treatments-xml/data/4D/05/87/4D0587CAFA20142941DE673FFAB22404.xml
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<document id="7411AA4E8708FBAB48E7150620D54009" ID-CLB-Dataset="135281" ID-DOI="10.11646/zootaxa.5178.6.4" ID-GBIF-Dataset="fb77111d-d7eb-4170-b94c-c897ac032fae" ID-ISSN="1175-5326" ID-Zenodo-Dep="7037497" ID-ZooBank="F4393595-212F-47BE-8D2A-82F5042A6E8E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661935507431" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2022" docId="4D0587CAFA20142941DE673FFAB22404" docLanguage="en" docName="zootaxa.5178.6.4.pdf" docOrigin="Zootaxa 5178 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tmesiphorus shihongliangi Yin 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="581" masterDocId="B13CFFB2FA21142D41496421FB34205D" masterDocTitle="A new species of Tmesiphorus LeConte from northern Myanmar (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="582" masterPageNumber="577" pageNumber="578" updateTime="1699374489107" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E7E34BB1DBD2BA11F86CC220E9829F44">A new species of Tmesiphorus LeConte from northern Myanmar (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="30E1E468B4DD0A32D11765CE0152E337" type="personal">
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<mods:namePart id="C5442FEC5EE7EB0940BFC5A1D73D2D17">Yin, Zi-Wei</mods:namePart>
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<mods:title id="05C907730B3435B03CAC0DD83F32C381">Zootaxa</mods:title>
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<mods:part id="85CEF79EB47E7DE6CCFFD31A69D9FBE3">
<mods:date id="A33584B5351E1B9F5923D61FD236AC1A">2022</mods:date>
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<mods:number id="8BC3EC51124E72600A6ADE89F6F1F262">2022-08-31</mods:number>
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<treatment id="4D0587CAFA20142941DE673FFAB22404" ID-DOI="http://doi.org/10.5281/zenodo.7043992" ID-GBIF-Taxon="199920578" ID-Zenodo-Dep="7043992" LSID="urn:lsid:plazi:treatment:4D0587CAFA20142941DE673FFAB22404" httpUri="http://treatment.plazi.org/id/4D0587CAFA20142941DE673FFAB22404" lastPageId="4" lastPageNumber="581" pageId="1" pageNumber="578">
<subSubSection id="8DB66557FA20142C41DE673FF97D2364" box="[151,585,798,825]" pageId="1" pageNumber="578" type="nomenclature">
<paragraph id="C51336DCFA20142C41DE673FF97D2364" blockId="1.[151,585,798,861]" box="[151,585,798,825]" pageId="1" pageNumber="578">
<heading id="9E5B81B0FA20142C41DE673FF97D2364" bold="true" box="[151,585,798,825]" fontSize="11" level="1" pageId="1" pageNumber="578" reason="1">
<emphasis id="F7D8EACEFA20142C41DE673FF97D2364" bold="true" box="[151,585,798,825]" pageId="1" pageNumber="578">
<taxonomicName id="02AC4D5FFA20142C41DE673FFAD12364" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[151,485,798,825]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="578" phylum="Arthropoda" rank="species" species="shihongliangi" status="sp. nov.">
<emphasis id="F7D8EACEFA20142C41DE673FFAD12364" bold="true" box="[151,485,798,825]" italics="true" pageId="1" pageNumber="578">Tmesiphorus shihongliangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECEB57B5FA20142C40A5673EF97D2364" box="[492,585,799,825]" pageId="1" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="8DB66557FA20142C41DE6762FA382300" box="[151,268,835,861]" pageId="1" pageNumber="578" type="description">
<paragraph id="C51336DCFA20142C41DE6762FA382300" blockId="1.[151,585,798,861]" box="[151,268,835,861]" pageId="1" pageNumber="578">
(
<figureCitation id="5D972A59FA20142C41D66762FBD22300" box="[159,230,835,861]" captionStart="FIGURE 1" captionStartId="2.[151,250,1473,1497]" captionTargetBox="[154,1436,234,1447]" captionTargetId="figure-35@2.[151,1436,234,1447]" captionTargetPageId="2" captionText="FIGURE 1. Tmesiphorus shihongliangi sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Right maxillary palpus, ventral. D. Left antennal club, ventral. E Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="1" pageNumber="578">Figs 1</figureCitation>
,
<figureCitation id="5D972A59FA20142C41BD6762FA372300" box="[244,259,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="1" pageNumber="578">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8DB66557FA20142C41DE67AAF97423B0" pageId="1" pageNumber="578" type="materials_examined">
<paragraph id="C51336DCFA20142C41DE67AAF97423B0" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="578">
<emphasis id="F7D8EACEFA20142C41DE67AAFA7423F9" bold="true" box="[151,320,906,933]" pageId="1" pageNumber="578">Type material</emphasis>
(13 exx.).
