257 lines
42 KiB
XML
257 lines
42 KiB
XML
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<mods:title id="F65AAB4CE607F2E6B35CDA312C21339F">New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera</mods:title>
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<mods:name id="CABB221875B93521963B6F7AF9F0A379" type="personal">
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<mods:namePart id="2962A4D162EEFC197AE258A5724F44D2">Ingrisch, Sigfrid</mods:namePart>
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<mods:title id="5225AA210E01116FA3AF821C2E7FC3F1">Zootaxa</mods:title>
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<mods:part id="F5F1733BC8BFA4A10BD7F8AC581916EE">
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<mods:date id="6FAD4B6A58BBEEBE744C46D2069B78DC">2018</mods:date>
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<mods:number id="81EB57E0857BB42FE19040CC2DEF2426">2018-11-05</mods:number>
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<mods:number id="133A8C56B435AB1627ACD5900BE5C27F">4510</mods:number>
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<mods:identifier id="5D9D8FE5F1E26F1977E6ECB0A93E8366" type="DOI">10.11646/zootaxa.4510.1.1</mods:identifier>
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<treatment id="535994569773FFA4FF75FBB2FE58BB77" ID-DOI="http://doi.org/10.5281/zenodo.5986916" ID-GBIF-Taxon="154497797" ID-Zenodo-Dep="5986916" LSID="urn:lsid:plazi:treatment:535994569773FFA4FF75FBB2FE58BB77" httpUri="http://treatment.plazi.org/id/535994569773FFA4FF75FBB2FE58BB77" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
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<subSubSection id="93EA76CB9773FFA5FF75FBB2FDE5BB75" box="[151,539,1087,1114]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="DB4F25409773FFA5FF75FBB2FDE5BB75" blockId="36.[151,585,1087,1147]" box="[151,539,1087,1114]" pageId="36" pageNumber="37">
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<heading id="8007922C9773FFA5FF75FBB2FDE5BB75" bold="true" box="[151,539,1087,1114]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
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<emphasis id="E984F9529773FFA5FF75FBB2FDE5BB75" bold="true" box="[151,539,1087,1114]" pageId="36" pageNumber="37">
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<taxonomicName id="1CF05EC39773FFA5FF75FBB2FE48BB76" ID-CoL="8NRKS" authority="Ingrisch, 2018" authorityName="Ingrisch" authorityYear="2018" box="[151,438,1087,1113]" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="E984F9529773FFA5FF75FBB2FE48BB76" bold="true" box="[151,438,1087,1113]" italics="true" pageId="36" pageNumber="37">Eugryllacris trabicauda</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F2B744299773FFA5FE5FFBCDFDE5BB75" box="[445,539,1088,1114]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="93EA76CB9773FFA5FF75FBEFFDB7BB54" box="[151,585,1122,1147]" pageId="36" pageNumber="37" type="description">
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<paragraph id="DB4F25409773FFA5FF75FBEFFDB7BB54" blockId="36.[151,585,1087,1147]" box="[151,585,1122,1147]" pageId="36" pageNumber="37">
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<figureCitation id="43CB39C59773FFA5FF75FBEFFF09BB54" box="[151,247,1122,1147]" captionStart="FIGURE 1" captionStartId="12.[151,250,1761,1783]" captionTargetBox="[153,1439,198,1734]" captionTargetId="figure@12.[148,1440,193,1740]" captionTargetPageId="12" captionText="FIGURE 1. A, Larnaca (L.) eugenii (Griffini, 1914); B, Eugryllacris trabicauda sp. nov.; C, Ocellarnaca braueri (Griffini, 1911); D, Gryllacris kinabaluensis Griffini, 1915; E, Ultragryllacris triangula sp. nov.; F, Woznessenskia bavi sp. nov.; G, Psilogryllacris sp. (Karny's 1931 "allotype" of "dalbertisi Griffini 1909"); H, Zalarnaca (Zalarnaca) maninjau sp. nov.—Left tegmen with main veins (A–G), apical area of right hind tibia and hind tarsus in oblique lateral view with ventral pre-apical spine marked (H). Abbreviations for veins: C costa, ScA subcosta anterior, ScP subcosta posterior, R radius, RS radius sector, MA media anterior, MP media posterior, CuA cubitus anterior, CuP cubitus posterior, An1 first anal vein, following anal veins unmarked." figureDoi="http://doi.org/10.5281/zenodo.10109014" httpUri="https://zenodo.org/record/10109014/files/figure.png" pageId="36" pageNumber="37">Figs. 1B</figureCitation>
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,
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<figureCitation id="43CB39C59773FFA5FEE6FBEEFEECBB54" box="[260,274,1123,1147]" captionStart="FIGURE 4" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[151,1437,195,1894]" captionTargetId="figure@27.[149,1438,193,1894]" captionTargetPageId="27" captionText="FIGURE 4. Genus Eugryllacris: Habitus dorsal view: A–B, E. trabicauda sp. nov. female holotype (A) and male paratype (B); C–D, E. vermicauda sp. nov. female holotype (C) male paratype (D); E, E. serricauda sp. nov. male holotype; F, E. guomashan sp. nov. female holotype; G, E. crassicauda sp. nov. female paratype; H, E. crassicauda cambodiana ssp. nov. female holotype; I, E. malaccensis (Griffini, 1908) male; J, E. serricauda sp. nov. female paratype. Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10109022" httpUri="https://zenodo.org/record/10109022/files/figure.png" pageId="36" pageNumber="37">4</figureCitation>
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A–B, 5H, 6A–B, 8A–B, 9O
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</paragraph>
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||
</subSubSection>
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<subSubSection id="93EA76CB9773FFA5FF75FB27FBA6BA00" pageId="36" pageNumber="37" type="materials_examined">
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||
<paragraph id="DB4F25409773FFA5FF75FB27FAD9BBC8" blockId="36.[151,1437,1194,2011]" pageId="36" pageNumber="37">
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||
<emphasis id="E984F9529773FFA5FF75FB27FE77BBEC" bold="true" box="[151,393,1194,1219]" pageId="36" pageNumber="37">Material examined.</emphasis>
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||
