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<document id="10D90DE681E4A945C03B63EE90A17E05" ID-DOI="10.1655/06-001.1" ID-Zenodo-Dep="14384794" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1733861782093" checkinUser="juliana" docAuthor="Torres-Carvajal, Omar" docDate="2007" docId="467D8791FFFAFFD6FF62FDF9154AFA1B" docLanguage="en" docName="HerpetolMonogr.21.1.76-178.pdf.pdf" docOrigin="Herpetological Monographs 21 (1)" docSource="https://doi.org/10.1655/06-001.1" docTitle="Stenocercus orientalis Fritts 1972" docType="treatment" docVersion="6" lastPageNumber="140" masterDocId="BA44FFE9FFBBFF97FFBCFFC41632FFF9" masterDocTitle="A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS" masterLastPageNumber="178" masterPageNumber="76" pageNumber="140" updateTime="1733923315685" updateUser="juliana">
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<mods:title id="EE9220B250103F6DC25F97FF7FB9815B">A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS</mods:title>
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<mods:namePart id="C0CF3AB1694F80F999FCE39F508315F6">Torres-Carvajal, Omar</mods:namePart>
<mods:affiliation id="90444C6B9932AADB497736BFCB0D5C5B">Department of Vertebrate Zoologŋ, National Museum of Natural Historŋ, Smithsonian Institution, Washington, DC 20560, USA &amp; Escuela de Biología, Pontificia UniƲersidad Católica del Ecuador, AƲenida 12 de Octubre ŋ Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:date id="C5B51A6F9236864CD7885CB5CC610312">2007</mods:date>
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<treatment id="467D8791FFFAFFD6FF62FDF9154AFA1B" ID-DOI="http://doi.org/10.5281/zenodo.14372749" ID-Zenodo-Dep="14372749" LSID="urn:lsid:plazi:treatment:467D8791FFFAFFD6FF62FDF9154AFA1B" httpUri="http://treatment.plazi.org/id/467D8791FFFAFFD6FF62FDF9154AFA1B" lastPageNumber="140" pageId="65" pageNumber="140">
<subSubSection id="86CE650CFFFAFFD6FF62FDF91427FDAC" box="[222,533,573,597]" pageId="65" pageNumber="140" type="nomenclature">
<paragraph id="CE6B3687FFFAFFD6FF62FDF91427FDAC" blockId="65.[122,633,573,1722]" box="[222,533,573,597]" pageId="65" pageNumber="140">
<heading id="952381EBFFFAFFD6FF62FDF91427FDAC" box="[222,533,573,597]" centered="true" fontSize="10" level="3" pageId="65" pageNumber="140" reason="4">
<taxonomicName id="09D44D04FFFAFFD6FF62FDF91427FDAC" ID-CoL="4ZWKH" authority="Fritts" authorityName="Fritts" authorityYear="1972" box="[222,533,573,597]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="FCA0EA95FFFAFFD6FF62FDF917FDFDAC" box="[222,463,573,597]" italics="true" pageId="65" pageNumber="140">Stenocercus orientalis</emphasis>
Fritts
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFAFFD6FEF5FD9F1798FD8A" box="[329,426,603,627]" pageId="65" pageNumber="140" type="description">
<paragraph id="CE6B3687FFFAFFD6FEF5FD9F1798FD8A" blockId="65.[122,633,573,1722]" box="[329,426,603,627]" pageId="65" pageNumber="140">
<heading id="952381EBFFFAFFD6FEF5FD9F1798FD8A" box="[329,426,603,627]" centered="true" fontSize="10" level="3" pageId="65" pageNumber="140" reason="4">
(