<materialsCitation id="75C43C81FA20142C40F667AAF83223F8" ID-GBIF-Occurrence="3897814302" box="[447,774,906,933]" country="Myanmar" location="MYANMAR" pageId="1" pageNumber="578" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="F7D8EACEFA20142C40F667AAF9D123F8" bold="true" box="[447,741,906,933]" pageId="1" pageNumber="578">
<typeStatus id="1A17887EFA20142C40F667AAF90423F8" box="[447,560,907,933]" pageId="1" pageNumber="578" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="BDBB764CFA20142C437667ABF9E823F9" box="[575,732,906,932]" name="Myanmar" pageId="1" pageNumber="578">MYANMAR</collectingCountry>
:
</emphasis>
<specimenCount id="D3AAFD55FA20142C43A667AAF83223F8" box="[751,774,907,933]" pageId="1" pageNumber="578" type="male"></specimenCount>
</materialsCitation>
,
<materialsCitation id="75C43C81FA20142C425167AAF8B92394" ID-GBIF-Occurrence="3897814301" collectingDate="2016-12-13" collectionCode="SNUC" collectorName="Shi H. L." country="Myanmar" elevation="1051" latitude="27.5634" location="Putao Dist." longitude="97.1016" pageId="1" pageNumber="578" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<collectingCountry id="BDBB764CFA20142C425167AAF8B823F8" box="[792,908,907,933]" name="Myanmar" pageId="1" pageNumber="578">Myanmar</collectingCountry>
,
<collectingRegion id="0768F83EFA20142C42D167AAFF1923F8" box="[920,1069,907,933]" country="Myanmar" name="Kachin" pageId="1" pageNumber="578">Kachin State</collectingRegion>
,
<location id="C0736007FA20142C457367AAFF8F23F8" LSID="urn:lsid:plazi:treatment:4D0587CAFA20142941DE673FFAB22404:C0736007FA20142C457367AAFF8F23F8" box="[1082,1211,907,933]" country="Myanmar" latitude="27.5634" name="Putao Dist." pageId="1" pageNumber="578" stateProvince="Kachin">Putao Dist.</location>
,
<locationDeviation id="D4A12FFEFA20142C458167AAFEAC23F8" box="[1224,1432,907,933]" location="Ziradum" pageId="1" pageNumber="578">
<quantity id="02549B39FA20142C458167AAFE3723F8" box="[1224,1283,907,933]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="578" unit="km" value="2.0">2 km</quantity>
SE Ziradum
</locationDeviation>
, N27.5634, E97.1.64,
<date id="B112101CFA20142C40CF678EF93C2394" box="[390,520,943,969]" pageId="1" pageNumber="578" value="2016-12-13">
<collectingDate id="A156E9F4FA20142C40CF678EF93C2394" box="[390,520,943,969]" pageId="1" pageNumber="578" value="2016-12-13">2016.xii.13</collectingDate>
</date>
,
<quantity id="02549B39FA20142C435B678FF95D2394" box="[530,617,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.051" pageId="1" pageNumber="578" unit="m" value="1051.0">
<elevation id="4E81D1EFFA20142C435B678FF95D2394" box="[530,617,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.051" pageId="1" pageNumber="578" unit="m" value="1051.0">1051 m</elevation>
</quantity>
,
<collectorName id="6859530AFA20142C433D678EF9E02394" box="[628,724,943,969]" pageId="1" pageNumber="578">Shi H.L.</collectorName>
, in log (
<collectionCode id="A3BDAE19FA20142C427C678EF8B22394" box="[821,902,943,969]" pageId="1" pageNumber="578">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="75C43C81FA20142C42DE678EF90823B0" ID-GBIF-Occurrence="3897814303" collectingDate="2016-12-13" collectionCode="SNUC, IZCAS" collectorName="Shi H. L." country="Myanmar" elevation="1051" latitude="27.5634" location="Putao Dist." pageId="1" pageNumber="578" specimenCount="12" specimenCount-female="9" specimenCount-male="3" stateProvince="Kachin" typeStatus="paratype">
<emphasis id="F7D8EACEFA20142C42DE678EFFF22394" bold="true" box="[919,1222,942,969]" pageId="1" pageNumber="578">
<typeStatus id="1A17887EFA20142C42DE678EFF272394" box="[919,1043,943,969]" pageId="1" pageNumber="578" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="BDBB764CFA20142C4556678FFF882395" box="[1055,1212,942,968]" name="Myanmar" pageId="1" pageNumber="578">MYANMAR</collectingCountry>
:
</emphasis>
<specimenCount id="D3AAFD55FA20142C4585678EFE382394" box="[1228,1292,943,969]" pageId="1" pageNumber="578" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="D3AAFD55FA20142C445E678EFE632394" box="[1303,1367,943,969]" pageId="1" pageNumber="578" type="female">9 ♀♀</specimenCount>
, same data as for holotype (
<collectionCode id="A3BDAE19FA20142C40CD67F2FAE723B0" box="[388,467,979,1005]" pageId="1" pageNumber="578">SNUC</collectionCode>
,
<collectionCode id="A3BDAE19FA20142C409767F2F90023B0" box="[478,564,979,1005]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="578">IZCAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8DB66557FA20142C418E67D7F82624B4" pageId="1" pageNumber="578" type="diagnosis">
<paragraph id="C51336DCFA20142C418E67D7F82624B4" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="578">
<emphasis id="F7D8EACEFA20142C418E67D7FA76244D" bold="true" box="[199,322,1014,1040]" pageId="1" pageNumber="578">Diagnosis.</emphasis>
Body length
<quantity id="02549B39FA20142C409E67D6F955244C" box="[471,609,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="1" pageNumber="578" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.12.2 mm</quantity>
; dorsal vestiture coarsely punctate and with short setae. Head sub-hexagonal, with small vertexal and frontal foveae; ocular canthi well developed. Pronotum broadest at anterior 1/3. Elytra distinctly ridged, discal stria forming broad impression with smooth area at base. Tergites 1 and 2 (IV and VI) each with one middle and one pair of lateral longitudinal carinae extending through tergal length.
