Walker, 1869) female from Kinabalu.—Face (A, F, H, N), habitus dorsal view (C, E, I, K), habitus lateral view (B, D, G, J), stridulatory pegs on second and third abdominal tergites, front is left (L), female abdominal apex with ovipositor and subgenital plate (M)." figureDoi="http://doi.org/10.5281/zenodo.10109168" httpUri="https://zenodo.org/record/10109168/files/figure.png" pageId="214" pageNumber="215">76M</figureCitation>
).
</paragraph>
<paragraph id="DB4F25409781FF57FF25F827FA87B8CA" blockId="214.[151,1437,540,2021]" pageId="214" pageNumber="215">
Measurements (
<specimenCount id="CDF6EEC99781FF57FE62F827FE1DB8ED" box="[384,483,1961,1986]" pageId="214" pageNumber="215" type="female">1 female</specimenCount>
from Kinabalu).—body w/wings: 36; body w/o wings: 29; pronotum: 6; tegmen: 27.5; hind femur: 15.5; tegmen width: 10.5; antenna: 110; ovipositor:
<quantity id="1C0888A59781FF57FC45F841FBFEB8CB" box="[935,1024,1996,2021]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="214" pageNumber="215" unit="mm" value="7.5">7.5 mm</quantity>
. For variation see
<bibRefCitation id="BF6158B19781FF57FB2FF840FA8BB8CA" author="Karny, H. H." box="[1229,1397,1996,2021]" pageId="214" pageNumber="215" pagination="1 - 67" refId="ref187404" refString="Karny, H. H. (1926 a) On Malaysian cricket-locusts (Gryllacridae). Journal of the Federated Malay States Museum, Kuala Lumpur, 13 [1925], 1 - 67, pls 1 - 2." type="journal article" year="1926">Karny (1926a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="93EA76CB9780FF56FF25FF1AFE96BD75" pageId="215" pageNumber="216" type="discussion">
<paragraph id="DB4F25409780FF56FF25FF1AFA62BD3C" blockId="215.[151,1437,151,603]" pageId="215" pageNumber="216">
<emphasis id="E984F9529780FF56FF25FF1AFEB2BF9F" bold="true" box="[199,332,151,176]" pageId="215" pageNumber="216">Discussion.</emphasis>
<taxonomicName id="1CF05EC39780FF56FEB1FF15FDE8BF9F" authorityName="Karny" authorityYear="1937" box="[339,534,152,176]" class="Insecta" family="Gryllacrididae" genus="Melanogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="E984F9529780FF56FEB1FF15FDE8BF9F" box="[339,534,152,176]" italics="true" pageId="215" pageNumber="216">Melanogryllacris</emphasis>
</taxonomicName>
is so far a monotypic genus. However, two varieties have been named by
<bibRefCitation id="BF6158B19780FF56FAB4FF15FF1ABFFB" author="Karny, H. H." pageId="215" pageNumber="216" pagination="1 - 67" refId="ref187404" refString="Karny, H. H. (1926 a) On Malaysian cricket-locusts (Gryllacridae). Journal of the Federated Malay States Museum, Kuala Lumpur, 13 [1925], 1 - 67, pls 1 - 2." type="journal article" year="1926">Karny (1926a</bibRefCitation>
and
<bibRefCitation id="BF6158B19780FF56FEC1FF31FE95BFFB" author="Karny, H. H." box="[291,363,188,212]" pageId="215" pageNumber="216" pagination="203 - 225" refId="ref187689" refString="Karny, H. H. (1928 e) Vorlaufige Mitteilung uber die wissenschaftlichen Ergebnisse meines Europa-Urlaubs. Entomologische Mitteilungen, Berlin-Dahlem, 17, 60 - 76, 203 - 225." type="journal article" year="1928">1928e</bibRefCitation>
). Since they were named before 1961, they are currently treated as subspecies (OSF 2017). However Karny's (1926) text suggests, that he intended to name an entity below the subspecies level. He explicitly mentions that he did not find any difference between the three males of
<taxonomicName id="1CF05EC39780FF56FC51FE8EFBE2BE35" baseAuthorityName="Walker" baseAuthorityYear="1869" box="[947,1052,259,282]" class="Insecta" family="Gryllacrididae" genus="Melanogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="E984F9529780FF56FC51FE8EFBE2BE35" box="[947,1052,259,282]" italics="true" pageId="215" pageNumber="216">M. atrata</emphasis>
</taxonomicName>
he studied, except that in the male that he named
<taxonomicName id="1CF05EC39780FF56FEA1FEAAFE1BBE11" box="[323,485,295,318]" pageId="215" pageNumber="216" rank="variety" variety="conjuncta">
var.