<figureCitation id="56EF2A02FFFAFFD6FEEDFD9F1790FD8A" box="[337,418,603,627]" captionStart="FIG" captionStartId="59.[143,157,1718,1737]" captionTargetBox="[122,1174,189,1701]" captionTargetId="figure-1@59.[122,1174,189,1701]" captionTargetPageId="59" captionText="FIG. 20.—Eight species of Stenocercus. (A) S. marmoratus (AMU); (B) S. melanopŋgus (PVE); (C) S. modestus (MLU); (D) S. nigromaculatus (WED); (E) S. nubicola (PVE); (F) S. ochoai (WED); (G) S. orientalis (PVE); (H) S. ornatissimus (MLU)." figureDoi="http://doi.org/10.5281/zenodo.14384850" httpUri="https://zenodo.org/record/14384850/files/figure.png" pageId="65" pageNumber="140">Fig. 20</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFAFFD6FFC6FD4617BDFD08" pageId="65" pageNumber="140" type="reference_group">
<paragraph id="CE6B3687FFFAFFD6FFC6FD4617BDFD08" blockId="65.[122,633,573,1722]" pageId="65" pageNumber="140">
<taxonomicName id="09D44D04FFFAFFD6FFC6FD461415FD60" authority="Fritts, 1972: 14" authorityName="Fritts" authorityPageNumber="14" authorityYear="1972" box="[122,551,641,665]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="FCA0EA95FFFAFFD6FFC6FD461742FD60" box="[122,368,641,665]" italics="true" pageId="65" pageNumber="140">Stenocercus orientalis</emphasis>
<bibRefCitation id="AA454B76FFFAFFD6FEC1FD451415FD60" author="FRITTS, T. H." box="[381,551,641,665]" journalOrPublisher="Ocassional Papers University of Kansas Museum of Natural History" pageId="65" pageNumber="140" pagination="1 - 21" part="10" refId="ref77696" refString="FRITTS, T. H. 1972. New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae). Ocassional Papers University of Kansas Museum of Natural History 10: 1 - 21." title="New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae)" type="journal article" year="1972">Fritts, 1972:14</bibRefCitation>
</taxonomicName>
.
<materialsCitation id="7EBC3CDAFFFAFFD6FD84FD4517BDFD08" collectionCode="KU" country="Peru" elevation="2340" location="Chachapoyas" municipality="Chachapoyas" pageId="65" pageNumber="140" specimenCode="KU 134466" specimenCount="1" specimenCount-male="1" stateProvince="Departamento Amazonas" typeStatus="holotype">
<typeStatus id="116F8825FFFAFFD6FD84FD4516FAFD4E" pageId="65" pageNumber="140">Holotype</typeStatus>
:
<specimenCode id="9E729EFCFFFAFFD6FF65FD5B176DFD4F" box="[217,351,670,694]" pageId="65" pageNumber="140">KU 134466</specimenCode>
, a
<specimenCount id="D8D2FD0EFFFAFFD6FE37FD5A17F2FD4F" box="[395,448,670,694]" pageId="65" pageNumber="140" type="male">male</specimenCount>
from
<collectingMunicipality id="2E0FACFDFFFAFFD6FDA6FD5A16E7FD2D" pageId="65" pageNumber="140">Chachapoyas</collectingMunicipality>
,
<quantity id="092C9B62FFFAFFD6FF54FD78170CFD2A" box="[232,318,700,724]" metricMagnitude="3" metricUnit="m" metricValue="2.34" pageId="65" pageNumber="140" unit="m" value="2340.0">
<elevation id="45F9D1B4FFFAFFD6FF54FD78170CFD2A" box="[232,318,700,724]" metricMagnitude="3" metricUnit="m" metricValue="2.34" pageId="65" pageNumber="140" unit="m" value="2340.0">2340 m</elevation>
</quantity>
,
<collectingRegion id="0C10F865FFFAFFD6FEEDFD781446FD2D" box="[337,628,700,724]" pageId="65" pageNumber="140">Departamento Amazonas</collectingRegion>
,
<collectingCountry id="B6C37617FFFAFFD6FF29FD1D16F8FD08" box="[149,202,729,753]" name="Peru" pageId="65" pageNumber="140">Perú</collectingCountry>
̕̕;