<emphasis id="F7D8EACEFA20142C45E16042FFD62420" box="[1192,1250,1123,1149]" italics="true" pageId="1" pageNumber="578">Male</emphasis>
: Antennomere 9 broadly and deeply impressed on ventral surface, 10 asymmetrical. Protrochanter spinose at base. Sternite 1 and 2 (IV and V) shallowly impressed at middle. Aedeagus elongate; median lobe divided into two parts at apex; endophallus elongate; parameres each with three apical setae.
</paragraph>
</subSubSection>
<subSubSection id="8DB66557FA201429418E60D3F9E32394" lastPageId="4" lastPageNumber="581" pageId="1" pageNumber="578" type="description">
<paragraph id="C51336DCFA20142C418E60D3F855256C" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="578">
<emphasis id="F7D8EACEFA20142C418E60D3FA6D2551" bold="true" box="[199,345,1266,1292]" pageId="1" pageNumber="578">Description.</emphasis>
<emphasis id="F7D8EACEFA20142C402860D2FAAF2550" box="[353,411,1267,1293]" italics="true" pageId="1" pageNumber="578">Male</emphasis>
. Body (
<figureCitation id="5D972A59FA20142C40B060D2F9652550" box="[505,593,1267,1293]" captionStart="FIGURE 1" captionStartId="2.[151,250,1473,1497]" captionTargetBox="[154,1436,234,1447]" captionTargetId="figure-35@2.[151,1436,234,1447]" captionTargetPageId="2" captionText="FIGURE 1. Tmesiphorus shihongliangi sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Right maxillary palpus, ventral. D. Left antennal club, ventral. E Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="1" pageNumber="578">Fig. 1A</figureCitation>
) length
<quantity id="02549B39FA20142C43FB60D2F86E2550" box="[690,858,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" metricValueMax="2.2" metricValueMin="2.14" pageId="1" pageNumber="578" unit="mm" value="2.17" valueMax="2.2" valueMin="2.14">2.142.20 mm</quantity>
; color reddish-brown, tarsi and mouthparts lighter. Dorsal vestiture coarsely punctate and covered with short setae.
</paragraph>
<paragraph id="C51336DCFA20142C418E611AFE0C26E0" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="578">
Head (
<figureCitation id="5D972A59FA20142C4058611AFA512508" box="[273,357,1339,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1473,1497]" captionTargetBox="[154,1436,234,1447]" captionTargetId="figure-35@2.[151,1436,234,1447]" captionTargetPageId="2" captionText="FIGURE 1. Tmesiphorus shihongliangi sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Right maxillary palpus, ventral. D. Left antennal club, ventral. E Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="1" pageNumber="578">Fig. 1B</figureCitation>
) sub-hexagonal, truncate at base, wider than long, length
<quantity id="02549B39FA20142C42AC611AFFBD2508" box="[997,1161,1339,1365]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="1" pageNumber="578" unit="mm" value="0.45" valueMax="0.46" valueMin="0.44">0.440.46 mm</quantity>
, width across eyes
<quantity id="02549B39FA20142C4414611AFA352524" metricMagnitude="-4" metricUnit="m" metricValue="4.95" metricValueMax="5.1" metricValueMin="4.8" pageId="1" pageNumber="578" unit="mm" value="0.495" valueMax="0.51" valueMin="0.48">0.48 0.51 mm</quantity>
; vertex convex, with small vertexal foveae (dorsal tentorial pits), with thin, incomplete median carina; antennal tubercles moderately raised; frons greatly impressed medially, confluent with clypeus at middle; clypeus sharply declined, smooth, anterior margin rounded and carinate; lacking ocular-mandibular carina. Venter with two small gular foveae (posterior tentorial pits) in shared transverse opening, lacking median carina. Maxillary palpus (
<figureCitation id="5D972A59FA20142C41D661CEFBC22654" box="[159,246,1519,1545]" captionStart="FIGURE 1" captionStartId="2.[151,250,1473,1497]" captionTargetBox="[154,1436,234,1447]" captionTargetId="figure-35@2.[151,1436,234,1447]" captionTargetPageId="2" captionText="FIGURE 1. Tmesiphorus shihongliangi sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Right maxillary palpus, ventral. D. Left antennal club, ventral. E Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="1" pageNumber="578">Fig. 1C</figureCitation>
) with palpomere 1 small, 2 pedunculate basally and enlarged apically, 3 lengthily subtriangular, 2 and 3 each with long, pencil-like, and apically setose protuberance on lateral margin, 4 greatly roundly and expanded laterally. Compound eyes greatly prominent, each composed of approximately 75 ommatidia; ocular canthus well developed. Antenna elongate, length
<quantity id="02549B39FA20142C437F627BF9EF2628" box="[566,731,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.135" metricValueMax="1.15" metricValueMin="1.12" pageId="1" pageNumber="578" unit="mm" value="1.135" valueMax="1.15" valueMin="1.12">1.121.15 mm</quantity>
, distinctly clubbed (
<figureCitation id="5D972A59FA20142C42F4627AFF272628" box="[957,1043,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1473,1497]" captionTargetBox="[154,1436,234,1447]" captionTargetId="figure-35@2.[151,1436,234,1447]" captionTargetPageId="2" captionText="FIGURE 1. Tmesiphorus shihongliangi sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Right maxillary palpus, ventral. D. Left antennal club, ventral. E Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="1" pageNumber="578">Fig. 1D</figureCitation>
) formed by greatly enlarged apical three antennomeres; antennomere 1 elongate, subcylindrical, 28 successively shorter, 8 shortest, 9 greatly enlarged, 10 much wider than 9, ventral surface broadly impressed, 11 smaller than 10, symmetrical at base.