<materialsCitation id="6B982F1D9773FFA5FE7AFB27FAD9BBC8" ID-GBIF-Occurrence="2028147453" collectingDate="2011-07-19" collectingDateMax="2011-07-23" collectingDateMin="2011-07-19" collectionCode="RBINS" collectorName="J. Constant & J. Bresseel" country="Vietnam" latitude="20.316668" location="Cuc Phuong National Park" longLatPrecision="925" longitude="105.60833" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Hoa Binh" typeStatus="holotype">
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||
<typeStatus id="044B9BE29773FFA5FE7AFB27FDFEBBEC" box="[408,512,1194,1219]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
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(female):
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<collectingCountry id="A3E765D09773FFA5FD65FB27FD14BBEC" box="[647,746,1194,1219]" name="Vietnam" pageId="36" pageNumber="37">Vietnam</collectingCountry>
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||
:
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<collectingRegion id="1934EBA29773FFA5FD1FFB26FC8FBBEC" box="[765,881,1194,1219]" country="Vietnam" name="Hoa Binh" pageId="36" pageNumber="37">Hoa Binh</collectingRegion>
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Prov.,
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<location id="DE2F739B9773FFA5FC30FB26FAE5BBEC" LSID="urn:lsid:plazi:treatment:535994569773FFA4FF75FBB2FE58BB77:DE2F739B9773FFA5FC30FB26FAE5BBEC" box="[978,1307,1194,1219]" country="Vietnam" latitude="20.316668" longLatPrecision="925" longitude="105.60833" name="Cuc Phuong National Park" pageId="36" pageNumber="37" stateProvince="Hoa Binh">Cuc Phuong National Park</location>
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, (
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||
<geoCoordinate id="BEC443879773FFA5FADAFB26FA69BBEC" box="[1336,1431,1195,1219]" direction="north" orientation="latitude" pageId="36" pageNumber="37" precision="925" value="20.316668">20°19'N</geoCoordinate>
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||
,
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||
<geoCoordinate id="BEC443879773FFA5FF75FB43FED9BBC8" box="[151,295,1230,1255]" direction="east" orientation="longitude" pageId="36" pageNumber="37" precision="15" value="105.60833">105°36'30''E</geoCoordinate>
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),
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||
<date id="AF4E03809773FFA5FEDEFB43FE19BBC8" box="[316,487,1230,1255]" pageId="36" pageNumber="37" value="2011-07-19" valueMax="2011-07-23" valueMin="2011-07-19">
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||
<collectingDate id="BF0AFA689773FFA5FEDEFB43FE19BBC8" box="[316,487,1230,1255]" pageId="36" pageNumber="37" value="2011-07-19" valueMax="2011-07-23" valueMin="2011-07-19">19–23.vii.2011</collectingDate>
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||
</date>
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, leg.
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<collectorName id="760540969773FFA5FDC0FB42FD61BBC8" box="[546,671,1231,1255]" pageId="36" pageNumber="37">J. Constant</collectorName>
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&
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<collectorName id="760540969773FFA5FD21FB42FCC4BBC8" box="[707,826,1230,1255]" pageId="36" pageNumber="37">J. Bresseel</collectorName>
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(I.G. 31.933)—
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<specimenCount id="CDF6EEC99773FFA5FC13FB42FBB3BBC8" box="[1009,1101,1230,1255]" pageId="36" pageNumber="37" type="female">1 female</specimenCount>
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||
(Brussels
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<collectionCode id="BDE1BD859773FFA5FB26FB42FAE5BBC8" box="[1220,1307,1231,1255]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="37" type="Museum">RBINS</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="DB4F25409773FFA5FF25FB7FFBA6BA00" blockId="36.[151,1437,1194,2011]" pageId="36" pageNumber="37">
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||
Other specimens: same data as
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<typeStatus id="044B9BE29773FFA5FDCBFB7FFD70BA24" box="[553,654,1266,1291]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="CDF6EEC99773FFA5FD7BFB7EFD18BA24" box="[665,742,1266,1291]" pageId="36" pageNumber="37" type="male">1 male</specimenCount>
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||
(
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<typeStatus id="044B9BE29773FFA5FD17FB7EFCA7BA24" box="[757,857,1267,1291]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
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) (
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<collectingRegion id="1934EBA29773FFA5FC92FB7FFC2EBA24" box="[880,976,1266,1291]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="36" pageNumber="37">Brussels</collectingRegion>
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RBINS);
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<materialsCitation id="6B982F1D9773FFA5FBA1FB7EFBAABA00" ID-GBIF-Occurrence="2028147482" collectingDate="2010-08-11" collectingDateMax="2010-08-18" collectingDateMin="2010-08-11" collectionCode="RBINS" collectorName="J. Constant & P. Limbourg" pageId="36" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Region de Bruxelles-Capitale" typeStatus="paratype">
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same locality,
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<date id="AF4E03809773FFA5FB07FB7FFA66BA24" box="[1253,1432,1266,1291]" pageId="36" pageNumber="37" value="2010-08-11" valueMax="2010-08-18" valueMin="2010-08-11">
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||
<collectingDate id="BF0AFA689773FFA5FB07FB7FFA66BA24" box="[1253,1432,1266,1291]" pageId="36" pageNumber="37" value="2010-08-11" valueMax="2010-08-18" valueMin="2010-08-11">11–18.viii.2010</collectingDate>
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</date>
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, leg.