<bibRefCitation id="AA454B76FFFAFFD6FF59FD1D17B9FD08" author="FRITTS, T. H." box="[229,395,729,753]" journalOrPublisher="San Diego Society of Natural History Memoir" pageId="65" pageNumber="140" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">Fritts, 1974:60</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFAFFD6FF29FD3C1416FBEE" pageId="65" pageNumber="140" type="diagnosis">
<paragraph id="CE6B3687FFFAFFD6FF29FD3C1416FBEE" blockId="65.[122,633,573,1722]" pageId="65" pageNumber="140">
<emphasis id="FCA0EA95FFFAFFD6FF29FD3C1733FCF6" box="[149,257,760,783]" italics="true" pageId="65" pageNumber="140">Diagnosis</emphasis>
.—
<taxonomicName id="09D44D04FFFAFFD6FE9EFD3C1423FCF6" authorityName="Fritts" authorityYear="1972" box="[290,529,759,783]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="FCA0EA95FFFAFFD6FE9EFD3C1423FCF6" box="[290,529,759,783]" italics="true" pageId="65" pageNumber="140">Stenocercus orientalis</emphasis>
</taxonomicName>
is distinguished from other species of
<taxonomicName id="09D44D04FFFAFFD6FE4BFCD1144AFCD5" box="[503,632,789,812]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="genus">
<emphasis id="FCA0EA95FFFAFFD6FE4BFCD1144AFCD5" box="[503,632,789,812]" italics="true" pageId="65" pageNumber="140">Stenocercus</emphasis>
</taxonomicName>
except
<taxonomicName id="09D44D04FFFAFFD6FF7AFCF6175AFCB3" authorityName="Boulenger" authorityYear="1900" box="[198,360,818,842]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="chrnsopngus">
<emphasis id="FCA0EA95FFFAFFD6FF7AFCF6175AFCB3" box="[198,360,818,842]" italics="true" pageId="65" pageNumber="140">S. chrysopygus</emphasis>
</taxonomicName>
,
<taxonomicName id="09D44D04FFFAFFD6FEC8FCF617D6FCB3" authorityName="Boulenger" authorityYear="1885" box="[372,484,818,842]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="FCA0EA95FFFAFFD6FEC8FCF617D6FCB3" box="[372,484,818,842]" italics="true" pageId="65" pageNumber="140">S. cupreus</emphasis>
</taxonomicName>
,
<emphasis id="FCA0EA95FFFAFFD6FE4CFCF61730FC9E" italics="true" pageId="65" pageNumber="140">
<taxonomicName id="09D44D04FFFAFFD6FE4CFCF6144AFCB3" authorityName="Cadle" authorityYear="1998" box="[496,632,818,842]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="latebrosus">S. latebrosus</taxonomicName>
<taxonomicName id="09D44D04FFFAFFD6FFC6FC941730FC9E" box="[122,258,847,871]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="modestus">S. modestus</taxonomicName>
</emphasis>
, and
<taxonomicName id="09D44D04FFFAFFD6FEF2FC9417C8FC9E" box="[334,506,848,871]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="ornatissimus">
<emphasis id="FCA0EA95FFFAFFD6FEF2FC9417C8FC9E" box="[334,506,848,871]" italics="true" pageId="65" pageNumber="140">S. ornatissimus</emphasis>
</taxonomicName>
by having granular scales on the posterior surface of thighs, conspicuous antehumeral and oblique neck folds, a distinct mite pocket under oblique neck fold, and by lacking a vertebral crest. Of these species,
<taxonomicName id="09D44D04FFFAFFD6FEA5FC2617AEFC03" authorityName="Fritts" authorityYear="1972" box="[281,412,994,1018]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="FCA0EA95FFFAFFD6FEA5FC2617AEFC03" box="[281,412,994,1018]" italics="true" pageId="65" pageNumber="140">S. orientalis</emphasis>
</taxonomicName>
is unique in having prominently keeled dorsal head scales.