</paragraph>
<paragraph id="C51336DCFA20142C418E62E6F889272C" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="578">
Pronotum (
<figureCitation id="5D972A59FA20142C400E62E6FAA926BC" box="[327,413,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1473,1497]" captionTargetBox="[154,1436,234,1447]" captionTargetId="figure-35@2.[151,1436,234,1447]" captionTargetPageId="2" captionText="FIGURE 1. Tmesiphorus shihongliangi sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Right maxillary palpus, ventral. D. Left antennal club, ventral. E Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="1" pageNumber="578">Fig. 1B</figureCitation>
) slightly wider than long, length
<quantity id="02549B39FA20142C425762E6F8F126BC" box="[798,965,1735,1761]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="1" pageNumber="578" unit="mm" value="0.45" valueMax="0.46" valueMin="0.44">0.440.46 mm</quantity>
, width
<quantity id="02549B39FA20142C455262E6FFF526BC" box="[1051,1217,1735,1761]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="4.9" metricValueMin="4.8" pageId="1" pageNumber="578" unit="mm" value="0.485" valueMax="0.49" valueMin="0.48">0.480.49 mm</quantity>
, widest at approximately apical 1/3; sides roundly expanded at widest point and evenly convergent posteriorly; disc moderately convex, lacking carina or sulcus; with small asetose median and setose lateral antebasal foveae; lacking basolateral foveae. Prosternum with anterior part slightly longer than coxal part at middle, with small, setose lateral procoxal foveae; lacking hypomeral ridge; margin of coxal cavity thinly carinate.
</paragraph>
<paragraph id="C51336DCFA20142F418E635AFA7A2088" blockId="1.[151,1437,906,2013]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="579" pageId="1" pageNumber="578">
Elytra much wider than long, length
<quantity id="02549B39FA20142C433C635BF82927C8" box="[629,797,1914,1941]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="1" pageNumber="578" unit="mm" value="0.51" valueMax="0.52" valueMin="0.5">0.500.52 mm</quantity>
, width
<quantity id="02549B39FA20142C423F635BFF2B27C8" box="[886,1055,1914,1941]" metricMagnitude="-4" metricUnit="m" metricValue="8.3" metricValueMax="8.5" metricValueMin="8.1" pageId="1" pageNumber="578" unit="mm" value="0.83" valueMax="0.85" valueMin="0.81">0.810.85 mm</quantity>
; each elytron (
<figureCitation id="5D972A59FA20142C4585635AFE1727C8" box="[1228,1315,1915,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1473,1497]" captionTargetBox="[154,1436,234,1447]" captionTargetId="figure-35@2.[151,1436,234,1447]" captionTargetPageId="2" captionText="FIGURE 1. Tmesiphorus shihongliangi sp. nov., male. A. Dorsal habitus. B. Head dorsum and pronotum. C. Right maxillary palpus, ventral. D. Left antennal club, ventral. E Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C." figureDoi="http://doi.org/10.5281/zenodo.7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="1" pageNumber="578">Fig. 1E</figureCitation>
) with two large, setose basal foveae; disc distinctly carinate, stria forming broad, deep impression with smooth basal region, extending posteriorly from outer basal fovea to approximately half of elytral length; humeral region also distinctly raised, lacking subhumeral fovea or marginal stria; posterolateral margin with broad notch. Metathoracic wings fully developed.
</paragraph>
<caption id="91D36654FA23142F41DE61E0FE5C25A0" ID-DOI="http://doi.org/10.5281/zenodo.7037501" ID-Zenodo-Dep="7037501" httpUri="https://zenodo.org/record/7037501/files/figure.png" pageId="2" pageNumber="579" startId="2.[151,250,1473,1497]" targetBox="[154,1436,234,1447]" targetPageId="2">
<paragraph id="C51336DCFA23142F41DE61E0FE5C25A0" blockId="2.[151,1436,1473,1533]" pageId="2" pageNumber="579">
<emphasis id="F7D8EACEFA23142F41DE61E0FA202584" bold="true" box="[151,276,1473,1497]" pageId="2" pageNumber="579">FIGURE 1.</emphasis>
<taxonomicName id="02AC4D5FFA23142F405361E0F91B2584" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[282,559,1473,1497]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="shihongliangi" status="sp. nov.">
<emphasis id="F7D8EACEFA23142F405361E0F91B2584" box="[282,559,1473,1497]" italics="true" pageId="2" pageNumber="579">Tmesiphorus shihongliangi</emphasis>
</taxonomicName>
<emphasis id="F7D8EACEFA23142F437F61E0F9BC2584" bold="true" box="[566,648,1473,1497]" pageId="2" pageNumber="579">
<taxonomicNameLabel id="ECEB57B5FA23142F437F61E0F9BC2584" box="[566,648,1473,1497]" pageId="2" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="F7D8EACEFA23142F439D61E0F9D82584" bold="true" box="[724,748,1473,1497]" pageId="2" pageNumber="579">A.</emphasis>
Dorsal habitus.
<emphasis id="F7D8EACEFA23142F42DD61E0F89F2584" bold="true" box="[916,939,1473,1497]" pageId="2" pageNumber="579">B.</emphasis>
Head dorsum and pronotum.
<emphasis id="F7D8EACEFA23142F459561E0FFC12584" bold="true" box="[1244,1269,1473,1497]" pageId="2" pageNumber="579">C.</emphasis>
Right maxillary palpus, ventral.
<emphasis id="F7D8EACEFA23142F407061C4FA6625A0" bold="true" box="[313,338,1509,1533]" pageId="2" pageNumber="579">D.</emphasis>
Left antennal club, ventral.