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<collectorName id="760540969773FFA5FF25FA9AFEBABA00" box="[199,324,1303,1327]" pageId="36" pageNumber="37">J. Constant</collectorName>
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&
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<collectorName id="760540969773FFA5FE85FA9AFE0CBA00" box="[359,498,1302,1327]" pageId="36" pageNumber="37">P. Limbourg</collectorName>
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||
(I.G. 31.668)—
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<specimenCount id="CDF6EEC99773FFA5FD4AFA9BFCFABA00" box="[680,772,1302,1327]" pageId="36" pageNumber="37" type="female">1 female</specimenCount>
|
||
(
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<typeStatus id="044B9BE29773FFA5FCF1FA9AFC89BA00" box="[787,887,1303,1327]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
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) (
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<collectingRegion id="1934EBA29773FFA5FC6FFA9BFC10BA00" box="[909,1006,1302,1327]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="36" pageNumber="37">Brussels</collectingRegion>
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<collectionCode id="BDE1BD859773FFA5FC17FA9AFBB2BA00" box="[1013,1100,1303,1327]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="37" type="Museum">RBINS</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="93EA76CB9773FFA5FF25FAB7FBF3B994" pageId="36" pageNumber="37" type="diagnosis">
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||
<paragraph id="DB4F25409773FFA5FF25FAB7FBF3B994" blockId="36.[151,1437,1194,2011]" pageId="36" pageNumber="37">
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||
<emphasis id="E984F9529773FFA5FF25FAB7FEBFBA7C" bold="true" box="[199,321,1338,1363]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
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With regard to the median process of the female seventh abdominal sternite, the new species resembles
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<taxonomicName id="1CF05EC39773FFA5FEEFFAD2FDA2BA58" authority="(Karny, 1926)" baseAuthorityName="Karny" baseAuthorityYear="1926" box="[269,604,1374,1399]" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cylindrigera">
|
||
<emphasis id="E984F9529773FFA5FEEFFAD2FE49BA59" box="[269,439,1374,1398]" italics="true" pageId="36" pageNumber="37">E. cylindrigera</emphasis>
|
||
(Karny, 1926)
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</taxonomicName>
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,
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<taxonomicName id="1CF05EC39773FFA5FD8AFAD2FC49BA58" authority="Bian & Shi, 2016" authorityName="Bian & Shi" authorityYear="2016" box="[616,951,1374,1399]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="elongata">
|
||
<emphasis id="E984F9529773FFA5FD8AFAD2FD17BA59" box="[616,745,1374,1398]" italics="true" pageId="36" pageNumber="37">E. elongata</emphasis>
|
||
Bian &
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<bibRefCitation id="BF6158B19773FFA5FCAAFAD3FC49BA58" author="Shi & Bian X. & Guo L. Y." box="[840,951,1374,1399]" pageId="36" pageNumber="37" pagination="1 - 8" refId="ref189328" refString="Shi F. M., Bian X. & Guo L. Y. (2016) First record of the genus Dialarnaca Gorochov, 2005 from China, with description of two new species (Orthoptera, Gryllacrididae, Gryllacridinae). ZooKeys, 558, 1 - 8. https: // doi. org / 10.3897 / zookeys. 482.8713" type="journal article" year="2016">Shi, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1CF05EC39773FFA5FC23FAD2FAE4BA58" authority="Bian & Shi, 2016" authorityName="Bian & Shi" authorityYear="2016" box="[961,1306,1374,1399]" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="longifissa">
|
||
<emphasis id="E984F9529773FFA5FC23FAD2FBB3BA59" box="[961,1101,1374,1398]" italics="true" pageId="36" pageNumber="37">E. longifissa</emphasis>
|
||
Bian &
|
||
<bibRefCitation id="BF6158B19773FFA5FB4EFAD3FAE4BA58" author="Shi & Bian X. & Guo L. Y." box="[1196,1306,1374,1399]" pageId="36" pageNumber="37" pagination="1 - 8" refId="ref189328" refString="Shi F. M., Bian X. & Guo L. Y. (2016) First record of the genus Dialarnaca Gorochov, 2005 from China, with description of two new species (Orthoptera, Gryllacrididae, Gryllacridinae). ZooKeys, 558, 1 - 8. https: // doi. org / 10.3897 / zookeys. 482.8713" type="journal article" year="2016">Shi, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="1CF05EC39773FFA5FAB6FAD2FE93BAB4" authority="Bian & Shi, 2016" authorityName="Bian & Shi" authorityYear="2016" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="xiei">
|
||
<emphasis id="E984F9529773FFA5FAB6FAD2FA62BA59" box="[1364,1436,1375,1398]" italics="true" pageId="36" pageNumber="37">E. xiei</emphasis>