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFAFFD6FF29FBDA1500FB42" pageId="65" pageNumber="140" type="description">
<paragraph id="CE6B3687FFFAFFD6FF29FBDA15DEFC6C" blockId="65.[122,633,573,1722]" lastBlockId="65.[664,1175,190,1506]" pageId="65" pageNumber="140">
<emphasis id="FCA0EA95FFFAFFD6FF29FBDA1724FBCC" box="[149,278,1054,1077]" italics="true" pageId="65" pageNumber="140">Description</emphasis>
.—(1) Maximum SVL in males 79 mm (
<emphasis id="FCA0EA95FFFAFFD6FF64FBFF16D4FBAB" box="[216,230,1083,1106]" italics="true" pageId="65" pageNumber="140">n</emphasis>
= 17); (2) maximum SVL in females 66 mm (
<emphasis id="FCA0EA95FFFAFFD6FF5CFB9D16DCFB89" box="[224,238,1113,1136]" italics="true" pageId="65" pageNumber="140">n</emphasis>
= 17); (3) vertebrals 4458; (4) paravertebrals 4862; (5) scales around midbody 4659; (6) supraoculars 56; (7) internasals 35; (8) postrostrals 47; (9) loreals 23; (10) gulars 2026; (11) subdigitals on Finger IV 1621; (12) subdigitals on Toe IV 2130; (13) posthumeral mite pocket present as one or more vertical folds or ridges; (14) postfemoral mite pocket distinct with slit-like opening; (15) parietal eye always visible through interparietal cornea; (16) scales on occipitoparietal region small, keeled, imbricate; (17) projecting angulate temporals absent; (18) row of enlarged supraoculars occupying most of supraocular region absent; (19) scales on frontonasal region weakly imbricate anteriorly; (20) no preauricular fringe present; (21) antehumeral and oblique neck folds present; (22) lateral nuchals less than half the size of dorsal nuchals; (23) posterior gulars cycloid, smooth, slightly imbricate, notched; (24) lateral and dorsal body scales similar in size; (25) vertebrals and adjacent paravertebrals similar in size; (26) dorsolateral crest absent; (27) ventrals smooth, imbricate; (28) scales on posterior surfaces of thighs granular; (29) inguinal granular pocket absent; (30) inguinal groove absent; (31) preanals not projected; (32) tail not strongly compressed laterally in adult males; (33) tail length 6569% of total length; (34) caudal whorls per autotomic segment three; (35) caudals not spinose; (36) dark brown stripe extending anterodorsally from subocular region to supraciliaries absent; (37) dark patch extensively covering gular region of females absent; (38) dark patch extensively covering gular region of adult males absent; (39) black patch on ventral surface of neck in adult males absent; (40) dark midventral longitudinal mark such as faint line, conspicuous stripe, or extensive patch in adult males absent; (41) dark patches on ventral surface of thighs in adult males absent; (42) postxiphisternal inscriptional ribs not in contact midventrally, Patterns 1B and 2C.