<emphasis id="F7D8EACEFA23142F433961C4F9B525A0" bold="true" box="[624,641,1509,1533]" pageId="2" pageNumber="579">E</emphasis>
Right elytron. Scale bars: 0.5 mm in A; 0.2 mm in B, D, E; 0.1 mm in C.
</paragraph>
</caption>
<paragraph id="C51336DCFA23142F418E620AFF7B26EC" blockId="2.[151,1437,1579,2038]" pageId="2" pageNumber="579">
Mesoventrite (
<figureCitation id="5D972A59FA23142F4024620AFAFC2618" box="[365,456,1579,1605]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="2" pageNumber="579">Fig. 2A</figureCitation>
) almost impunctate, with pair of admesal carinae, with setose mediobasal sulcus and large, setose lateral mesoventral foveae; inter-coxal process short. Metaventrite (
<figureCitation id="5D972A59FA23142F4557626EFF432634" box="[1054,1143,1615,1641]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="2" pageNumber="579">Fig. 2A</figureCitation>
) confluent with mesoventrite, coarsely punctate, with punctures much sparser laterally, broadly impressed posteromedially; with large, setose lateral mesocoxal foveae; posterior margin roundly and deeply emarginate at middle.
</paragraph>
<paragraph id="C51336DCFA23142F418E629DF8DD26A4" blockId="2.[151,1437,1579,2038]" pageId="2" pageNumber="579">
Legs elongate; protrochanter (
<figureCitation id="5D972A59FA23142F4350629AF95B2688" box="[537,623,1723,1749]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="2" pageNumber="579">Fig. 2B</figureCitation>
) with sharp, short spine at ventral margin, protibia distinctly curved and slightly expanded at middle; middle and hind legs lacking spine or tubercle.
</paragraph>
<paragraph id="C51336DCFA231429418E6322FF4E2140" blockId="2.[151,1437,1579,2038]" lastBlockId="4.[151,1437,150,1113]" lastPageId="4" lastPageNumber="581" pageId="2" pageNumber="579">
Abdomen enlarged, coarsely punctate dorso-ventrally, widest at lateral margins of tergite 1 (IV), length
<quantity id="02549B39FA23142F44146322FBCB271F" metricMagnitude="-4" metricUnit="m" metricValue="8.049999999999999" metricValueMax="8.2" metricValueMin="7.9" pageId="2" pageNumber="579" unit="mm" value="0.8049999999999999" valueMax="0.82" valueMin="0.79">0.79 0.82 mm</quantity>
, width
<quantity id="02549B39FA23142F401C6306FAC9271F" box="[341,509,1831,1858]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="8.8" metricValueMin="8.2" pageId="2" pageNumber="579" unit="mm" value="0.85" valueMax="0.88" valueMin="0.82">0.820.88 mm</quantity>
. Tergite 1 (IV) in dorsal view as long as 2 (V) (
<figureCitation id="5D972A59FA23142F456F6306FFB4271C" box="[1062,1152,1831,1857]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="2" pageNumber="579">Fig. 2C</figureCitation>
), with deep, setose basal sulcus, 2 with thin basal sulcus, tergites 1 and 2 each with one median and two lateral longitudinal carinae extending through tergal lengths, 14 (IVVII) each with one pair of basolateral foveae, 13 each with one pair of complete and 4 with triangular paratergites, 5 (VIII) with posterior margin broadly rounded, with one pair of small basolateral foveae, posterior margin roundly emarginate at middle. Sternite 2 (IV) (
<figureCitation id="5D972A59FA23142F428C6396FF29278C" box="[965,1053,1975,2001]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="2" pageNumber="579">Fig. 2D</figureCitation>
) longest, with broad, setose basal sulcus interrupted at middle and one pair of basolateral foveae at lateral ends of sulcus, 3 (V) (
<figureCitation id="5D972A59FA23142F459463FAFE0227A8" box="[1245,1334,2011,2037]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="2" pageNumber="579">Fig. 2D</figureCitation>
) slightly shorter than 2 at middle, 4 (VI) and 5 (VII) each short, combined shorter than 3 at middle, tergites 2 and 3 shallowly impressed at middle, 35 each with thin, asetose basal sulcus and one pair of basolateral foveae, 3 and 5 each with short setose fringe at basolateral margin, 6 (VIII) transverse, broadly emarginate at posterior margin, 7 (IX) (
<figureCitation id="5D972A59FA251429442664FEFB832140" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="4" pageNumber="581">Fig. 2E</figureCitation>
) composed of one moderately sclerotized median and two membranous lateral plates.
</paragraph>
<caption id="91D36654FA22142E41DE63B6F9DE27AA" ID-DOI="http://doi.org/10.5281/zenodo.7037503" ID-Zenodo-Dep="7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="3" pageNumber="580" startId="3.[151,250,1943,1967]" targetBox="[175,1412,182,1916]" targetPageId="3">
<paragraph id="C51336DCFA22142E41DE63B6F9DE27AA" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="580">
<emphasis id="F7D8EACEFA22142E41DE63B6FA2027F2" bold="true" box="[151,276,1943,1967]" pageId="3" pageNumber="580">FIGURE 2.</emphasis>
<taxonomicName id="02AC4D5FFA22142E405263B9F90427ED" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[283,560,1944,1968]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="shihongliangi" status="sp. nov.">
<emphasis id="F7D8EACEFA22142E405263B9F90427ED" box="[283,560,1944,1968]" italics="true" pageId="3" pageNumber="580">Tmesiphorus shihongliangi</emphasis>
</taxonomicName>
<emphasis id="F7D8EACEFA22142E437E63B9F9BC27ED" bold="true" box="[567,648,1944,1968]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="ECEB57B5FA22142E437E63B9F9BC27ED" box="[567,648,1944,1968]" pageId="3" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="F7D8EACEFA22142E439D63B6F9D927F2" bold="true" box="[724,749,1943,1967]" pageId="3" pageNumber="580">A.</emphasis>
Meso- and metaventrite.