<emphasis id="E984F9529780FF56FE94FEAAFE1BBE11" box="[374,485,295,318]" italics="true" pageId="215" pageNumber="216">conjuncta</emphasis>
</taxonomicName>
the media (= media anterior as used in this paper) fused for a short distance with radius sector while in the other two males that he later (
<bibRefCitation id="BF6158B19780FF56FD1FFEC4FC6EBE4E" author="Karny, H. H." box="[765,912,329,353]" pageId="215" pageNumber="216" pagination="203 - 225" refId="ref187689" refString="Karny, H. H. (1928 e) Vorlaufige Mitteilung uber die wissenschaftlichen Ergebnisse meines Europa-Urlaubs. Entomologische Mitteilungen, Berlin-Dahlem, 17, 60 - 76, 203 - 225." type="journal article" year="1928">Karny 1928e</bibRefCitation>
) named
<taxonomicName id="1CF05EC39780FF56FC12FEC7FB78BE4E" box="[1008,1158,329,353]" pageId="215" pageNumber="216" rank="variety" variety="disjuncta">
var.
<emphasis id="E984F9529780FF56FBFDFEC4FB78BE4E" box="[1055,1158,329,353]" italics="true" pageId="215" pageNumber="216">disjuncta</emphasis>
</taxonomicName>
, the media was free over the whole length. Regarding the individual variability with regard to this character in
<taxonomicName id="1CF05EC39780FF56FB85FEE1FAF2BEAA" authorityName="Blanchard" authorityYear="1845" box="[1127,1292,364,389]" class="Insecta" family="Gryllacrididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="family">Gryllacrididae</taxonomicName>
, I would not pose much importance on this character. However, a re-investigation of the specimens concerned and including Walkers'
<typeStatus id="044B9BE29780FF56FF1DFE39FED1BEE3" box="[255,303,436,460]" pageId="215" pageNumber="216">type</typeStatus>
from
<collectingCountry id="A3E765D09780FF56FE94FE3EFE27BEE3" box="[374,473,435,460]" name="Malaysia" pageId="215" pageNumber="216">Sarawak</collectingCountry>
, with emphasis on a detailed examination of the male abdominal appendages, would be necessary to settle this question and eventually draw any taxonomic consequences. It would also be necessary to re-investigate Walkers'
<typeStatus id="044B9BE29780FF56FE7FFE76FE33BD3C" box="[413,461,507,531]" pageId="215" pageNumber="216">type</typeStatus>
of
<taxonomicName id="1CF05EC39780FF56FE11FE76FDA2BD3D" baseAuthorityName="Walker" baseAuthorityYear="1869" box="[499,604,507,530]" class="Insecta" family="Gryllacrididae" genus="Melanogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="E984F9529780FF56FE11FE76FDA2BD3D" box="[499,604,507,530]" italics="true" pageId="215" pageNumber="216">M. atrata</emphasis>
</taxonomicName>
for the venation, as that character has not been studied by
<bibRefCitation id="BF6158B19780FF56FB0CFE77FA69BD3C" author="Griffini, A." box="[1262,1431,506,531]" pageId="215" pageNumber="216" pagination="82 - 94" refId="ref186022" refString="Griffini, A. (1910) Revision des types de certaines Gryllacris decrites par F. Walker, existant au Musee d' Oxford (Orth.). Deutsche Entomologische Zeitschrift, 1910, 82 - 94; Berlin." type="journal article" year="1910">Griffini (1910)</bibRefCitation>
.
</paragraph>
<paragraph id="DB4F25409780FF56FF25FD90FE96BD75" blockId="215.[151,1437,151,603]" pageId="215" pageNumber="216">
The two females studied agree in general characters with the redescription of the male
<typeStatus id="044B9BE29780FF56FB72FD90FB0CBD19" box="[1168,1266,541,566]" pageId="215" pageNumber="216" type="holotype">holotype</typeStatus>
of
<emphasis id="E984F9529780FF56FAF4FD92FA65BD19" box="[1302,1435,543,566]" italics="true" pageId="215" pageNumber="216">
M. a.
<taxonomicName id="1CF05EC39780FF56FAB5FD92FA65BD19" baseAuthorityName="Walker" baseAuthorityYear="1869" box="[1367,1435,543,566]" class="Insecta" family="Gryllacrididae" genus="Melanogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="species" species="atrata">atrata</taxonomicName>
</emphasis>
by
<bibRefCitation id="BF6158B19780FF56FF58FDCFFE9DBD75" author="Griffini, A." box="[186,355,578,603]" pageId="215" pageNumber="216" pagination="82 - 94" refId="ref186022" refString="Griffini, A. (1910) Revision des types de certaines Gryllacris decrites par F. Walker, existant au Musee d' Oxford (Orth.). Deutsche Entomologische Zeitschrift, 1910, 82 - 94; Berlin." type="journal article" year="1910">Griffini (1910)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>