</paragraph>
<paragraph id="CE6B3687FFFAFFD6FD0FFC5F1500FB42" blockId="65.[664,1175,190,1506]" pageId="65" pageNumber="140">
<emphasis id="FCA0EA95FFFAFFD6FD0FFC5F1577FC4A" box="[691,837,923,947]" italics="true" pageId="65" pageNumber="140">Color in life</emphasis>
.—Dorsum brown with dark brown or black blotches, occasionally forming chevrons in males; light lateral stripe extends posteriorly from subocular region to dorsal aspect of tympanum and continues dorsolaterally to approximately the level of fore limb insertion; gular region and venter beige in males and grayish beige in females; ventrally, pelvic region and thighs yellow in males (
<bibRefCitation id="AA454B76FFFAFFD6FD1CFB671514FB42" author="FRITTS, T. H." box="[672,806,1187,1211]" journalOrPublisher="Ocassional Papers University of Kansas Museum of Natural History" pageId="65" pageNumber="140" pagination="1 - 21" part="10" refId="ref77696" refString="FRITTS, T. H. 1972. New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae). Ocassional Papers University of Kansas Museum of Natural History 10: 1 - 21." title="New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae)" type="journal article" year="1972">Fritts, 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFAFFD6FD0FFB051545FAED" pageId="65" pageNumber="140" type="biology_ecology">
<paragraph id="CE6B3687FFFAFFD6FD0FFB051545FAED" blockId="65.[664,1175,190,1506]" pageId="65" pageNumber="140">
<emphasis id="FCA0EA95FFFAFFD6FD0FFB051559FB20" box="[691,875,1217,1241]" italics="true" pageId="65" pageNumber="140">Natural History</emphasis>
.—This species has been found in open areas at the bases of shrubs (
<bibRefCitation id="AA454B76FFFAFFD6FD1CFB381517FAED" author="FRITTS, T. H." box="[672,805,1276,1300]" journalOrPublisher="Ocassional Papers University of Kansas Museum of Natural History" pageId="65" pageNumber="140" pagination="1 - 21" part="10" refId="ref77696" refString="FRITTS, T. H. 1972. New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae). Ocassional Papers University of Kansas Museum of Natural History 10: 1 - 21." title="New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae)" type="journal article" year="1972">Fritts, 1972</bibRefCitation>
,
<bibRefCitation id="AA454B76FFFAFFD6FC8EFB381559FAED" author="FRITTS, T. H." box="[818,875,1276,1300]" journalOrPublisher="San Diego Society of Natural History Memoir" pageId="65" pageNumber="140" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">1974</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFAFFD6FD0FFADE154AFA1B" pageId="65" pageNumber="140" type="distribution">
<paragraph id="CE6B3687FFFAFFD6FD0FFADE154AFA1B" blockId="65.[664,1175,190,1506]" pageId="65" pageNumber="140">
<emphasis id="FCA0EA95FFFAFFD6FD0FFADE1509FACB" box="[691,827,1306,1330]" italics="true" pageId="65" pageNumber="140">Distribution</emphasis>
.—
<taxonomicName id="09D44D04FFFAFFD6FCE0FADF1257FACB" authorityName="Fritts" authorityYear="1972" box="[860,1125,1306,1330]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="65" pageNumber="140" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="FCA0EA95FFFAFFD6FCE0FADF1257FACB" box="[860,1125,1306,1330]" italics="true" pageId="65" pageNumber="140">Stenocercus orientalis</emphasis>
</taxonomicName>
is known from the eastern Cordillera of the central Andes in northern Peru (6
<emphasis id="FCA0EA95FFFAFFD6FB87FA911277FA94" bold="true" box="[1083,1093,1365,1389]" pageId="65" pageNumber="140">°</emphasis>
S5
<emphasis id="FCA0EA95FFFAFFD6FBD2FA91124AFA94" bold="true" box="[1134,1144,1365,1389]" pageId="65" pageNumber="140">°</emphasis>
S). This species occurs in the valley of Río Utcubamba (Atlantic drainage) at elevations between 22002900 m in Departamento Amazonas (
<figureCitation id="56EF2A02FFFAFFD6FCA5FA0E1559FA1B" box="[793,875,1482,1506]" captionStart="FIG" captionStartId="51.[319,333,1042,1061]" captionTargetBox="[122,1176,189,1025]" captionTargetId="figure-299@51.[122,1176,189,1025]" captionTargetPageId="51" captionText="FIG. 19.—Distribution of six species of Stenocercus in Ecuador and Peru." figureDoi="http://doi.org/10.5281/zenodo.14384848" httpUri="https://zenodo.org/record/14384848/files/figure.png" pageId="65" pageNumber="140">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>