<emphasis id="F7D8EACEFA22142E42BA63B9FF3E27ED" bold="true" box="[1011,1034,1944,1968]" pageId="3" pageNumber="580">B.</emphasis>
Protrochanter.
<emphasis id="F7D8EACEFA22142E45EE63B6FF8B27F2" bold="true" box="[1191,1215,1943,1967]" pageId="3" pageNumber="580">C.</emphasis>
Tergites 1 (IV) and 2 (V).
<emphasis id="F7D8EACEFA22142E418E639DFBD42789" bold="true" box="[199,224,1980,2004]" pageId="3" pageNumber="580">D.</emphasis>
Sternites 2 (IV) and 3 (V).
<emphasis id="F7D8EACEFA22142E40B2639DF9262789" bold="true" box="[507,530,1980,2004]" pageId="3" pageNumber="580">E.</emphasis>
Sternite 7 (IX).
<emphasis id="F7D8EACEFA22142E43F3639DF9DF2789" bold="true" box="[698,747,1980,2004]" pageId="3" pageNumber="580">FH</emphasis>
Aedeagus, dorsal (F), lateral (G), and ventral (H).
<emphasis id="F7D8EACEFA22142E45B9639DFE342789" bold="true" box="[1264,1280,1980,2004]" pageId="3" pageNumber="580">I.</emphasis>
Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I.
</paragraph>
</caption>
<paragraph id="C51336DCFA251429418E6506FA6721F0" blockId="4.[151,1437,150,1113]" pageId="4" pageNumber="581">
Aedeagus (
<figureCitation id="5D972A59FA251429400F6506FA8A211C" box="[326,446,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="4" pageNumber="581">Fig. 2FH</figureCitation>
)
<quantity id="02549B39FA25142940846506F907211C" box="[461,563,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="581" unit="mm" value="0.42">0.42 mm</quantity>
long, asymmetrical; median lobe with large basal capsule and oval foramen, ventral stalk broad dorso-ventrally, curved ventrally in lateral view, apical part greatly concave at middle and divided into two parts; endophallus composed of single elongate sclerite; parameres short (
<figureCitation id="5D972A59FA25142945C0654EFFEC21D4" box="[1161,1240,367,393]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[175,1412,182,1916]" captionTargetId="figure-15@3.[174,1413,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Tmesiphorus shihongliangi sp. nov., male. A. Meso- and metaventrite. B. Protrochanter. C. Tergites 1 (IV) and 2 (V). D. Sternites 2 (IV) and 3 (V). E. Sternite 7 (IX). FH Aedeagus, dorsal (F), lateral (G), and ventral (H). I. Left paramere. Scale bars: 0.2 mm in A, CH; 0.1 mm in B; 0.05 mm in I." figureDoi="http://doi.org/10.5281/zenodo.7037503" httpUri="https://zenodo.org/record/7037503/files/figure.png" pageId="4" pageNumber="581">Fig. 2I</figureCitation>
), each with three thin apical setae.
</paragraph>
<paragraph id="C51336DCFA251429418E6596F93F22D8" blockId="4.[151,1437,150,1113]" pageId="4" pageNumber="581">
<emphasis id="F7D8EACEFA251429418E6596FA2F218C" box="[199,283,439,465]" italics="true" pageId="4" pageNumber="581">Female</emphasis>
. Similar to male in external morphology; antenna slightly shorter, club less enlarged and lacking modifications; each compound eye composed of approximately 60 ommatidia; elytra relatively shorter and more constricted at bases; metathoracic wings also well-developed; protrochanter lacking ventral spine; sternite 2 (IV) and 3 (V) flat at middle. Measurements (as for male): body length
<quantity id="02549B39FA251429420B6603F8DD2260" box="[834,1001,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.16" metricValueMax="2.17" metricValueMin="2.15" pageId="4" pageNumber="581" unit="mm" value="2.16" valueMax="2.17" valueMin="2.15">2.152.17 mm</quantity>
; length/width of head 0.440.46/
<quantity id="02549B39FA25142944176602FA35223C" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.699999999999999" pageId="4" pageNumber="581" unit="mm" value="0.475" valueMax="0.48" valueMin="0.47">0.47 0.48 mm</quantity>
, pronotum 0.430.45/
<quantity id="02549B39FA25142940B66667F993223C" box="[511,679,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.699999999999999" pageId="4" pageNumber="581" unit="mm" value="0.475" valueMax="0.48" valueMin="0.47">0.470.48 mm</quantity>
, elytra 0.49/
<quantity id="02549B39FA25142942736666F8D6223C" box="[826,994,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.1" metricValueMin="7.9" pageId="4" pageNumber="581" unit="mm" value="0.8" valueMax="0.81" valueMin="0.79">0.790.81 mm</quantity>
; abdomen 0.800.84/0.0.86
<quantity id="02549B39FA25142944676666FEAC223C" box="[1326,1432,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="4" pageNumber="581" unit="mm" value="0.87">0.87 mm</quantity>
; length of antenna
<quantity id="02549B39FA2514294028664AF93322D8" box="[353,519,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.065" metricValueMax="1.07" metricValueMin="1.06" pageId="4" pageNumber="581" unit="mm" value="1.065" valueMax="1.07" valueMin="1.06">1.061.07 mm</quantity>
.