|
||
Bian &
|
||
<bibRefCitation id="BF6158B19773FFA5FF16FA0FFE93BAB4" author="Shi & Bian X. & Guo L. Y." box="[244,365,1410,1435]" pageId="36" pageNumber="37" pagination="1 - 8" refId="ref189328" refString="Shi F. M., Bian X. & Guo L. Y. (2016) First record of the genus Dialarnaca Gorochov, 2005 from China, with description of two new species (Orthoptera, Gryllacrididae, Gryllacridinae). ZooKeys, 558, 1 - 8. https: // doi. org / 10.3897 / zookeys. 482.8713" type="journal article" year="2016">Shi, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
, all from
|
||
<collectingCountry id="A3E765D09773FFA5FE3CFA0FFDDBBAB4" box="[478,549,1410,1435]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
|
||
. It differs by the narrow but distinct apical excision of that process and more strikingly by the very long and straight ovipositor while the species compared have a much shorter and upcurved ovipositor. Such a long and straight ovipositor is so far only known within the genus from
|
||
<taxonomicName id="1CF05EC39773FFA5FB46FA46FEB3B928" authority="(Pictet & Saussure, 1891)" baseAuthorityName="Pictet & Saussure" baseAuthorityYear="1891" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="vaginalis">
|
||
<emphasis id="E984F9529773FFA5FB46FA46FAD2BACC" box="[1188,1324,1483,1507]" italics="true" pageId="36" pageNumber="37">E. vaginalis</emphasis>
|
||
(
|
||
<bibRefCitation id="BF6158B19773FFA5FADEFA47FEBBB928" author="Pictet, A. & Saussure, H. de" pageId="36" pageNumber="37" pagination="1888 - 1893" refId="ref189136" refString="Pictet, A. & Saussure, H. de (1891 - 2) De quelques Orthopteres nouveaux. Bulletin de la Societe Entomologique Suisse, 8 [1888 - 1893], 293 - 318, pls 1 - 2." type="journal article" year="1891">Pictet & Saussure, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="A3E765D09773FFA5FE78FA63FE2DB928" box="[410,467,1518,1543]" name="India" pageId="36" pageNumber="37">India</collectingCountry>
|
||
and
|
||
<taxonomicName id="1CF05EC39773FFA5FDF3FA62FD45B929" authority="Ingrisch, 2018" authorityName="Ingrisch" authorityYear="2018" box="[529,699,1518,1542]" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crassicauda" status="sp. nov.">
|
||
<emphasis id="E984F9529773FFA5FDF3FA62FD45B929" box="[529,699,1518,1542]" italics="true" pageId="36" pageNumber="37">E. crassicauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E984F9529773FFA5FD24FA63FCDAB928" bold="true" box="[710,804,1518,1543]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="F2B744299773FFA5FD24FA63FCDAB928" box="[710,804,1518,1543]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<collectingCountry id="A3E765D09773FFA5FC8DFA63FC2DB928" box="[879,979,1518,1543]" name="Thailand" pageId="36" pageNumber="37">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="A3E765D09773FFA5FBF3FA63FB79B928" box="[1041,1159,1518,1543]" name="Cambodia" pageId="36" pageNumber="37">Cambodia</collectingCountry>
|
||
,
|
||
<taxonomicName id="1CF05EC39773FFA5FB74FA62FACAB929" authority="Ingrisch, 2018" authorityName="Ingrisch" authorityYear="2018" box="[1174,1332,1518,1542]" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="trabicauda" status="sp. nov.">
|
||
<emphasis id="E984F9529773FFA5FB74FA62FACAB929" box="[1174,1332,1518,1542]" italics="true" pageId="36" pageNumber="37">E. trabicauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E984F9529773FFA5FADCFA63FA62B928" bold="true" box="[1342,1436,1518,1543]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="F2B744299773FFA5FADCFA63FA62B928" box="[1342,1436,1518,1543]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from both by the uncolored, subtransparent hind wings, an even longer ovipositor (
|
||
<figureCitation id="43CB39C59773FFA5FB72F99FFB14B904" box="[1168,1258,1554,1579]" captionStart="FIGURE 9" captionStartId="33.[151,250,1827,1849]" captionTargetBox="[150,1436,197,1806]" captionTargetId="figure@33.[149,1438,195,1806]" captionTargetPageId="33" captionText="FIGURE 9. Genus Eugryllacris: A, E. inversa sp. nov. male holotype; B, E. maculipennis (Stål, 1877) det. Brunner female; C, E. maculipennis specularis (Karny, 1925) female type; D, E. gandaki sp. nov. female holotype; E, E. malaccensis (Griffini, 1908) male; F, E. crassicauda cambodiana ssp. nov. holotype; G, E. vermicauda sp. nov. holotype; H, E. inversa sp. nov. paratype; I, E. gandaki sp. nov.; J, E. sulcata sp. nov. holotype; K, E. crassicauda sp. nov. paratype; L, E. serricauda sp. nov. paratype; M, E. guomashan sp. nov. holotype; N, E. poultoniana (Griffini, 1909); O, E. trabicauda sp. nov. holotype.— Habitus in dorsal view (A–E) and lateral view (F), ovipositor (G–O). Scales 10 mm; images of ovipositor adapted to scale at bottom." figureDoi="http://doi.org/10.5281/zenodo.10109032" httpUri="https://zenodo.org/record/10109032/files/figure.png" pageId="36" pageNumber="37">Fig. 9O</figureCitation>
|
||
), the subgenital plate being distinctly excised at tip and the shape of the seventh abdominal sternite (
|
||
<figureCitation id="43CB39C59773FFA5FB69F9BBFB22B960" box="[1163,1244,1590,1615]" captionStart="FIGURE 8" captionStartId="31.[151,250,1576,1598]" captionTargetBox="[151,1437,195,1554]" captionTargetId="figure@31.[148,1439,193,1555]" captionTargetPageId="31" captionText="FIGURE 8. Genus Eugryllacris: A–B, E. trabicauda sp. nov. holotype; C–D, E. vermicauda sp. nov. holotype; E–F, E. sulcata sp. nov. holotype; G–H, E. guomashan sp. nov. holotype.—Apical area of female abdomen with seventh abdominal sternite, subgenital plate and base of ovipositor in lateral (A, C, E, G) and ventral view (B, D, F, H)." figureDoi="http://doi.org/10.5281/zenodo.10109030" httpUri="https://zenodo.org/record/10109030/files/figure.png" pageId="36" pageNumber="37">Figs. 8</figureCitation>
|
||
A–B). The male abdominal apex of
|
||
<taxonomicName id="1CF05EC39773FFA5FE9BF9D6FDE8B95C" authority="Ingrisch, 2018" authorityName="Ingrisch" authorityYear="2018" box="[377,534,1627,1651]" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="trabicauda" status="sp. nov.">
|
||
<emphasis id="E984F9529773FFA5FE9BF9D6FDE8B95C" box="[377,534,1627,1651]" italics="true" pageId="36" pageNumber="37">E. trabicauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E984F9529773FFA5FDC3F9D7FD80B95C" bold="true" box="[545,638,1626,1651]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="F2B744299773FFA5FDC3F9D7FD80B95C" box="[545,638,1626,1651]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to that of
|
||
<taxonomicName id="1CF05EC39773FFA5FC96F9D6FBE4B95C" authority="Ingrisch, 2018" authorityName="Ingrisch" authorityYear="2018" box="[884,1050,1627,1651]" class="Insecta" family="Gryllacrididae" genus="Eugryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="vermicauda" status="sp. nov.">
|
||
<emphasis id="E984F9529773FFA5FC96F9D6FBE4B95C" box="[884,1050,1627,1651]" italics="true" pageId="36" pageNumber="37">E. vermicauda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E984F9529773FFA5FBC7F9D7FB7CB95C" bold="true" box="[1061,1154,1626,1651]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="F2B744299773FFA5FBC7F9D7FB7CB95C" box="[1061,1154,1626,1651]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
It differs by the pair of projections in the middle of the split apical margin of the ninth tergite below the obtuse bumps, which are shorter and conical (
|
||
<figureCitation id="43CB39C59773FFA5FEC4F92FFE82B994" box="[294,380,1698,1723]" captionStart="FIGURE 6" captionStartId="29.[151,250,1580,1602]" captionTargetBox="[162,1435,185,1560]" captionTargetId="figure@29.[158,1437,185,1560]" captionTargetPageId="29" captionText="FIGURE 6. Genus Eugryllacris: A–B, E. trabicauda sp. nov. paratype; C–D, E. vermicauda sp. nov. paratype; E–F, E. serricauda sp. nov. holotype; G–I, E. panteli (Bolivar, 1900) male type; J–K, E. malaccensis (Griffini, 1908); L–M, E. inversa sp. nov. holotype.—Male abdominal apex in about ventral (A, C, H, K, L), apical (E, J, M), and oblique lateral view (G, on both specimens studied distorted), tip of ninth abdominal tergite with projections in higher magnification (B, D, F), subgenital plate ventral view (H) and dorsal view (I: process of subgenital plate terminates into a down curved acute tip; tip of process vertical to disc of plate). Scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10109026" httpUri="https://zenodo.org/record/10109026/files/figure.png" pageId="36" pageNumber="37">Fig. 6B</figureCitation>
|
||
) instead of compressed, longer and little curved (
|
||
<figureCitation id="43CB39C59773FFA5FC45F92FFBFFB994" box="[935,1025,1698,1723]" captionStart="FIGURE 6" captionStartId="29.[151,250,1580,1602]" captionTargetBox="[162,1435,185,1560]" captionTargetId="figure@29.[158,1437,185,1560]" captionTargetPageId="29" captionText="FIGURE 6. Genus Eugryllacris: A–B, E. trabicauda sp. nov. paratype; C–D, E. vermicauda sp. nov. paratype; E–F, E. serricauda sp. nov. holotype; G–I, E. panteli (Bolivar, 1900) male type; J–K, E. malaccensis (Griffini, 1908); L–M, E. inversa sp. nov. holotype.—Male abdominal apex in about ventral (A, C, H, K, L), apical (E, J, M), and oblique lateral view (G, on both specimens studied distorted), tip of ninth abdominal tergite with projections in higher magnification (B, D, F), subgenital plate ventral view (H) and dorsal view (I: process of subgenital plate terminates into a down curved acute tip; tip of process vertical to disc of plate). Scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10109026" httpUri="https://zenodo.org/record/10109026/files/figure.png" pageId="36" pageNumber="37">Fig. 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93EA76CB9773FFA4FF25F94BFB12BB20" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="DB4F25409773FFA5FF25F94BFEA4B840" blockId="36.[151,1437,1194,2011]" pageId="36" pageNumber="37">