</paragraph>
<paragraph id="C51336DCFA251429418E66AFF9E32394" blockId="4.[151,1437,150,1113]" pageId="4" pageNumber="581">
<emphasis id="F7D8EACEFA251429418E66AFFA9B22F4" bold="true" box="[199,431,654,681]" pageId="4" pageNumber="581">Comparative notes.</emphasis>
Two
<taxonomicName id="02AC4D5FFA25142940A566AEF9E022F4" box="[492,724,655,681]" pageId="4" pageNumber="581">
<emphasis id="F7D8EACEFA25142940A566AEF94922F4" box="[492,637,655,681]" italics="true" pageId="4" pageNumber="581">Tmesiphorus</emphasis>
species
</taxonomicName>
described more than a century ago are known to occur in
<collectingCountry id="BDBB764CFA251429441866AEFBF32290" name="Myanmar" pageId="4" pageNumber="581">Myanmar</collectingCountry>
:
<taxonomicName id="02AC4D5FFA251429419B6692F9342290" authority="(Motschulsky)" baseAuthorityName="Motschulsky" baseAuthorityYear="1851" box="[210,512,691,717]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="paulinae">
<emphasis id="F7D8EACEFA251429419B6692FBD62291" box="[210,226,691,716]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA25142941B96692FA662290" box="[240,338,691,717]" italics="true" pageId="4" pageNumber="581">paulinae</emphasis>
(Motschulsky)
</taxonomicName>
, and
<taxonomicName id="02AC4D5FFA25142943726692F81C2290" authority="Schaufuss" authorityName="Schaufuss" box="[571,808,691,717]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="F7D8EACEFA25142943726692F97F2291" box="[571,587,691,716]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA25142943106692F99B2290" box="[601,687,691,717]" italics="true" pageId="4" pageNumber="581">raffrayi</emphasis>
Schaufuss
</taxonomicName>
(
<bibRefCitation id="A13D4B2DFA251429427F6692FF3F2290" author="Motschulsky, V." box="[822,1035,690,717]" pageId="4" pageNumber="581" pagination="479 - 511" refId="ref3055" refString="Motschulsky, V. (1851) Enumeration des nouvelles especes de Coleopteres. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 24 (4), 479 - 511." type="journal article" year="1851">Motschulsky 1851</bibRefCitation>
;
<bibRefCitation id="A13D4B2DFA251429455E6692FF9B2290" author="Raffray, A." box="[1047,1199,691,717]" pageId="4" pageNumber="581" pagination="229 - 251" refId="ref3280" refString="Raffray, A. (1883) Pselaphides nouveaux ou peu connus. Deuxieme memoire. Revue d'Entomologie 2, 229 - 251, pls. 4 - 5." type="journal article" year="1883">Raffray 1883</bibRefCitation>
; Schaufuss, C. 1888;
<bibRefCitation id="A13D4B2DFA25142941DE66F6FA1B22AC" author="Blattny, C." box="[151,303,726,753]" pageId="4" pageNumber="581" pagination="179 - 222" refId="ref2524" refString="Blattny, C. (1925) Revise Pselaphidu sbirky Helferovy Nar. Musea v Praze. Revision der Pselaphiden der Collection Helfer im National-Museum in Prag. Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze, 3 (26), 179 - 222." type="journal article" year="1925">Blattný 1925</bibRefCitation>
).
<taxonomicName id="02AC4D5FFA251429400D66F6F94C22AC" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[324,632,727,753]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="shihongliangi" status="sp. nov.">
<emphasis id="F7D8EACEFA251429400D66F6F94C22AC" box="[324,632,727,753]" italics="true" pageId="4" pageNumber="581">Tmesiphorus shihongliangi</emphasis>
</taxonomicName>
<emphasis id="F7D8EACEFA25142943C866F6F9E922AC" bold="true" box="[641,733,727,753]" pageId="4" pageNumber="581">
<taxonomicNameLabel id="ECEB57B5FA25142943C866F6F9E922AC" box="[641,733,727,753]" pageId="4" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be readily separated from them by the smaller body size (
<quantity id="02549B39FA25142941D666DAFA1D2348" box="[159,297,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="4" pageNumber="581" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.12.2 mm</quantity>
vs.
<quantity id="02549B39FA251429401066DAFA852348" box="[345,433,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="581" unit="mm" value="2.6">2.6 mm</quantity>
for
<taxonomicName id="02AC4D5FFA25142940AB66DAF9572348" baseAuthorityName="Motschulsky" baseAuthorityYear="1851" box="[482,611,763,789]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="paulinae">
<emphasis id="F7D8EACEFA25142940AB66DAFAC62349" box="[482,498,763,788]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA251429434866DAF9572348" box="[513,611,763,789]" italics="true" pageId="4" pageNumber="581">paulinae</emphasis>
</taxonomicName>
and
<quantity id="02549B39FA25142943D566DAF8362348" box="[668,770,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.66" pageId="4" pageNumber="581" unit="mm" value="2.66">2.66 mm</quantity>
for
<taxonomicName id="02AC4D5FFA251429427A66DAF89C2348" box="[819,936,763,789]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="F7D8EACEFA251429427A66DAF8772349" box="[819,835,763,788]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA251429421B66DAF89C2348" box="[850,936,763,789]" italics="true" pageId="4" pageNumber="581">raffrayi</emphasis>
</taxonomicName>
), and presence of three longitudinal carinae on tergites 1 (IV) and 2 (V). Both
<taxonomicName id="02AC4D5FFA2514294357673EF9AB2364" baseAuthorityName="Motschulsky" baseAuthorityYear="1851" box="[542,671,799,825]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="paulinae">
<emphasis id="F7D8EACEFA2514294357673EF91A2365" box="[542,558,799,824]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA2514294374673EF9AB2364" box="[573,671,799,825]" italics="true" pageId="4" pageNumber="581">paulinae</emphasis>
</taxonomicName>
and
<taxonomicName id="02AC4D5FFA2514294391673EF8782364" box="[728,844,799,825]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="F7D8EACEFA2514294391673EF9DC2365" box="[728,744,799,824]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA25142943BF673EF8782364" box="[758,844,799,825]" italics="true" pageId="4" pageNumber="581">raffrayi</emphasis>
</taxonomicName>
possess only one pair of lateral carinae on tergite 1, and lack any carina on tergite 2. The new species is morphologically similar to
<taxonomicName id="02AC4D5FFA25142945426762FFCE2300" authority="Jeannel" authorityName="Jeannel" authorityYear="1960" box="[1035,1274,835,861]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="F7D8EACEFA25142945426762FF2F2301" box="[1035,1051,835,860]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA25142945606762FFAB2300" box="[1065,1183,835,861]" italics="true" pageId="4" pageNumber="581">championi</emphasis>
Jeannel
</taxonomicName>
from northern
<collectingCountry id="BDBB764CFA25142941DE6746FBE523DC" box="[151,209,871,897]" name="India" pageId="4" pageNumber="581">India</collectingCountry>
by sharing the pattern of abdominal carinae.
<taxonomicName id="02AC4D5FFA25142943816746F8E223DC" authorityName="Jeannel" authorityYear="1960" box="[712,982,871,897]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="F7D8EACEFA25142943816746F8E223DC" box="[712,982,871,897]" italics="true" pageId="4" pageNumber="581">Tmesiphorus championi</emphasis>
</taxonomicName>
has a slightly larger body size (
<quantity id="02549B39FA251429447C6746FEA423DC" box="[1333,1424,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="581" unit="mm" value="2.4">2.4 mm</quantity>
), and
<bibRefCitation id="A13D4B2DFA251429418F67AAFA4923F8" author="Jeannel, R." box="[198,381,907,933]" pageId="4" pageNumber="581" pagination="403 - 456" refId="ref2785" refString="Jeannel, R. (1960) Sur les Pselaphides (Coleoptera) de l'Inde septentrionale. Bulletin of the British Museum (Natural History), Entomology, 9, 403 - 456. https: // doi. org / 10.5962 / bhl. part. 27559" type="journal article" year="1960">Jeannels (1960)</bibRefCitation>
original figures of the male antennal clubs and the aedeagus provided enough details to clearly separate this species from
<taxonomicName id="02AC4D5FFA25142940F6678EF9422394" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[447,630,943,969]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="species" species="shihongliangi" status="sp. nov.">
<emphasis id="F7D8EACEFA25142940F6678EFAFB2395" box="[447,463,943,968]" italics="true" pageId="4" pageNumber="581">T</emphasis>
.
<emphasis id="F7D8EACEFA2514294095678EF9422394" box="[476,630,943,969]" italics="true" pageId="4" pageNumber="581">shihongliangi</emphasis>
</taxonomicName>
<emphasis id="F7D8EACEFA2514294334678EF9E32394" bold="true" box="[637,727,943,969]" pageId="4" pageNumber="581">
<taxonomicNameLabel id="ECEB57B5FA2514294334678EF9E32394" box="[637,727,943,969]" pageId="4" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="8DB66557FA251429418E67F3FE70244C" pageId="4" pageNumber="581" type="distribution">
<paragraph id="C51336DCFA251429418E67F3F9F723B0" blockId="4.[151,1437,150,1113]" box="[199,707,978,1005]" pageId="4" pageNumber="581">
<emphasis id="F7D8EACEFA251429418E67F3FA5523B1" bold="true" box="[199,353,978,1004]" pageId="4" pageNumber="581">Distribution.</emphasis>
North
<collectingCountry id="BDBB764CFA25142940F967F2F91523B0" box="[432,545,979,1005]" name="Myanmar" pageId="4" pageNumber="581">Myanmar</collectingCountry>
:
<collectingRegion id="0768F83EFA251429436267F2F98B23B0" box="[555,703,979,1005]" country="Myanmar" name="Kachin" pageId="4" pageNumber="581">Kachin State</collectingRegion>
.
</paragraph>
<paragraph id="C51336DCFA251429418E67D7FE70244C" blockId="4.[151,1437,150,1113]" box="[199,1348,1014,1041]" pageId="4" pageNumber="581">
<emphasis id="F7D8EACEFA251429418E67D7FA7F244D" bold="true" box="[199,331,1014,1040]" pageId="4" pageNumber="581">Bionomics.</emphasis>
All individuals were collected under bark of large fallen logs in a broad-leaved rain forest.
</paragraph>
</subSubSection>
<subSubSection id="8DB66557FA251429418E603AFAB22404" pageId="4" pageNumber="581" type="etymology">
<paragraph id="C51336DCFA251429418E603AFAB22404" blockId="4.[151,1437,150,1113]" pageId="4" pageNumber="581">
<emphasis id="F7D8EACEFA251429418E603AFA7A2468" bold="true" box="[199,334,1051,1077]" pageId="4" pageNumber="581">Etymology.</emphasis>
The new species is named after Hong-Liang Shi, who collected all the material and kindly offered them to me for study.
</paragraph>
</subSubSection>
</treatment>
</document>