|
||
<emphasis id="E984F9529773FFA5FF25F94BFEA7B9F0" bold="true" box="[199,345,1734,1759]" pageId="36" pageNumber="37">Description.</emphasis>
|
||
Large species (
|
||
<figureCitation id="43CB39C59773FFA5FDEFF94BFDA0B9F0" box="[525,606,1734,1759]" captionStart="FIGURE 4" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[151,1437,195,1894]" captionTargetId="figure@27.[149,1438,193,1894]" captionTargetPageId="27" captionText="FIGURE 4. Genus Eugryllacris: Habitus dorsal view: A–B, E. trabicauda sp. nov. female holotype (A) and male paratype (B); C–D, E. vermicauda sp. nov. female holotype (C) male paratype (D); E, E. serricauda sp. nov. male holotype; F, E. guomashan sp. nov. female holotype; G, E. crassicauda sp. nov. female paratype; H, E. crassicauda cambodiana ssp. nov. female holotype; I, E. malaccensis (Griffini, 1908) male; J, E. serricauda sp. nov. female paratype. Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10109022" httpUri="https://zenodo.org/record/10109022/files/figure.png" pageId="36" pageNumber="37">Figs. 4</figureCitation>
|
||
A–B). Head: Face ovoid; fastigium verticis distinctly wider than scapus; ocelli distinct, of light color; fastigium frontis separated from fastigium verticis by a very fine suture; a pair of short, weak furrows, starting at internal angle of antennal scrobae, outline the fastigium frontis; subocular furrows weak and restricted to upper part (
|
||
<figureCitation id="43CB39C59773FFA5FDC5F8BFFD7AB864" box="[551,644,1842,1867]" captionStart="FIGURE 5" captionStartId="28.[151,250,1671,1693]" captionTargetBox="[152,1435,195,1649]" captionTargetId="figure@28.[152,1435,193,1649]" captionTargetPageId="28" captionText="FIGURE 5. Genus Eugryllacris: A, E. gandaki sp. nov. female holotype; B, E. serricauda sp. nov. male holotype; C, E. crassicauda sp. nov. female paratype; D, E. crassicauda cambodiana sp. nov. female holotype; E, E. sulcata sp. nov. female holotype; F, E. malaccensis (Griffini, 1908) male; G, E. vermicauda sp. nov. female holotype; H, E. trabicauda sp. nov. female holotype; I, E. inversa sp. nov. male holotype; J, E. guomashan sp. nov. female holotype; K, E. sulcata sp. nov. female holotype.—Face (A–J), habitus dorsal view (K). Scales 5 mm (face); 10 mm (habitus).—Abbreviations: as antennal scrobus, cf clypeo-frontal suture, cl clypeus, e compound eye, ff fastigium frontis with median ocellus, fv fastigium verticis, la labrum, lo lateral ocellus, lp labial palpus, ma mandible, mo median ocellus, mp maxillary palpus, sa sub-antennal furrow, so subocular furrow, sc scapus. The arrow in H points at the spine at anterior coxa." figureDoi="http://doi.org/10.5281/zenodo.10109024" httpUri="https://zenodo.org/record/10109024/files/figure.png" pageId="36" pageNumber="37">Fig. 5H</figureCitation>
|
||
). Abdominal tergites two and three with few, very minute, indistinct stridulatory pegs.
|
||
</paragraph>
|
||
<paragraph id="DB4F25409773FFA5FF25F8F7FB74B8F4" blockId="36.[151,1437,1194,2011]" pageId="36" pageNumber="37">
|
||
Wings surpassing hind knees. Tegmen (
|
||
<figureCitation id="43CB39C59773FFA5FD7EF8F7FD06B8BC" box="[668,760,1914,1939]" captionStart="FIGURE 1" captionStartId="12.[151,250,1761,1783]" captionTargetBox="[153,1439,198,1734]" captionTargetId="figure@12.[148,1440,193,1740]" captionTargetPageId="12" captionText="FIGURE 1. A, Larnaca (L.) eugenii (Griffini, 1914); B, Eugryllacris trabicauda sp. nov.; C, Ocellarnaca braueri (Griffini, 1911); D, Gryllacris kinabaluensis Griffini, 1915; E, Ultragryllacris triangula sp. nov.; F, Woznessenskia bavi sp. nov.; G, Psilogryllacris sp. (Karny's 1931 "allotype" of "dalbertisi Griffini 1909"); H, Zalarnaca (Zalarnaca) maninjau sp. nov.—Left tegmen with main veins (A–G), apical area of right hind tibia and hind tarsus in oblique lateral view with ventral pre-apical spine marked (H). Abbreviations for veins: C costa, ScA subcosta anterior, ScP subcosta posterior, R radius, RS radius sector, MA media anterior, MP media posterior, CuA cubitus anterior, CuP cubitus posterior, An1 first anal vein, following anal veins unmarked." figureDoi="http://doi.org/10.5281/zenodo.10109014" httpUri="https://zenodo.org/record/10109014/files/figure.png" pageId="36" pageNumber="37">Fig. 1B</figureCitation>
|
||
): Radius with two branches, both forked near tip; media anterior fused in basal area with radius; media posterior absent; cubitus anterior forks before mid-length into two veins, CuA1 and CuA2; cubitus posterior undivided, free throughout; with 4–5 anal veins.
|
||
</paragraph>
|
||
<paragraph id="DB4F25409772FFA4FF25FF15FD58BE6F" blockId="37.[151,1437,151,1112]" pageId="37" pageNumber="38">Legs: Fore coxa with a spine at fore margin; fore and mid femora unarmed; fore and mid tibiae with four pairs of large, on mid tibiae comparatively short, ventral spines and one pair of smaller ventral spurs; hind femur with 4– 9 external and 3–9 internal spines on ventral margins; hind tibia with spaced spines on both dorsal margins, ventral margins with one pre-apical spine each; with 3 apical spurs on both sides [male left hind tibia regenerated: shorter, with almost no spines, right hind tibia normal].</paragraph>
|
||
<paragraph id="DB4F25409772FFA4FF25FEC1FB45BEDA" blockId="37.[151,1437,151,1112]" pageId="37" pageNumber="38">Coloration. General color discolored brownish, might have been green when alive, legs with remnants of green; vertex unicolored; disc of pronotum unicolored, hind margin hardly darker. Legs of general color. Face unicolored brownish, maybe green when alive; eyes red brown; clypeo-frontal sutures darkened. Tegmen semitransparent with darker veins; except along anterior and apical margins with yellowish brown infumation (probably green when alive); hind wing semi-transparent white with brown veins and lighter cross-veins.</paragraph>
|
||
<paragraph id="DB4F25409772FFA4FF25FE72FA62BD87" blockId="37.[151,1437,151,1112]" pageId="37" pageNumber="38">
|
||
Male. Eighth abdominal tergite prolonged. Ninth abdominal tergite semi-globular: prolonged and down-curved laterally and apically; in descending posterior area furrowed in midline, at end of furrow split and on both sides of split area with an obtuse bump and below bump with a small conical projection pointing mediad (
|
||
<figureCitation id="43CB39C59772FFA4FAE5FDCAFAA9BD4F" box="[1287,1367,583,608]" captionStart="FIGURE 6" captionStartId="29.[151,250,1580,1602]" captionTargetBox="[162,1435,185,1560]" captionTargetId="figure@29.[158,1437,185,1560]" captionTargetPageId="29" captionText="FIGURE 6. Genus Eugryllacris: A–B, E. trabicauda sp. nov. paratype; C–D, E. vermicauda sp. nov. paratype; E–F, E. serricauda sp. nov. holotype; G–I, E. panteli (Bolivar, 1900) male type; J–K, E. malaccensis (Griffini, 1908); L–M, E. inversa sp. nov. holotype.—Male abdominal apex in about ventral (A, C, H, K, L), apical (E, J, M), and oblique lateral view (G, on both specimens studied distorted), tip of ninth abdominal tergite with projections in higher magnification (B, D, F), subgenital plate ventral view (H) and dorsal view (I: process of subgenital plate terminates into a down curved acute tip; tip of process vertical to disc of plate). Scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10109026" httpUri="https://zenodo.org/record/10109026/files/figure.png" pageId="37" pageNumber="38">Figs. 6</figureCitation>
|
||
A–B). Epiproct and paraproctes distinct. Subgenital plate almost twice as wide as long; lateral margins convex; apical margin wide, slightly obtuse-triangularly excised; curved styli inserted laterally just before apical margin (
|
||
<figureCitation id="43CB39C59772FFA4FAD4FD02FA71BD87" box="[1334,1423,655,680]" captionStart="FIGURE 6" captionStartId="29.[151,250,1580,1602]" captionTargetBox="[162,1435,185,1560]" captionTargetId="figure@29.[158,1437,185,1560]" captionTargetPageId="29" captionText="FIGURE 6. Genus Eugryllacris: A–B, E. trabicauda sp. nov. paratype; C–D, E. vermicauda sp. nov. paratype; E–F, E. serricauda sp. nov. holotype; G–I, E. panteli (Bolivar, 1900) male type; J–K, E. malaccensis (Griffini, 1908); L–M, E. inversa sp. nov. holotype.—Male abdominal apex in about ventral (A, C, H, K, L), apical (E, J, M), and oblique lateral view (G, on both specimens studied distorted), tip of ninth abdominal tergite with projections in higher magnification (B, D, F), subgenital plate ventral view (H) and dorsal view (I: process of subgenital plate terminates into a down curved acute tip; tip of process vertical to disc of plate). Scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10109026" httpUri="https://zenodo.org/record/10109026/files/figure.png" pageId="37" pageNumber="38">Fig. 6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DB4F25409772FFA4FF25FD39FDC9BC8B" blockId="37.[151,1437,151,1112]" pageId="37" pageNumber="38">
|
||
Female. Seventh abdominal sternite with roundly converging lateral margins and a long, narrow, parallel-sided apical projection with obtuse, in middle slightly concave tip covering base of subgenital plate (
|
||
<figureCitation id="43CB39C59772FFA4FAE7FD5AFAA8BDDF" box="[1285,1366,727,752]" captionStart="FIGURE 8" captionStartId="31.[151,250,1576,1598]" captionTargetBox="[151,1437,195,1554]" captionTargetId="figure@31.[148,1439,193,1555]" captionTargetPageId="31" captionText="FIGURE 8. Genus Eugryllacris: A–B, E. trabicauda sp. nov. holotype; C–D, E. vermicauda sp. nov. holotype; E–F, E. sulcata sp. nov. holotype; G–H, E. guomashan sp. nov. holotype.—Apical area of female abdomen with seventh abdominal sternite, subgenital plate and base of ovipositor in lateral (A, C, E, G) and ventral view (B, D, F, H)." figureDoi="http://doi.org/10.5281/zenodo.10109030" httpUri="https://zenodo.org/record/10109030/files/figure.png" pageId="37" pageNumber="38">Figs. 8</figureCitation>
|
||
A–B). Subgenital plate with nearly straight converging lateral margins, apical margin in middle concave, on both sides obtuse angular; surface in more than basal third grooved and in middle towards base membranous (
|
||
<figureCitation id="43CB39C59772FFA4FB1AFC92FAAEBC17" box="[1272,1360,799,824]" captionStart="FIGURE 8" captionStartId="31.[151,250,1576,1598]" captionTargetBox="[151,1437,195,1554]" captionTargetId="figure@31.[148,1439,193,1555]" captionTargetPageId="31" captionText="FIGURE 8. Genus Eugryllacris: A–B, E. trabicauda sp. nov. holotype; C–D, E. vermicauda sp. nov. holotype; E–F, E. sulcata sp. nov. holotype; G–H, E. guomashan sp. nov. holotype.—Apical area of female abdomen with seventh abdominal sternite, subgenital plate and base of ovipositor in lateral (A, C, E, G) and ventral view (B, D, F, H)." figureDoi="http://doi.org/10.5281/zenodo.10109030" httpUri="https://zenodo.org/record/10109030/files/figure.png" pageId="37" pageNumber="38">Fig. 8B</figureCitation>
|
||
). Base of ovipositor with a small sclerotised bump between subgenital plate and very base of ovipositor. Ovipositor elongate, only faintly curved dorsad, margins slightly and gradually narrowing towards tip; dorsal margin with a slight expansion before tip (
|
||
<figureCitation id="43CB39C59772FFA4FE32FC01FDD4BC8B" box="[464,554,908,933]" captionStart="FIGURE 9" captionStartId="33.[151,250,1827,1849]" captionTargetBox="[150,1436,197,1806]" captionTargetId="figure@33.[149,1438,195,1806]" captionTargetPageId="33" captionText="FIGURE 9. Genus Eugryllacris: A, E. inversa sp. nov. male holotype; B, E. maculipennis (Stål, 1877) det. Brunner female; C, E. maculipennis specularis (Karny, 1925) female type; D, E. gandaki sp. nov. female holotype; E, E. malaccensis (Griffini, 1908) male; F, E. crassicauda cambodiana ssp. nov. holotype; G, E. vermicauda sp. nov. holotype; H, E. inversa sp. nov. paratype; I, E. gandaki sp. nov.; J, E. sulcata sp. nov. holotype; K, E. crassicauda sp. nov. paratype; L, E. serricauda sp. nov. paratype; M, E. guomashan sp. nov. holotype; N, E. poultoniana (Griffini, 1909); O, E. trabicauda sp. nov. holotype.— Habitus in dorsal view (A–E) and lateral view (F), ovipositor (G–O). Scales 10 mm; images of ovipositor adapted to scale at bottom." figureDoi="http://doi.org/10.5281/zenodo.10109032" httpUri="https://zenodo.org/record/10109032/files/figure.png" pageId="37" pageNumber="38">Fig. 9O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DB4F25409772FFA4FF25FC3DFB12BB20" blockId="37.[151,1437,151,1112]" pageId="37" pageNumber="38">
|
||
Measurements (
|
||
<specimenCount id="CDF6EEC99772FFA4FE9EFC3DFE36BCE7" box="[380,456,943,968]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="CDF6EEC99772FFA4FE36FC3DFDBCBCE7" box="[468,578,943,968]" pageId="37" pageNumber="38" type="female">2 females</specimenCount>
|
||
).—body w/wings: male 54, female 49–54; body w/o wings: male 37, female 36–41; pronotum: male 8.5, female 9.0–9.5; tegmen: male 39, female 36–37; tegmen width: male 14, female 13; hind femur: male 22.5, female 21.5–23.0; antenna: female 115–190; ovipositor: female
|
||
<quantity id="1C0888A59772FFA4FB8EFC75FB16BB20" box="[1132,1256,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="37" pageNumber="38" unit="mm" value="39.0" valueMax="40.0" valueMin="38.0">38–40 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93EA76CB9772FFA4FF25FB91FE58BB77" pageId="37" pageNumber="38" type="etymology">
|
||
<paragraph id="DB4F25409772FFA4FF25FB91FE58BB77" blockId="37.[151,1437,151,1112]" pageId="37" pageNumber="38">
|
||
<emphasis id="E984F9529772FFA4FF25FB91FEB0BB1A" bold="true" box="[199,334,1052,1077]" pageId="37" pageNumber="38">Etymology.</emphasis>
|
||
The new species is named for the shape of the female seventh abdominal sternite; from Latin
|
||
<emphasis id="E984F9529772FFA4FA80FB91FA62BB1B" box="[1378,1436,1052,1076]" italics="true" pageId="37" pageNumber="38">trabs</emphasis>
|
||
(beam) and
|
||
<emphasis id="E984F9529772FFA4FEFEFBCDFE9EBB77" box="[284,352,1088,1112]" italics="true" pageId="37" pageNumber="38">cauda</emphasis>
|
||
(tail).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |