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<document id="44206D0FA712CAA247039887288F9D99" ID-DOI="10.11646/zootaxa.1479.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10088161" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625847322231" checkinUser="felipe" docAuthor="Pilato, Giovanni &amp; Kaczmarek, Łukasz" docDate="2007" docId="154287C2FF8AFFDCFF019A65C7F3D0CE" docLanguage="en" docName="zt01479p007.pdf.imf" docOrigin="Zootaxa 1479 (1)" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Macrobiotus alvaroi Pilato &amp; Kaczmarek 2007, sp. n." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="E97BFFBAFF8BFFD9FF969F70C663D73A" masterDocTitle="Macrobiotus alvaroi, a new species of eutardigrade (Tardigrada, Macrobiotidae) of the polyopus group from Costa Rica (Central America)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1699509752474" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="97EFE83B811EAAB1A061C7F082278763" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B068DB306011E473B30ACAC8A23247C7">
<mods:title id="561A7488B40AE53143CBE3098D132FD7">Macrobiotus alvaroi, a new species of eutardigrade (Tardigrada, Macrobiotidae) of the polyopus group from Costa Rica (Central America)</mods:title>
</mods:titleInfo>
<mods:name id="0A3F6AE765FB4E95C089B1079BE6EB17" type="personal">
<mods:role id="97EBCC0B74B3391BAD9D16C1F63F605E">
<mods:roleTerm id="5B0423ADF6C92B9F52EDE801C1F578D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57860F7A74BD0DD4A96838399F5F47AB">Pilato, Giovanni</mods:namePart>
<mods:affiliation id="49D3E6C6E6DE6EE83BC97C63A4BA140E">Department of Animal Biology “ Marcello La Greca ”, University of Catania, Italy. E-mail: pilato @ unict. it Department of Animal Taxonomy &amp; Ecology, Institute of Environmental Biology, A. Mickiewicz University, Umultowska 89, 61 - 614 Pozna, Poland. E-mail: kczmar @ amu. edu. pl</mods:affiliation>
</mods:name>
<mods:name id="6F1195D9C2CFB4AD67C2845AEFA976CF" type="personal">
<mods:role id="E2D92DE06CAC2AFC2EF18462100113A9">
<mods:roleTerm id="15B7F3E9103A7633913A04DD035475E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E43EA855E9B9F013C7E936934E93EF2">Kaczmarek, Łukasz</mods:namePart>
</mods:name>
<mods:typeOfResource id="DCCAEADAED66C8E537AB7DDDFA1B256B">text</mods:typeOfResource>
<mods:relatedItem id="E7D7635CFC37A80C11B6B4DC4F483925" type="host">
<mods:titleInfo id="7E0DBB5F35FCF67106F431052DC5817D">
<mods:title id="682B2C479BA17B7681D6DAEAAF925144">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0C1E9A0E25DDB4545617035359AF9B7B">
<mods:date id="1084964DD443CF1517C2EE7E524F2B20">2007</mods:date>
<mods:detail id="5605421ACFD674F8F9BAD265FC5FDF9C" type="pubDate">
<mods:number id="8A593459FFC7D477B198ED336E04F692">2007-05-21</mods:number>
</mods:detail>
<mods:detail id="8EEDE35AEF1A7646F90D06687C13FB22" type="volume">
<mods:number id="6B7951D42A56AC0B6725C26FA0AB29E8">1479</mods:number>
</mods:detail>
<mods:detail id="8AF8E8AB4C84D2A5860DAD90D75EF83E" type="issue">
<mods:number id="8FD37807CB19353B3C42C885A5AFF48A">1</mods:number>
</mods:detail>
<mods:extent id="223B53A230961973DA3CAE27EABF64AD" unit="page">
<mods:start id="B9E85F5FDDF9607F60131EECBD69DECB">1</mods:start>
<mods:end id="8ADFAA919B4133F0D8A6DC329441980A">7</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6AD8E97837AD6865FA2C42922F1D4BFC">journal article</mods:classification>
<mods:identifier id="BB68B5E33A709B9E965FFFAC3E226ED5" type="DOI">10.11646/zootaxa.1479.1.1</mods:identifier>
<mods:identifier id="231C01042D039684C692953B8E86D3A2" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="261789CE585627172A59184C99770F22" type="Zenodo-Dep">10088161</mods:identifier>
<mods:identifier id="7CE07C85E674797201ACC0F6ADDBA6ED" type="ZooBank">E7356049-19D8-43C0-88B8-8FFDE373F7BF</mods:identifier>
</mods:mods>
<treatment id="154287C2FF8AFFDCFF019A65C7F3D0CE" LSID="urn:lsid:plazi:treatment:154287C2FF8AFFDCFF019A65C7F3D0CE" httpUri="http://treatment.plazi.org/id/154287C2FF8AFFDCFF019A65C7F3D0CE" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
<subSubSection id="D5F1655FFF8AFFD8FF019A65C7B6D215" box="[151,469,1301,1327]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="9D5436D4FF8AFFD8FF019A65C7B6D215" blockId="1.[151,469,1301,1362]" box="[151,469,1301,1327]" pageId="1" pageNumber="2">
<heading id="C61C81B8FF8AFFD8FF019A65C7B6D215" bold="true" box="[151,469,1301,1327]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="AF9FEAC6FF8AFFD8FF019A65C7B6D215" bold="true" box="[151,469,1301,1327]" pageId="1" pageNumber="2">
<taxonomicName id="5AEB4D57FF8AFFD8FF019A65C7EFD215" ID-CoL="7bc18580-e951-4a38-b107-9bf3ee202066" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[151,396,1301,1327]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="2" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8AFFD8FF019A65C7EFD215" bold="true" box="[151,396,1301,1327]" italics="true" pageId="1" pageNumber="2">Macrobiotus alvaroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4AC57BDFF8AFFD8FE059A65C7B6D215" box="[403,469,1301,1327]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D5F1655FFF8AFFD8FF019A48C714D268" box="[151,375,1336,1362]" pageId="1" pageNumber="2" type="description">
<paragraph id="9D5436D4FF8AFFD8FF019A48C714D268" blockId="1.[151,469,1301,1362]" box="[151,375,1336,1362]" pageId="1" pageNumber="2">
(
<figureCitation id="05D02A51FF8AFFD8FF099A48C687D268" box="[159,228,1336,1362]" captionStart="FIGURE 1" captionStartId="2.[151,255,1898,1922]" captionTargetBox="[182,1404,473,1862]" captionTargetId="figure-188@2.[174,1409,467,1875]" captionTargetPageId="2" captionText="FIGURE 1. Macrobiotus alvaroi sp. n. A–B, buccopharyngeal apparatus (holotype). C, claws of the second and third pairs of legs. D, egg (the reticular design of the egg shell is focused). E, detail of the egg (the shape of the egg processes is visible). Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088163" httpUri="https://zenodo.org/record/10088163/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
,
<figureCitation id="05D02A51FF8AFFD8FF659A48C70DD268" box="[243,366,1336,1362]" captionStart="FIGURE 2" captionStartId="4.[151,255,1978,2002]" captionTargetBox="[233,1364,184,1960]" captionTargetId="figure-47@4.[225,1372,178,1977]" captionTargetPageId="4" captionText="FIGURE 2. Eggs, and their details, of: A–B, M. insularis. C–D, M. ocotensis. E–F, M. mandalae. G–H, M. alvaroi sp. n. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088165" httpUri="https://zenodo.org/record/10088165/files/figure.png" pageId="1" pageNumber="2">Fig. 2G, H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D5F1655FFF8AFFD8FF019AF8C403D2A5" box="[151,608,1416,1439]" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="9D5436D4FF8AFFD8FF019AF8C403D2A5" blockId="1.[151,608,1416,1439]" box="[151,608,1416,1439]" pageId="1" pageNumber="2">
<emphasis id="AF9FEAC6FF8AFFD8FF019AF8C6C6D2A5" box="[151,165,1416,1439]" italics="true" pageId="1" pageNumber="2">=</emphasis>
<taxonomicName id="5AEB4D57FF8AFFD8FF339AF8C403D2A5" ID-CoL="3X2V7" authority="Kaczmarek, 2003" authorityName="Kaczmarek" authorityYear="2003" box="[165,608,1416,1439]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="2" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="AF9FEAC6FF8AFFD8FF339AF8C7F4D2A5" box="[165,407,1416,1439]" italics="true" pageId="1" pageNumber="2">Macrobiotus polyopus</emphasis>
<bibRefCitation id="F97A4B25FF8AFFD8FE0B9AF8C403D2A5" author="Kaczmarek, L." box="[413,608,1416,1439]" pageId="1" pageNumber="2" pagination="1 - 4" refId="ref3219" refString="Kaczmarek, L. (2003) New records of and key to Tardigrada from Costa Rica. Zootaxa, 177, 1 - 4." type="journal article" year="2003">Kaczmarek, 2003</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D5F1655FFF8AFFDCFF019AA0C7F3D0CE" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="9D5436D4FF8AFFD8FF019AA0C702D128" blockId="1.[151,1436,1488,2034]" pageId="1" pageNumber="2">
<emphasis id="AF9FEAC6FF8AFFD8FF019AA0C7E3D2D3" box="[151,384,1488,1513]" italics="true" pageId="1" pageNumber="2">Material examined:</emphasis>
<specimenCount id="8BEDFD5DFF8AFFD8FE1C9AA0C47FD2D0" box="[394,540,1488,1514]" pageId="1" pageNumber="2" type="generic">8 specimens</specimenCount>
and
<specimenCount id="8BEDFD5DFF8AFFD8FDCF9AA0C4C4D2D0" box="[601,679,1488,1514]" pageId="1" pageNumber="2" type="egg">6 eggs</specimenCount>
(
<specimenCount id="8BEDFD5DFF8AFFD8FD2E9AA0C574D2D0" box="[696,791,1488,1514]" pageId="1" pageNumber="2" type="egg">one egg</specimenCount>
with embryo), additional material examined:
<specimenCount id="8BEDFD5DFF8AFFD8FAA89AA0C6B6D128" pageId="1" pageNumber="2" type="generic">2 specimens</specimenCount>
and
<specimenCount id="8BEDFD5DFF8AFFD8FE9B9A88C73ED128" box="[269,349,1528,1554]" pageId="1" pageNumber="2" type="egg">3 eggs</specimenCount>
.
</paragraph>
<paragraph id="9D5436D4FF8AFFD8FF519950C4A4D068" blockId="1.[151,1436,1488,2034]" pageId="1" pageNumber="2">
<emphasis id="AF9FEAC6FF8AFFD8FF519950C7FAD103" box="[199,409,1568,1593]" italics="true" pageId="1" pageNumber="2">Species diagnosis</emphasis>
: cuticle smooth with circular and elliptical pores, without dots on the legs; eye spots present; ventral lamina longer than the double length of the stylet sheaths; buccal armature with peribuccal lamellae and a system of very thin transversal ridges (three dorsal and three ventral); pharyngeal bulb with apophyses, two rod-shaped macroplacoids and a small microplacoid; claws of
<emphasis id="AF9FEAC6FF8AFFD8FBDF99E8C2DBD18B" box="[1097,1208,1688,1713]" italics="true" pageId="1" pageNumber="2">hufelandi</emphasis>
<typeStatus id="42508876FF8AFFD8FB5499E8C297D188" box="[1218,1268,1688,1714]" pageId="1" pageNumber="2">type</typeStatus>
(the common basal portion short); smooth lunules present; eggs, laid freely, with 23–26 small processes around the circumference and about
<quantity id="5A139B31FF8AFFD8FEE29998C7A2D038" box="[372,449,1768,1794]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="2" unit="in" value="100.0">100 in</quantity>
the hemisphere; egg processes with a basal portion in shape of a hexagonal prism and a conical distal portion with an apical, sharpened, point with almost invisible annulations; egg shell with a regular reticular design with hexagonal meshes.
</paragraph>
<paragraph id="9D5436D4FF8AFFD8FF519810C492D098" blockId="1.[151,1436,1488,2034]" pageId="1" pageNumber="2">
<emphasis id="AF9FEAC6FF8AFFD8FF519810C466D043" box="[199,517,1888,1913]" italics="true" pageId="1" pageNumber="2">
Description of the
<typeStatus id="42508876FF8AFFD8FE369810C466D043" box="[416,517,1888,1913]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
</emphasis>
: Body length 290.0, colourless, eye spots present, cuticle smooth with circular and elliptical pores. Granulation on the legs absent.
</paragraph>
<paragraph id="9D5436D4FF8AFFDBFF5198C0C426D610" blockId="1.[151,1436,1488,2034]" lastBlockId="2.[151,1436,152,418]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
Mouth opening terminal with 10 small peribuccal lamellae, buccal armature very simple and difficult to see due to the small dimensions of the buccal cavity; system of very thin transversal ridges visible (three dor- sal and three ventral; medioventral ridge very short). Buccal tube 30.8 long and 3.1 wide [
<emphasis id="AF9FEAC6FF89FFDBFB289FE8C292D78B" box="[1214,1265,152,177]" italics="true" pageId="2" pageNumber="3">10.1</emphasis>
] (
<figureCitation id="05D02A51FF89FFDBFA9A9FE8C3EFD788" box="[1292,1420,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1898,1922]" captionTargetBox="[182,1404,473,1862]" captionTargetId="figure-188@2.[174,1409,467,1875]" captionTargetPageId="2" captionText="FIGURE 1. Macrobiotus alvaroi sp. n. A–B, buccopharyngeal apparatus (holotype). C, claws of the second and third pairs of legs. D, egg (the reticular design of the egg shell is focused). E, detail of the egg (the shape of the egg processes is visible). Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088163" httpUri="https://zenodo.org/record/10088163/files/figure.png" pageId="2" pageNumber="3">Fig. 1A, B</figureCitation>
); ventral lamina 19.1 [
<emphasis id="AF9FEAC6FF89FFDBFE199FB0C7A1D7E3" box="[399,450,192,217]" italics="true" pageId="2" pageNumber="3">62.0</emphasis>
] long (longer than the double length of the stylet sheaths). Stylet supports inserted on the buccal tube at 72.9 % of its length. Large thickening, to which the pharyngeal apophyses are connected, present at the end of the buccal tube.
</paragraph>
<paragraph id="9D5436D4FF89FFDBFF519E48C2C7D698" blockId="2.[151,1436,152,418]" pageId="2" pageNumber="3">
Pharyngeal bulb (
<figureCitation id="05D02A51FF89FFDBFE039E48C78DD668" box="[405,494,312,338]" captionStart="FIGURE 1" captionStartId="2.[151,255,1898,1922]" captionTargetBox="[182,1404,473,1862]" captionTargetId="figure-188@2.[174,1409,467,1875]" captionTargetPageId="2" captionText="FIGURE 1. Macrobiotus alvaroi sp. n. A–B, buccopharyngeal apparatus (holotype). C, claws of the second and third pairs of legs. D, egg (the reticular design of the egg shell is focused). E, detail of the egg (the shape of the egg processes is visible). Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088163" httpUri="https://zenodo.org/record/10088163/files/figure.png" pageId="2" pageNumber="3">Fig. 1B</figureCitation>
) with apophyses, two rod-shaped macroplacoids and a small microplacoid. First macroplacoid with a central constriction, 7.1 [
<emphasis id="AF9FEAC6FF89FFDBFD559E10C495D643" box="[707,758,352,377]" italics="true" pageId="2" pageNumber="3">23.1</emphasis>
] long, second one shorter 4.7 [
<emphasis id="AF9FEAC6FF89FFDBFBE49E10C2C6D643" box="[1138,1189,352,377]" italics="true" pageId="2" pageNumber="3">15.2</emphasis>
] long. Microplacoid small, 1.6 [
<emphasis id="AF9FEAC6FF89FFDBFE8D9EF8C75CD69B" box="[283,319,392,417]" italics="true" pageId="2" pageNumber="3">5.2</emphasis>
] long. Entire placoid row 14.7 [
<emphasis id="AF9FEAC6FF89FFDBFD2F9EF8C48FD69B" box="[697,748,392,417]" italics="true" pageId="2" pageNumber="3">47.7</emphasis>
] long, macroplacoid row 12.7 [
<emphasis id="AF9FEAC6FF89FFDBFBF69EF8C2F0D69B" box="[1120,1171,392,417]" italics="true" pageId="2" pageNumber="3">41.2</emphasis>
].
</paragraph>
<caption id="C994665CFF89FFDBFF01981AC7BBD0FD" ID-DOI="http://doi.org/10.5281/zenodo.10088163" ID-Zenodo-Dep="10088163" httpUri="https://zenodo.org/record/10088163/files/figure.png" pageId="2" pageNumber="3" startId="2.[151,255,1898,1922]" targetBox="[182,1404,473,1862]" targetPageId="2">
<paragraph id="9D5436D4FF89FFDBFF01981AC7BBD0FD" blockId="2.[151,1436,1898,1991]" pageId="2" pageNumber="3">
<emphasis id="AF9FEAC6FF89FFDBFF01981AC778D0B8" bold="true" box="[151,283,1898,1922]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
<taxonomicName id="5AEB4D57FF89FFDBFEB5981BC79CD0B8" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[291,511,1899,1922]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="3" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF89FFDBFEB5981BC79CD0B8" box="[291,511,1899,1922]" italics="true" pageId="2" pageNumber="3">Macrobiotus alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF89FFDBFD91981AC426D0B8" bold="true" box="[519,581,1898,1922]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="B4AC57BDFF89FFDBFD91981AC426D0B8" box="[519,581,1898,1922]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A–B, buccopharyngeal apparatus (holotype). C, claws of the second and third pairs of legs. D, egg (the reticular design of the egg shell is focused). E, detail of the egg (the shape of the egg processes is visible). Scale bar = 10 µm.
</paragraph>
</caption>
<paragraph id="9D5436D4FF88FFDAFF519FE8C23CD668" blockId="3.[151,1436,152,418]" pageId="3" pageNumber="4">
Claws of
<emphasis id="AF9FEAC6FF88FFDAFEAA9FE8C7C9D78B" box="[316,426,152,177]" italics="true" pageId="3" pageNumber="4">hufelandi</emphasis>
<typeStatus id="42508876FF88FFDAFE239FE8C789D788" box="[437,490,152,178]" pageId="3" pageNumber="4">type</typeStatus>
, short (particularly the common basal portion) but not weak (
<figureCitation id="05D02A51FF88FFDAFB409FE8C349D788" box="[1238,1322,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1898,1922]" captionTargetBox="[182,1404,473,1862]" captionTargetId="figure-188@2.[174,1409,467,1875]" captionTargetPageId="2" captionText="FIGURE 1. Macrobiotus alvaroi sp. n. A–B, buccopharyngeal apparatus (holotype). C, claws of the second and third pairs of legs. D, egg (the reticular design of the egg shell is focused). E, detail of the egg (the shape of the egg processes is visible). Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088163" httpUri="https://zenodo.org/record/10088163/files/figure.png" pageId="3" pageNumber="4">Fig.1C</figureCitation>
); internal and external claws of the second and third pairs of legs 7.6 [
<emphasis id="AF9FEAC6FF88FFDAFCFB9FB0C5C3D7E3" box="[877,928,192,217]" italics="true" pageId="3" pageNumber="4">24.7</emphasis>
] and 8.0 [
<emphasis id="AF9FEAC6FF88FFDAFB889FB0C232D7E3" box="[1054,1105,192,217]" italics="true" pageId="3" pageNumber="4">26.0</emphasis>
] long respectively; anterior and posterior claws of the fourth pair of legs 8.5 [
<emphasis id="AF9FEAC6FF88FFDAFD639F98C54BD63B" box="[757,808,232,257]" italics="true" pageId="3" pageNumber="4">27.6</emphasis>
] and 8.7 [
<emphasis id="AF9FEAC6FF88FFDAFC3E9F98C5B8D63B" box="[936,987,232,257]" italics="true" pageId="3" pageNumber="4">28.2</emphasis>
] long respectively. Main branches of claws with well developed accessory points. Small, smooth, lunules present on all legs. Two faint cuticular thickenings, difficult to see, present near the lunules on the first three pairs of legs.
</paragraph>
<paragraph id="9D5436D4FF88FFDAFF519E10C2A2D698" blockId="3.[151,1436,152,418]" pageId="3" pageNumber="4">
The
<typeStatus id="42508876FF88FFDAFF699E10C70CD640" box="[255,367,352,378]" pageId="3" pageNumber="4" type="paratype">paratypes</typeStatus>
are similar to the
<typeStatus id="42508876FF88FFDAFDC09E10C4DED640" box="[598,701,352,378]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
in both qualitative and metric characters; the dimensions of some structures of the smallest and of the largest measured specimens are given in
<tableCitation id="D069036FFF88FFDAFBF59EF8C2D8D698" box="[1123,1211,392,418]" captionStart="TABLE 1" captionStartId="3.[151,241,471,495]" captionText="TABLE 1. Measurements [in µm] and pt values of selected morphological structures of the smallest and the largest (the holotype) measured specimens of Macrobiotus alvaroi sp. n. mounted in Hoyer’s medium." pageId="3" pageNumber="4">Table 1</tableCitation>
.
</paragraph>
<caption id="C994665CFF88FFDAFF019EA7C203D52B" pageId="3" pageNumber="4" startId="3.[151,241,471,495]">
<paragraph id="9D5436D4FF88FFDAFF019EA7C203D52B" blockId="3.[151,1436,471,530]" pageId="3" pageNumber="4">
<emphasis id="AF9FEAC6FF88FFDAFF019EA7C769D6D5" bold="true" box="[151,266,471,495]" pageId="3" pageNumber="4">TABLE 1.</emphasis>
Measurements [in µm] and
<emphasis id="AF9FEAC6FF88FFDAFDAE9EA8C42ED6D5" box="[568,589,472,495]" italics="true" pageId="3" pageNumber="4">pt</emphasis>
values of selected morphological structures of the smallest and the largest (the holotype) measured specimens of
<taxonomicName id="5AEB4D57FF88FFDAFD959E8AC4BED52B" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[515,733,506,529]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="4" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF88FFDAFD959E8AC4BED52B" box="[515,733,506,529]" italics="true" pageId="3" pageNumber="4">Macrobiotus alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF88FFDAFD729E8AC57CD528" bold="true" box="[740,799,506,530]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="B4AC57BDFF88FFDAFD729E8AC57CD528" box="[740,799,506,530]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
mounted in Hoyer’s medium.
</paragraph>
</caption>
<paragraph id="9D5436D4FF88FFDAFF099D42C282D3A5" pageId="3" pageNumber="4">
<table id="EFEBC474FF880026FF099D42C282D3A5" box="[159,1249,562,1183]" gridcols="5" gridrows="15" pageId="3" pageNumber="4">
<tr id="23DB3496FF880026FF099D42C282D573" box="[159,1249,562,585]" gridrow="0" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099D42C7B6D573" box="[159,469,562,585]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">Character</th>
<th id="600A5DEAFF880026FD9C9D42C425D573" box="[522,582,562,585]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">µm</th>
<th id="600A5DEAFF880026FD739D42C577D573" box="[741,788,562,585]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">pt</th>
<th id="600A5DEAFF880026FC569D42C59FD573" box="[960,1020,562,585]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">µm</th>
<th id="600A5DEAFF880026FB249D42C282D573" box="[1202,1249,562,585]" gridcol="4" gridrow="0" pageId="3" pageNumber="4">pt</th>
</tr>
<tr id="23DB3496FF880026FF099D2DC282D54E" box="[159,1249,605,628]" gridrow="1" pageId="3" pageNumber="4" rowspan-2="1">
<th id="600A5DEAFF880026FF099D2DC7B6D54E" box="[159,469,605,628]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Body length</th>
<td id="600A5DEAFF880026FD9C9D2DC425D54E" box="[522,582,605,628]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">261.0</td>
<td id="600A5DEAFF880026FC569D2DC59FD54E" box="[960,1020,605,628]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">290.0</td>
<td id="600A5DEAFF880026FB249D2DC282D54E" box="[1202,1249,605,628]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">-</td>
</tr>
<tr id="23DB3496FF880026FF099DF8C282D5A5" box="[159,1249,648,671]" gridrow="2" pageId="3" pageNumber="4" rowspan-2="1">
<th id="600A5DEAFF880026FF099DF8C7B6D5A5" box="[159,469,648,671]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Buccal tube length</th>
<td id="600A5DEAFF880026FD9C9DF8C425D5A5" box="[522,582,648,671]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">29.3</td>
<td id="600A5DEAFF880026FC569DF8C59FD5A5" box="[960,1020,648,671]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">30.8</td>
<td id="600A5DEAFF880026FB249DF8C282D5A5" box="[1202,1249,648,671]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">-</td>
</tr>
<tr id="23DB3496FF880026FF099DC2C282D5F3" box="[159,1249,690,713]" gridrow="3" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099DC2C7B6D5F3" box="[159,469,690,713]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Stylet support insertion point</th>
<td id="600A5DEAFF880026FD9C9DC2C425D5F3" box="[522,582,690,713]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">21.5</td>
<td id="600A5DEAFF880026FD739DC2C577D5F3" box="[741,788,690,713]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">73.4</td>
<td id="600A5DEAFF880026FC569DC2C59FD5F3" box="[960,1020,690,713]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">-</td>
<td id="600A5DEAFF880026FB249DC2C282D5F3" box="[1202,1249,690,713]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">72.9</td>
</tr>
<tr id="23DB3496FF880026FF099DADC282D5CE" box="[159,1249,733,756]" gridrow="4" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099DADC7B6D5CE" box="[159,469,733,756]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Buccal tube external width</th>
<td id="600A5DEAFF880026FD9C9DADC425D5CE" box="[522,582,733,756]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">3.2</td>
<td id="600A5DEAFF880026FD739DADC577D5CE" box="[741,788,733,756]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">10.9</td>
<td id="600A5DEAFF880026FC569DADC59FD5CE" box="[960,1020,733,756]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">3.1</td>
<td id="600A5DEAFF880026FB249DADC282D5CE" box="[1202,1249,733,756]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">10.1</td>
</tr>
<tr id="23DB3496FF880026FF099C78C282D425" box="[159,1249,776,799]" gridrow="5" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099C78C7B6D425" box="[159,469,776,799]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Ventral lamina</th>
<td id="600A5DEAFF880026FD9C9C78C425D425" box="[522,582,776,799]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">17.1</td>
<td id="600A5DEAFF880026FD739C78C577D425" box="[741,788,776,799]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">58.4</td>
<td id="600A5DEAFF880026FC569C78C59FD425" box="[960,1020,776,799]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">19.1</td>
<td id="600A5DEAFF880026FB249C78C282D425" box="[1202,1249,776,799]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">62.0</td>
</tr>
<tr id="23DB3496FF880026FF099C42C282D473" box="[159,1249,818,841]" gridrow="6" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099C42C7B6D473" box="[159,469,818,841]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">Macroplacoid 1</th>
<td id="600A5DEAFF880026FD9C9C42C425D473" box="[522,582,818,841]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">6.3</td>
<td id="600A5DEAFF880026FD739C42C577D473" box="[741,788,818,841]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">21.5</td>
<td id="600A5DEAFF880026FC569C42C59FD473" box="[960,1020,818,841]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">7.1</td>
<td id="600A5DEAFF880026FB249C42C282D473" box="[1202,1249,818,841]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">23.1</td>
</tr>
<tr id="23DB3496FF880026FF099C2DC282D44E" box="[159,1249,861,884]" gridrow="7" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099C2DC7B6D44E" box="[159,469,861,884]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">Macroplacoid 2</th>
<td id="600A5DEAFF880026FD9C9C2DC425D44E" box="[522,582,861,884]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">4.2</td>
<td id="600A5DEAFF880026FD739C2DC577D44E" box="[741,788,861,884]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">14.3</td>
<td id="600A5DEAFF880026FC569C2DC59FD44E" box="[960,1020,861,884]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">4.7</td>
<td id="600A5DEAFF880026FB249C2DC282D44E" box="[1202,1249,861,884]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">15.2</td>
</tr>
<tr id="23DB3496FF880026FF099CF8C282D4A5" box="[159,1249,904,927]" gridrow="8" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099CF8C7B6D4A5" box="[159,469,904,927]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">Microplacoid</th>
<td id="600A5DEAFF880026FD9C9CF8C425D4A5" box="[522,582,904,927]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">2.1</td>
<td id="600A5DEAFF880026FD739CF8C577D4A5" box="[741,788,904,927]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">7.2</td>
<td id="600A5DEAFF880026FC569CF8C59FD4A5" box="[960,1020,904,927]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">1.6</td>
<td id="600A5DEAFF880026FB249CF8C282D4A5" box="[1202,1249,904,927]" gridcol="4" gridrow="8" pageId="3" pageNumber="4">5.2</td>
</tr>
<tr id="23DB3496FF880026FF099CC2C282D4F3" box="[159,1249,946,969]" gridrow="9" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099CC2C7B6D4F3" box="[159,469,946,969]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">Macroplacoid row</th>
<td id="600A5DEAFF880026FD9C9CC2C425D4F3" box="[522,582,946,969]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">12.0</td>
<td id="600A5DEAFF880026FD739CC2C577D4F3" box="[741,788,946,969]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">41.0</td>
<td id="600A5DEAFF880026FC569CC2C59FD4F3" box="[960,1020,946,969]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">12.7</td>
<td id="600A5DEAFF880026FB249CC2C282D4F3" box="[1202,1249,946,969]" gridcol="4" gridrow="9" pageId="3" pageNumber="4">41.2</td>
</tr>
<tr id="23DB3496FF880026FF099CADC282D4CE" box="[159,1249,989,1012]" gridrow="10" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099CADC7B6D4CE" box="[159,469,989,1012]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">Placoid row</th>
<td id="600A5DEAFF880026FD9C9CADC425D4CE" box="[522,582,989,1012]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">13.7</td>
<td id="600A5DEAFF880026FD739CADC577D4CE" box="[741,788,989,1012]" gridcol="2" gridrow="10" pageId="3" pageNumber="4">46.8</td>
<td id="600A5DEAFF880026FC569CADC59FD4CE" box="[960,1020,989,1012]" gridcol="3" gridrow="10" pageId="3" pageNumber="4">14.7</td>
<td id="600A5DEAFF880026FB249CADC282D4CE" box="[1202,1249,989,1012]" gridcol="4" gridrow="10" pageId="3" pageNumber="4">47.7</td>
</tr>
<tr id="23DB3496FF880026FF099B78C282D325" box="[159,1249,1032,1055]" gridrow="11" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099B78C7B6D325" box="[159,469,1032,1055]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">Internal claws II and III</th>
<td id="600A5DEAFF880026FD9C9B78C425D325" box="[522,582,1032,1055]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">-</td>
<td id="600A5DEAFF880026FD739B78C577D325" box="[741,788,1032,1055]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">-</td>
<td id="600A5DEAFF880026FC569B78C59FD325" box="[960,1020,1032,1055]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">7.6</td>
<td id="600A5DEAFF880026FB249B78C282D325" box="[1202,1249,1032,1055]" gridcol="4" gridrow="11" pageId="3" pageNumber="4">24.7</td>
</tr>
<tr id="23DB3496FF880026FF099B42C282D373" box="[159,1249,1074,1097]" gridrow="12" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099B42C7B6D373" box="[159,469,1074,1097]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">External claws II and III</th>
<td id="600A5DEAFF880026FD9C9B42C425D373" box="[522,582,1074,1097]" gridcol="1" gridrow="12" pageId="3" pageNumber="4">8.0</td>
<td id="600A5DEAFF880026FD739B42C577D373" box="[741,788,1074,1097]" gridcol="2" gridrow="12" pageId="3" pageNumber="4">27.3</td>
<td id="600A5DEAFF880026FC569B42C59FD373" box="[960,1020,1074,1097]" gridcol="3" gridrow="12" pageId="3" pageNumber="4">8.0</td>
<td id="600A5DEAFF880026FB249B42C282D373" box="[1202,1249,1074,1097]" gridcol="4" gridrow="12" pageId="3" pageNumber="4">26.0</td>
</tr>
<tr id="23DB3496FF880026FF099B2DC282D34E" box="[159,1249,1117,1140]" gridrow="13" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099B2DC7B6D34E" box="[159,469,1117,1140]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">Anterior claws IV</th>
<td id="600A5DEAFF880026FD9C9B2DC425D34E" box="[522,582,1117,1140]" gridcol="1" gridrow="13" pageId="3" pageNumber="4">-</td>
<td id="600A5DEAFF880026FD739B2DC577D34E" box="[741,788,1117,1140]" gridcol="2" gridrow="13" pageId="3" pageNumber="4">-</td>
<td id="600A5DEAFF880026FC569B2DC59FD34E" box="[960,1020,1117,1140]" gridcol="3" gridrow="13" pageId="3" pageNumber="4">8.5</td>
<td id="600A5DEAFF880026FB249B2DC282D34E" box="[1202,1249,1117,1140]" gridcol="4" gridrow="13" pageId="3" pageNumber="4">27.6</td>
</tr>
<tr id="23DB3496FF880026FF099BF8C282D3A5" box="[159,1249,1160,1183]" gridrow="14" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099BF8C7B6D3A5" box="[159,469,1160,1183]" gridcol="0" gridrow="14" pageId="3" pageNumber="4">Posterior claws IV</th>
<td id="600A5DEAFF880026FD9C9BF8C425D3A5" box="[522,582,1160,1183]" gridcol="1" gridrow="14" pageId="3" pageNumber="4">9,7</td>
<td id="600A5DEAFF880026FD739BF8C577D3A5" box="[741,788,1160,1183]" gridcol="2" gridrow="14" pageId="3" pageNumber="4">29.7</td>
<td id="600A5DEAFF880026FC569BF8C59FD3A5" box="[960,1020,1160,1183]" gridcol="3" gridrow="14" pageId="3" pageNumber="4">8.7</td>
<td id="600A5DEAFF880026FB249BF8C282D3A5" box="[1202,1249,1160,1183]" gridcol="4" gridrow="14" pageId="3" pageNumber="4">28.2</td>
</tr>
</table>
</paragraph>
<paragraph id="9D5436D4FF88FFDAFF519B80C771D2E8" blockId="3.[151,1436,1264,1570]" pageId="3" pageNumber="4">
Eggs laid freely. Spherical, with large number of small processes (
<figureCitation id="05D02A51FF88FFDAFC4E9B80C23AD230" box="[984,1113,1264,1290]" captionStart="FIGURE 1" captionStartId="2.[151,255,1898,1922]" captionTargetBox="[182,1404,473,1862]" captionTargetId="figure-188@2.[174,1409,467,1875]" captionTargetPageId="2" captionText="FIGURE 1. Macrobiotus alvaroi sp. n. A–B, buccopharyngeal apparatus (holotype). C, claws of the second and third pairs of legs. D, egg (the reticular design of the egg shell is focused). E, detail of the egg (the shape of the egg processes is visible). Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088163" httpUri="https://zenodo.org/record/10088163/files/figure.png" pageId="3" pageNumber="4">Fig. 1D, E</figureCitation>
;
<figureCitation id="05D02A51FF88FFDAFBF09B80C280D230" box="[1126,1251,1264,1290]" captionStart="FIGURE 2" captionStartId="4.[151,255,1978,2002]" captionTargetBox="[233,1364,184,1960]" captionTargetId="figure-47@4.[225,1372,178,1977]" captionTargetPageId="4" captionText="FIGURE 2. Eggs, and their details, of: A–B, M. insularis. C–D, M. ocotensis. E–F, M. mandalae. G–H, M. alvaroi sp. n. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088165" httpUri="https://zenodo.org/record/10088165/files/figure.png" pageId="3" pageNumber="4">Fig. 2G, H</figureCitation>
) (23-26 around the circumference, about
<quantity id="5A139B31FF88FFDAFE529A68C473D208" box="[452,528,1304,1330]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="4" unit="in" value="100.0">100 in</quantity>
the hemisphere) with basal portion in shape of hexagonal prism. Distal portion of processes conical with apical, sharpened, point and with almost invisible annulations. The base of the processes are joined to one another by thin striae forming a very regular reticular design with hexagonal meshes. Egg shell (excluding reticular design) smooth. Measurements of some structures of the eggs are given in
<tableCitation id="D069036FFF88FFDAFF239AC8C76CD2E8" box="[181,271,1464,1490]" captionStart="TABLE 2" captionStartId="3.[151,241,1623,1647]" captionText="TABLE 2. Measurements [in µm] of selected morphological structures of 4 eggs of Macrobiotus alvaroi sp. n. mounted in Hoyer’s medium." pageId="3" pageNumber="4">Table 2</tableCitation>
.
</paragraph>
<paragraph id="9D5436D4FF88FFDAFF519A90C5F5D118" blockId="3.[151,1436,1264,1570]" pageId="3" pageNumber="4">
<emphasis id="AF9FEAC6FF88FFDAFF519A90C70BD2C3" box="[199,360,1504,1529]" italics="true" pageId="3" pageNumber="4">Type locality:</emphasis>
<materialsCitation id="2D833C89FF88FFDAFEE79A90C5F5D118" collectingDate="2000-08" collectorName="Ziemowit Olszanowski." country="Costa Rica" location="La Selva' Biological Station" municipality="Central America" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Heredia">
<collectingMunicipality id="7D30ACAEFF88FFDAFEE79A90C45BD2C0" box="[369,568,1504,1530]" pageId="3" pageNumber="4">Central America</collectingMunicipality>
,
<collectingCountry id="E5FC7644FF88FFDAFDD29A90C4AAD2C0" box="[580,713,1504,1530]" name="Costa Rica" pageId="3" pageNumber="4">Costa Rica</collectingCountry>
,
<collectingRegion id="5F2FF836FF88FFDAFD409A90C5C5D2C0" box="[726,934,1504,1530]" country="Costa Rica" name="Heredia" pageId="3" pageNumber="4">Heredia Province</collectingRegion>
, ‘
<location id="9834600FFF88FFDAFC2E9A90C372D2C0" LSID="urn:lsid:plazi:treatment:154287C2FF8AFFDCFF019A65C7F3D0CE:9834600FFF88FFDAFC2E9A90C372D2C0" box="[952,1297,1504,1530]" country="Costa Rica" municipality="Central America" name="La Selva' Biological Station" pageId="3" pageNumber="4" stateProvince="Heredia">La Selva’ Biological Station</location>
, moss from the tropical wet forest,
<date id="E9551014FF88FFDAFE309978C45DD118" box="[422,574,1544,1570]" pageId="3" pageNumber="4" value="2000-08">
<collectingDate id="F911E9FCFF88FFDAFE309978C45DD118" box="[422,574,1544,1570]" pageId="3" pageNumber="4" value="2000-08">August 2000</collectingDate>
</date>
, leg.
<collectorName id="301E5302FF88FFDAFDEB9978C5F5D118" box="[637,918,1544,1570]" pageId="3" pageNumber="4">Ziemowit Olszanowski.</collectorName>
</materialsCitation>
</paragraph>
<caption id="C994665CFF88FFDAFF019927C70FD1AD" pageId="3" pageNumber="4" startId="3.[151,241,1623,1647]">
<paragraph id="9D5436D4FF88FFDAFF019927C70FD1AD" blockId="3.[151,1435,1623,1687]" pageId="3" pageNumber="4">
<emphasis id="AF9FEAC6FF88FFDAFF019927C76AD155" bold="true" box="[151,265,1623,1647]" pageId="3" pageNumber="4">TABLE 2.</emphasis>
Measurements [in µm] of selected morphological structures of 4 eggs of
<taxonomicName id="5AEB4D57FF88FFDAFB8A9928C295D155" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[1052,1270,1624,1647]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="4" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF88FFDAFB8A9928C295D155" box="[1052,1270,1624,1647]" italics="true" pageId="3" pageNumber="4">Macrobiotus alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF88FFDAFB6A9927C35BD155" bold="true" box="[1276,1336,1623,1647]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="B4AC57BDFF88FFDAFB6A9927C35BD155" box="[1276,1336,1623,1647]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
mounted in Hoyer’s medium.
</paragraph>
</caption>
<paragraph id="9D5436D4FF88FFDAFF0999C8C34BD0F5" pageId="3" pageNumber="4">
<table id="EFEBC474FF880026FF0999C8C34BD0F5" box="[159,1320,1720,1999]" gridcols="5" gridrows="7" pageId="3" pageNumber="4">
<tr id="23DB3496FF880026FF0999C8C34BD1F5" box="[159,1320,1720,1743]" gridrow="0" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF0999C8C40BD1F5" box="[159,616,1720,1743]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">Character</th>
<th id="600A5DEAFF880026FD0E99C8C48BD1F5" box="[664,744,1720,1743]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">1</th>
<th id="600A5DEAFF880026FCCE99C8C5CBD1F5" box="[856,936,1720,1743]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">2</th>
<th id="600A5DEAFF880026FB8E99C8C20BD1F5" box="[1048,1128,1720,1743]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">3</th>
<th id="600A5DEAFF880026FB4199C8C34BD1F5" box="[1239,1320,1720,1743]" gridcol="4" gridrow="0" pageId="3" pageNumber="4">4</th>
</tr>
<tr id="23DB3496FF880026FF099992C34BD1C3" box="[159,1320,1762,1785]" gridrow="1" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099992C40BD1C3" box="[159,616,1762,1785]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Diameter of egg without processes</th>
<td id="600A5DEAFF880026FD0E9992C48BD1C3" box="[664,744,1762,1785]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">53.3</td>
<td id="600A5DEAFF880026FCCE9992C5CBD1C3" box="[856,936,1762,1785]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">55.9</td>
<td id="600A5DEAFF880026FB8E9992C20BD1C3" box="[1048,1128,1762,1785]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">58.7</td>
<td id="600A5DEAFF880026FB419992C34BD1C3" box="[1239,1320,1762,1785]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">53.1</td>
</tr>
<tr id="23DB3496FF880026FF09987DC34BD01E" box="[159,1320,1805,1828]" gridrow="2" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF09987DC40BD01E" box="[159,616,1805,1828]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Diameter of egg with processes</th>
<td id="600A5DEAFF880026FD0E987DC48BD01E" box="[664,744,1805,1828]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">62.2</td>
<td id="600A5DEAFF880026FCCE987DC5CBD01E" box="[856,936,1805,1828]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">62.2</td>
<td id="600A5DEAFF880026FB8E987DC20BD01E" box="[1048,1128,1805,1828]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">70.0</td>
<td id="600A5DEAFF880026FB41987DC34BD01E" box="[1239,1320,1805,1828]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">63.0</td>
</tr>
<tr id="23DB3496FF880026FF099848C34BD075" box="[159,1320,1848,1871]" gridrow="3" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099848C40BD075" box="[159,616,1848,1871]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Processes height</th>
<td id="600A5DEAFF880026FD0E9848C48BD075" box="[664,744,1848,1871]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">4.6–4.8</td>
<td id="600A5DEAFF880026FCCE9848C5CBD075" box="[856,936,1848,1871]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">3.6–4.8</td>
<td id="600A5DEAFF880026FB8E9848C20BD075" box="[1048,1128,1848,1871]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">5.2–5.6</td>
<td id="600A5DEAFF880026FB419848C34BD075" box="[1239,1320,1848,1871]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">5.5–6.1</td>
</tr>
<tr id="23DB3496FF880026FF099812C34BD043" box="[159,1320,1890,1913]" gridrow="4" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF099812C40BD043" box="[159,616,1890,1913]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Distance between processes</th>
<td id="600A5DEAFF880026FD0E9812C48BD043" box="[664,744,1890,1913]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">2.6–3.3</td>
<td id="600A5DEAFF880026FCCE9812C5CBD043" box="[856,936,1890,1913]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">3.6–4.1</td>
<td id="600A5DEAFF880026FB8E9812C20BD043" box="[1048,1128,1890,1913]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">3.7–4.0</td>
<td id="600A5DEAFF880026FB419812C34BD043" box="[1239,1320,1890,1913]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">-</td>
</tr>
<tr id="23DB3496FF880026FF0998FDC34BD09E" box="[159,1320,1933,1956]" gridrow="5" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF0998FDC40BD09E" box="[159,616,1933,1956]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Number of processes on the circumference</th>
<td id="600A5DEAFF880026FD0E98FDC48BD09E" box="[664,744,1933,1956]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">25</td>
<td id="600A5DEAFF880026FCCE98FDC5CBD09E" box="[856,936,1933,1956]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">23</td>
<td id="600A5DEAFF880026FB8E98FDC20BD09E" box="[1048,1128,1933,1956]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">26</td>
<td id="600A5DEAFF880026FB4198FDC34BD09E" box="[1239,1320,1933,1956]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">25</td>
</tr>
<tr id="23DB3496FF880026FF0998C8C34BD0F5" box="[159,1320,1976,1999]" gridrow="6" pageId="3" pageNumber="4">
<th id="600A5DEAFF880026FF0998C8C40BD0F5" box="[159,616,1976,1999]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">Basal diameter of processes</th>
<td id="600A5DEAFF880026FD0E98C8C48BD0F5" box="[664,744,1976,1999]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">3.3–4.1</td>
<td id="600A5DEAFF880026FCCE98C8C5CBD0F5" box="[856,936,1976,1999]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">3.4–4.5</td>
<td id="600A5DEAFF880026FB8E98C8C20BD0F5" box="[1048,1128,1976,1999]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">3.9–4.8</td>
<td id="600A5DEAFF880026FB4198C8C34BD0F5" box="[1239,1320,1976,1999]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">3.2–4.0</td>
</tr>
</table>
</paragraph>
<caption id="C994665CFF8FFFDDFF0198CAC701D0CE" ID-DOI="http://doi.org/10.5281/zenodo.10088165" ID-Zenodo-Dep="10088165" httpUri="https://zenodo.org/record/10088165/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,255,1978,2002]" targetBox="[233,1364,184,1960]" targetPageId="4">
<paragraph id="9D5436D4FF8FFFDDFF0198CAC701D0CE" blockId="4.[151,1436,1978,2036]" pageId="4" pageNumber="5">
<emphasis id="AF9FEAC6FF8FFFDDFF0198CAC77BD0E8" bold="true" box="[151,280,1978,2002]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
Eggs, and their details, of: A–B,
<taxonomicName id="5AEB4D57FF8FFFDDFDEC98CBC499D0E8" authorityName="Pilato" authorityYear="2006" box="[634,762,1979,2002]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="AF9FEAC6FF8FFFDDFDEC98CBC499D0E8" box="[634,762,1979,2002]" italics="true" pageId="4" pageNumber="5">M. insularis</emphasis>
</taxonomicName>
. C–D,
<taxonomicName id="5AEB4D57FF8FFFDDFCD198CBC5AED0E8" authorityName="Pilato" authorityYear="2006" box="[839,973,1979,2002]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="AF9FEAC6FF8FFFDDFCD198CBC5AED0E8" box="[839,973,1979,2002]" italics="true" pageId="4" pageNumber="5">M. ocotensis</emphasis>
</taxonomicName>
. E–F,
<taxonomicName id="5AEB4D57FF8FFFDDFB8798CBC2FED0E8" box="[1041,1181,1979,2002]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="mandalae">
<emphasis id="AF9FEAC6FF8FFFDDFB8798CBC2FED0E8" box="[1041,1181,1979,2002]" italics="true" pageId="4" pageNumber="5">M. mandalae</emphasis>
</taxonomicName>
. G–H,
<taxonomicName id="5AEB4D57FF8FFFDDFB7D98CBC339D0E8" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[1259,1370,1979,2002]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="5" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8FFFDDFB7D98CBC339D0E8" box="[1259,1370,1979,2002]" italics="true" pageId="4" pageNumber="5">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8FFFDDFAF698CAC3FFD0E8" bold="true" box="[1376,1436,1978,2002]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="B4AC57BDFF8FFFDDFAF698CAC3FFD0E8" box="[1376,1436,1978,2002]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Scale bar = 10 µm.
</paragraph>
</caption>
<paragraph id="9D5436D4FF8EFFDCFF519FE8C44BD638" blockId="5.[151,1436,152,858]" pageId="5" pageNumber="6">
<emphasis id="AF9FEAC6FF8EFFDCFF519FE8C7D9D78B" box="[199,442,152,177]" italics="true" pageId="5" pageNumber="6">Additional material:</emphasis>
<materialsCitation id="2D833C89FF8EFFDCFE529FE8C44BD638" collectingDate="2003-01-10" collectorName="La Tigra &amp; Lukasz Kaczmarek." country="Costa Rica" county="Central America" location="Ciudad Quesada" municipality="La Tigra" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Alajuela">
<collectingCounty id="74354E58FF8EFFDCFE529FE8C4EFD788" box="[452,652,152,178]" pageId="5" pageNumber="6">Central America</collectingCounty>
,
<collectingCountry id="E5FC7644FF8EFFDCFD0C9FE8C543D788" box="[666,800,152,178]" name="Costa Rica" pageId="5" pageNumber="6">Costa Rica</collectingCountry>
,
<collectingRegion id="5F2FF836FF8EFFDCFCB89FE8C265D788" box="[814,1030,152,178]" country="Costa Rica" name="Alajuela" pageId="5" pageNumber="6">Alajuela Province</collectingRegion>
, moss sample was collected from stone near the road from
<collectorName id="301E5302FF8EFFDCFE289FB0C440D7E0" box="[446,547,192,218]" pageId="5" pageNumber="6">
<collectingMunicipality id="7D30ACAEFF8EFFDCFE289FB0C440D7E0" box="[446,547,192,218]" pageId="5" pageNumber="6">La Tigra</collectingMunicipality>
</collectorName>
to
<location id="9834600FFF8EFFDCFDDF9FB0C569D7E0" LSID="urn:lsid:plazi:treatment:154287C2FF8AFFDCFF019A65C7F3D0CE:9834600FFF8EFFDCFDDF9FB0C569D7E0" box="[585,778,192,218]" country="Costa Rica" county="Central America" municipality="La Tigra" name="Ciudad Quesada" pageId="5" pageNumber="6" stateProvince="Alajuela">Ciudad Quesada</location>
(San Carlos) 16.0 km before
<location id="9834600FFF8EFFDCFBF19FB0C348D7E0" LSID="urn:lsid:plazi:treatment:154287C2FF8AFFDCFF019A65C7F3D0CE:9834600FFF8EFFDCFBF19FB0C348D7E0" box="[1127,1323,192,218]" country="Costa Rica" county="Central America" municipality="La Tigra" name="Ciudad Quesada" pageId="5" pageNumber="6" stateProvince="Alajuela">Ciudad Quesada</location>
,
<date id="E9551014FF8EFFDCFAA19FB0C763D638" pageId="5" pageNumber="6" value="2003-01-10">
<collectingDate id="F911E9FCFF8EFFDCFAA19FB0C763D638" pageId="5" pageNumber="6" value="2003-01-10">10 January 2003</collectingDate>
</date>
, leg.
<collectorName id="301E5302FF8EFFDCFEA99F98C44BD638" box="[319,552,232,258]" pageId="5" pageNumber="6">Łukasz Kaczmarek.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF519E60C40CD698" blockId="5.[151,1436,152,858]" pageId="5" pageNumber="6">
<emphasis id="AF9FEAC6FF8EFFDCFF519E60C7C3D613" box="[199,416,272,297]" italics="true" pageId="5" pageNumber="6">Type depositories:</emphasis>
<materialsCitation id="2D833C89FF8EFFDCFE3F9E60C40CD698" collectorName="Binda &amp; Pilato &amp; Ecology, A." country="Poland" county="Pozna" location="Department of Animal Biology" municipality="Department of Animal Taxonomy" pageId="5" pageNumber="6" specimenCount="8" specimenCount-egg="3" typeStatus="holotype">
<typeStatus id="42508876FF8EFFDCFE3F9E60C475D610" box="[425,534,272,298]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
(slide N. 5100),
<specimenCount id="8BEDFD5DFF8EFFDCFD709E60C50DD610" box="[742,878,272,298]" pageId="5" pageNumber="6" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
and
<specimenCount id="8BEDFD5DFF8EFFDCFC3D9E60C26AD610" box="[939,1033,272,298]" pageId="5" pageNumber="6" type="egg">one egg</specimenCount>
are deposited in the collection of
<collectorName id="301E5302FF8EFFDCFF019E48C6BED668" box="[151,221,312,338]" pageId="5" pageNumber="6">Binda</collectorName>
&amp;
<collectorName id="301E5302FF8EFFDCFE909E48C729D668" box="[262,330,312,338]" pageId="5" pageNumber="6">Pilato</collectorName>
(
<location id="9834600FFF8EFFDCFECA9E48C4B7D668" LSID="urn:lsid:plazi:treatment:154287C2FF8AFFDCFF019A65C7F3D0CE:9834600FFF8EFFDCFECA9E48C4B7D668" box="[348,724,312,338]" country="Poland" county="Pozna" municipality="Department of Animal Taxonomy" name="Department of Animal Biology" pageId="5" pageNumber="6">Department of Animal Biology</location>
&quot;
<location id="9834600FFF8EFFDCFD739E48C5AED668" LSID="urn:lsid:plazi:treatment:154287C2FF8AFFDCFF019A65C7F3D0CE:9834600FFF8EFFDCFD739E48C5AED668" box="[741,973,312,338]" country="Poland" county="Pozna" municipality="Department of Animal Taxonomy" name="Marcello La Greca" pageId="5" pageNumber="6">Marcello La Greca</location>
&quot;
<location id="9834600FFF8EFFDCFC769E48C285D668" LSID="urn:lsid:plazi:treatment:154287C2FF8AFFDCFF019A65C7F3D0CE:9834600FFF8EFFDCFC769E48C285D668" box="[992,1254,312,338]" country="Poland" county="Pozna" municipality="Department of Animal Taxonomy" name="University of Catania" pageId="5" pageNumber="6">University of Catania</location>
);
<specimenCount id="8BEDFD5DFF8EFFDCFB6A9E48C6AED640" pageId="5" pageNumber="6" type="egg" typeStatus="paratypes">2 paratypes 5 eggs</specimenCount>
and also additional material are deposited at the
<collectingMunicipality id="7D30ACAEFF8EFFDCFCA69E10C2A8D640" box="[816,1227,352,378]" pageId="5" pageNumber="6">Department of Animal Taxonomy</collectingMunicipality>
and
<collectorName id="301E5302FF8EFFDCFA9B9E10C3FFD640" box="[1293,1436,352,378]" pageId="5" pageNumber="6">Ecology, A.</collectorName>
Mickiewicz University,
<collectingCounty id="74354E58FF8EFFDCFE279EF8C79BD698" box="[433,504,392,418]" pageId="5" pageNumber="6">Pozna</collectingCounty>
ṅ,
<collectingCountry id="E5FC7644FF8EFFDCFD809EF8C408D698" box="[534,619,392,418]" name="Poland" pageId="5" pageNumber="6">Poland</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF519EC0C49AD6C8" blockId="5.[151,1436,152,858]" pageId="5" pageNumber="6">
<emphasis id="AF9FEAC6FF8EFFDCFF519EC0C727D6F3" box="[199,324,432,457]" italics="true" pageId="5" pageNumber="6">Etymology</emphasis>
: The new species is dedicated to the great Costa Rican zoologist and coordinator in National Biodiversity Institute (
<collectionCode id="FBFAAE11FF8EFFDCFE089EA8C789D6C8" box="[414,490,472,498]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="5" pageNumber="6">INBio</collectionCode>
) Dr. Alvaro Herrera
<collectionCode id="FBFAAE11FF8EFFDCFD769EA8C496D6C8" box="[736,757,472,498]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="6" type="Museum">V</collectionCode>
.
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF509D70C7AED523" blockId="5.[151,1436,152,858]" box="[198,461,512,537]" pageId="5" pageNumber="6">
<emphasis id="AF9FEAC6FF8EFFDCFF509D70C7AED523" box="[198,461,512,537]" italics="true" pageId="5" pageNumber="6">Differential diagnosis.</emphasis>
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF509D58C364D583" blockId="5.[151,1436,152,858]" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFF509D58C720D57B" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[198,323,552,577]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFF509D58C720D57B" box="[198,323,552,577]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFED89D57C7F1D57B" bold="true" box="[334,402,551,577]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFED89D57C7F1D57B" box="[334,402,551,577]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="5AEB4D57FF8EFFDCFDA39D58C4AAD57B" authorityName="Kaczmarek" authorityYear="2003" box="[565,713,552,577]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="AF9FEAC6FF8EFFDCFDA39D58C4AAD57B" box="[565,713,552,577]" italics="true" pageId="5" pageNumber="6">M. polyopus</emphasis>
</taxonomicName>
in the following features: buccal tube with a longer ventral lamina; microplacoid present; claws not particularly weak and with accessory points not particularly thin; in some characters of the eggs (
<tableCitation id="D069036FFF8EFFDCFE639D08C431D5A8" box="[501,594,632,658]" captionStart="TABLE 3" captionStartId="5.[151,241,900,924]" captionText="TABLE 3. Measurements [in µm] of selected morphological structures of the eggs of species of the polyopus group." pageId="5" pageNumber="6">Table 3</tableCitation>
): processes smaller and more numerous (25 around the circumference and about
<quantity id="5A139B31FF8EFFDCFE879DD0C738D580" box="[273,347,672,698]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="6" unit="in" value="100.0">100 in</quantity>
the hemisphere in
<taxonomicName id="5AEB4D57FF8EFFDCFDAC9DD0C4D7D583" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[570,692,672,697]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFDAC9DD0C4D7D583" box="[570,692,672,697]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFD2A9DEFC49DD583" bold="true" box="[700,766,671,697]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFD2A9DEFC49DD583" box="[700,766,671,697]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and 18 and 39 respectively in
<emphasis id="AF9FEAC6FF8EFFDCFBF29DD0C364D583" box="[1124,1287,672,697]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFBF29DD0C298D583" authorityName="Kaczmarek" authorityYear="2003" box="[1124,1275,672,697]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="polyopus">M. polyopus</taxonomicName>
).
</emphasis>
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF509DB8C23ED460" blockId="5.[151,1436,152,858]" pageId="5" pageNumber="6">
The new species differs from
<taxonomicName id="5AEB4D57FF8EFFDCFDB29DB8C4AED5DB" authorityName="Pilato" authorityYear="1974" box="[548,717,712,737]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="mandalaae">
<emphasis id="AF9FEAC6FF8EFFDCFDB29DB8C4AED5DB" box="[548,717,712,737]" italics="true" pageId="5" pageNumber="6">M. mandalaae</emphasis>
</taxonomicName>
in lacking granulation on the legs IV, and in some characters of the eggs: processes more numerous (about
<quantity id="5A139B31FF8EFFDCFD3D9D80C496D430" box="[683,757,752,778]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="6" unit="in" value="100.0">100 in</quantity>
the hemisphere in
<taxonomicName id="5AEB4D57FF8EFFDCFC479D80C228D433" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[977,1099,752,777]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFC479D80C228D433" box="[977,1099,752,777]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFBC49D9FC2F7D433" bold="true" box="[1106,1172,751,777]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFBC49D9FC2F7D433" box="[1106,1172,751,777]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<quantity id="5A139B31FF8EFFDCFB5D9D80C352D430" box="[1227,1329,752,778]" metricMagnitude="0" metricUnit="m" metricValue="1.8415" metricValueMax="1.905" metricValueMin="1.778" pageId="5" pageNumber="6" unit="in" value="72.5" valueMax="75.0" valueMin="70.0">70–75 in</quantity>
<taxonomicName id="5AEB4D57FF8EFFDCFAAF9D80C684D40B" authorityName="Pilato" authorityYear="1974" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="mandalaae">
<emphasis id="AF9FEAC6FF8EFFDCFAAF9D80C684D40B" italics="true" pageId="5" pageNumber="6">M. mandalaae</emphasis>
</taxonomicName>
) and slightly more spaced, with a thinner and more sharpened distal portion; processes never bifurcated, with annulations very difficult to see (
<figureCitation id="05D02A51FF8EFFDCFD379C30C5E5D460" box="[673,902,832,858]" captionStart="FIGURE 2" captionStartId="4.[151,255,1978,2002]" captionTargetBox="[233,1364,184,1960]" captionTargetId="figure-47@4.[225,1372,178,1977]" captionTargetPageId="4" captionText="FIGURE 2. Eggs, and their details, of: A–B, M. insularis. C–D, M. ocotensis. E–F, M. mandalae. G–H, M. alvaroi sp. n. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088165" httpUri="https://zenodo.org/record/10088165/files/figure.png" pageId="5" pageNumber="6">Fig. 2G, H and E, F</figureCitation>
, see also
<tableCitation id="D069036FFF8EFFDCFC639C30C22ED460" box="[1013,1101,832,858]" captionStart="TABLE 3" captionStartId="5.[151,241,900,924]" captionText="TABLE 3. Measurements [in µm] of selected morphological structures of the eggs of species of the polyopus group." pageId="5" pageNumber="6">Table 3</tableCitation>
).
</paragraph>
<caption id="C994665CFF8EFFDCFF019CF4C31BD4A6" box="[151,1400,900,924]" pageId="5" pageNumber="6" startId="5.[151,241,900,924]">
<paragraph id="9D5436D4FF8EFFDCFF019CF4C31BD4A6" blockId="5.[151,1400,900,924]" box="[151,1400,900,924]" pageId="5" pageNumber="6">
<emphasis id="AF9FEAC6FF8EFFDCFF019CF4C76AD4A6" bold="true" box="[151,265,900,924]" pageId="5" pageNumber="6">TABLE 3.</emphasis>
Measurements [in µm] of selected morphological structures of the eggs of species of the
<taxonomicName id="5AEB4D57FF8EFFDCFB5D9CF5C348D4A6" authorityName="Kaczmarek" authorityYear="2003" box="[1227,1323,901,924]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="AF9FEAC6FF8EFFDCFB5D9CF5C348D4A6" box="[1227,1323,901,924]" italics="true" pageId="5" pageNumber="6">polyopus</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph id="9D5436D4FF8EFFDCFF099CCDC37DD27E" pageId="5" pageNumber="6">
<table id="EFEBC474FF8E0026FF099CCCC3E2D27E" box="[159,1409,956,1348]" gridcols="6" gridrows="8" pageId="5" pageNumber="6">
<tr id="23DB3496FF8E0026FF099CCCC3E2D4EE" box="[159,1409,956,980]" gridrow="0" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099CCCC7BAD4EE" box="[159,473,956,980]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">Character</th>
<th id="600A5DEAFF8E0026FE649CCCC415D4EE" box="[498,630,956,980]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFE649CCDC415D4EE" authorityName="Kaczmarek" authorityYear="2003" box="[498,630,957,980]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="AF9FEAC6FF8EFFDCFE649CCDC415D4EE" box="[498,630,957,980]" italics="true" pageId="5" pageNumber="6">M. polyopus</emphasis>
</taxonomicName>
</th>
<th id="600A5DEAFF8E0026FD019CCCC553D4EE" box="[663,816,956,980]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFD019CCDC553D4EE" authorityName="Pilato" authorityYear="1974" box="[663,816,957,980]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="mandalaae">
<emphasis id="AF9FEAC6FF8EFFDCFD019CCDC553D4EE" box="[663,816,957,980]" italics="true" pageId="5" pageNumber="6">M. mandalaae</emphasis>
</taxonomicName>
</th>
<th id="600A5DEAFF8E0026FCC19CCCC5B9D4EE" box="[855,986,956,980]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFCCA9CCDC5B9D4EE" authorityName="Pilato" authorityYear="2006" box="[860,986,957,980]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="AF9FEAC6FF8EFFDCFCCA9CCDC5B9D4EE" box="[860,986,957,980]" italics="true" pageId="5" pageNumber="6">M. insularis</emphasis>
</taxonomicName>
</th>
<th id="600A5DEAFF8E0026FB999CCCC2F6D4EE" box="[1039,1173,956,980]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFB999CCDC2F6D4EE" authorityName="Pilato" authorityYear="2006" box="[1039,1173,957,980]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="AF9FEAC6FF8EFFDCFB999CCDC2F6D4EE" box="[1039,1173,957,980]" italics="true" pageId="5" pageNumber="6">M. ocotensis</emphasis>
</taxonomicName>
</th>
<th id="600A5DEAFF8E0026FB599CCCC3E2D4EE" box="[1231,1409,956,980]" gridcol="5" gridrow="0" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFB599CCDC35DD4EE" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[1231,1342,957,980]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFB599CCDC35DD4EE" box="[1231,1342,957,980]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFAD39CCCC3E2D4EE" bold="true" box="[1349,1409,956,980]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFAD39CCCC3E2D4EE" box="[1349,1409,956,980]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="23DB3496FF8E0026FF099C98C3E2D4C5" box="[159,1409,1000,1023]" gridrow="1" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099C98C7BAD4C5" box="[159,473,1000,1023]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">Diameter without processes</th>
<td id="600A5DEAFF8E0026FE649C98C415D4C5" box="[498,630,1000,1023]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">-</td>
<td id="600A5DEAFF8E0026FD019C98C553D4C5" box="[663,816,1000,1023]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">45.0</td>
<td id="600A5DEAFF8E0026FCC19C98C5B9D4C5" box="[855,986,1000,1023]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">78.7</td>
<td id="600A5DEAFF8E0026FB999C98C2F6D4C5" box="[1039,1173,1000,1023]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">63.0</td>
<td id="600A5DEAFF8E0026FB599C98C3E2D4C5" box="[1231,1409,1000,1023]" gridcol="5" gridrow="1" pageId="5" pageNumber="6">53.3–58.7</td>
</tr>
<tr id="23DB3496FF8E0026FF099B62C3E2D313" box="[159,1409,1042,1065]" gridrow="2" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099B62C7BAD313" box="[159,473,1042,1065]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">Diameter with processes</th>
<td id="600A5DEAFF8E0026FE649B62C415D313" box="[498,630,1042,1065]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">65.0</td>
<td id="600A5DEAFF8E0026FD019B62C553D313" box="[663,816,1042,1065]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">55.0</td>
<td id="600A5DEAFF8E0026FCC19B62C5B9D313" box="[855,986,1042,1065]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">94.0</td>
<td id="600A5DEAFF8E0026FB999B62C2F6D313" box="[1039,1173,1042,1065]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">79.9</td>
<td id="600A5DEAFF8E0026FB599B62C3E2D313" box="[1231,1409,1042,1065]" gridcol="5" gridrow="2" pageId="5" pageNumber="6">62.2 – 70.0</td>
</tr>
<tr id="23DB3496FF8E0026FF099B4DC3E2D34D" box="[159,1409,1085,1143]" gridrow="3" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099B4DC7BAD34D" box="[159,473,1085,1143]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">Number of processes on the circumference</th>
<td id="600A5DEAFF8E0026FE649B4DC415D34D" box="[498,630,1085,1143]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">18*</td>
<td id="600A5DEAFF8E0026FD019B4DC553D34D" box="[663,816,1085,1143]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">24–26</td>
<td id="600A5DEAFF8E0026FCC19B4DC5B9D34D" box="[855,986,1085,1143]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">13–14</td>
<td id="600A5DEAFF8E0026FB999B4DC2F6D34D" box="[1039,1173,1085,1143]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">25</td>
<td id="600A5DEAFF8E0026FB599B4DC3E2D34D" box="[1231,1409,1085,1143]" gridcol="5" gridrow="3" pageId="5" pageNumber="6">23–26</td>
</tr>
<tr id="23DB3496FF8E0026FF099BFAC3E2D3FE" box="[159,1409,1162,1220]" gridrow="4" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099BFAC7BAD3FE" box="[159,473,1162,1220]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Number of processes on the hemisphere</th>
<td id="600A5DEAFF8E0026FE649BFAC415D3FE" box="[498,630,1162,1220]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">39*</td>
<td id="600A5DEAFF8E0026FD019BFAC553D3FE" box="[663,816,1162,1220]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">70–75</td>
<td id="600A5DEAFF8E0026FCC19BFAC5B9D3FE" box="[855,986,1162,1220]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">35</td>
<td id="600A5DEAFF8E0026FB999BFAC2F6D3FE" box="[1039,1173,1162,1220]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">about 100</td>
<td id="600A5DEAFF8E0026FB599BFAC3E2D3FE" box="[1231,1409,1162,1220]" gridcol="5" gridrow="4" pageId="5" pageNumber="6">About 100</td>
</tr>
<tr id="23DB3496FF8E0026FF099BA8C3E2D3D5" box="[159,1409,1240,1263]" gridrow="5" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099BA8C7BAD3D5" box="[159,473,1240,1263]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">Processes height</th>
<td id="600A5DEAFF8E0026FE649BA8C415D3D5" box="[498,630,1240,1263]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">-</td>
<td id="600A5DEAFF8E0026FD019BA8C553D3D5" box="[663,816,1240,1263]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">3.4–4.7</td>
<td id="600A5DEAFF8E0026FCC19BA8C5B9D3D5" box="[855,986,1240,1263]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">7.3–8.8</td>
<td id="600A5DEAFF8E0026FB999BA8C2F6D3D5" box="[1039,1173,1240,1263]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">4.7–6.0</td>
<td id="600A5DEAFF8E0026FB599BA8C3E2D3D5" box="[1231,1409,1240,1263]" gridcol="5" gridrow="5" pageId="5" pageNumber="6">3.6–6.1</td>
</tr>
<tr id="23DB3496FF8E0026FF099A72C3E2D223" box="[159,1409,1282,1305]" gridrow="6" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099A72C7BAD223" box="[159,473,1282,1305]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">Basal diameter of the process</th>
<td id="600A5DEAFF8E0026FE649A72C415D223" box="[498,630,1282,1305]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">-</td>
<td id="600A5DEAFF8E0026FD019A72C553D223" box="[663,816,1282,1305]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">3.2–4.5</td>
<td id="600A5DEAFF8E0026FCC19A72C5B9D223" box="[855,986,1282,1305]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">10.9–13.2</td>
<td id="600A5DEAFF8E0026FB999A72C2F6D223" box="[1039,1173,1282,1305]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">4.9–6.0</td>
<td id="600A5DEAFF8E0026FB599A72C3E2D223" box="[1231,1409,1282,1305]" gridcol="5" gridrow="6" pageId="5" pageNumber="6">3.3–4.1</td>
</tr>
<tr id="23DB3496FF8E0026FF099A5DC3E2D27E" box="[159,1409,1325,1348]" gridrow="7" pageId="5" pageNumber="6">
<th id="600A5DEAFF8E0026FF099A5DC7BAD27E" box="[159,473,1325,1348]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">Meshes of the egg shell</th>
<td id="600A5DEAFF8E0026FE649A5DC415D27E" box="[498,630,1325,1348]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">-</td>
<td id="600A5DEAFF8E0026FD019A5DC553D27E" box="[663,816,1325,1348]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">smooth</td>
<td id="600A5DEAFF8E0026FCC19A5DC5B9D27E" box="[855,986,1325,1348]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">dotted</td>
<td id="600A5DEAFF8E0026FB999A5DC2F6D27E" box="[1039,1173,1325,1348]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">smooth</td>
<td id="600A5DEAFF8E0026FB599A5DC3E2D27E" box="[1231,1409,1325,1348]" gridcol="5" gridrow="7" pageId="5" pageNumber="6">smooth</td>
</tr>
</table>
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF019A1DC446D2BE" blockId="5.[151,549,1389,1412]" box="[151,549,1389,1412]" pageId="5" pageNumber="6">
*from the drawing of
<bibRefCitation id="F97A4B25FF8EFFDCFE179A1DC442D2BE" author="Marcus, E." box="[385,545,1389,1412]" pageId="5" pageNumber="6" pagination="1 - 230" refId="ref3245" refString="Marcus, E. (1928) Bartierchen (Tardigrada). In: Dahl F (ed.): Die Tierwelt Deutschlands und der angrenzenden Meeresteile, Gustav Fischer, Jena, 12, IV, 1 - 230." type="journal article" year="1928">Marcus (1928)</bibRefCitation>
.
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF509ADBC701D1E6" blockId="5.[151,1437,1450,2036]" pageId="5" pageNumber="6">
<taxonomicName id="5AEB4D57FF8EFFDCFF509ADBC723D2FE" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[198,320,1451,1476]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFF509ADBC723D2FE" box="[198,320,1451,1476]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFEDE9ADAC7E1D2FE" bold="true" box="[328,386,1450,1476]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFEDE9ADAC7E1D2FE" box="[328,386,1450,1476]" pageId="5" pageNumber="6" rank="species">sp. n</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="5AEB4D57FF8EFFDCFD8B9ADBC4C9D2FE" authorityName="Pilato" authorityYear="2006" box="[541,682,1451,1476]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="AF9FEAC6FF8EFFDCFD8B9ADBC4C9D2FE" box="[541,682,1451,1476]" italics="true" pageId="5" pageNumber="6">M. insularis</emphasis>
</taxonomicName>
in having shorter and narrower buccal tube. Due to the different length of the buccal tube, some structures,
<emphasis id="AF9FEAC6FF8EFFDCFD349AA3C4A8D2D6" box="[674,715,1491,1516]" italics="true" pageId="5" pageNumber="6">e.g.</emphasis>
, the placoids and the claws, have different values of the
<emphasis id="AF9FEAC6FF8EFFDCFA139AA3C3FFD2D6" box="[1413,1436,1491,1516]" italics="true" pageId="5" pageNumber="6">pt</emphasis>
index (see
<tableCitation id="D069036FFF8EFFDCFE8E9A8AC717D12E" box="[280,372,1530,1556]" captionStart="TABLE 4" captionStartId="6.[151,241,151,175]" captionText="TABLE 4. Measurements [in µm] and pt values of selected morphological structures of specimens of M. mandalaae, M. insularis, M. ocotensis and M. alvaroi sp. n. with similar body length." pageId="5" pageNumber="6">Table 4</tableCitation>
). It also differs in some characters of the eggs (
<figureCitation id="05D02A51FF8EFFDCFC3D9A8AC2C3D12E" box="[939,1184,1530,1556]" captionStart="FIGURE 2" captionStartId="4.[151,255,1978,2002]" captionTargetBox="[233,1364,184,1960]" captionTargetId="figure-47@4.[225,1372,178,1977]" captionTargetPageId="4" captionText="FIGURE 2. Eggs, and their details, of: A–B, M. insularis. C–D, M. ocotensis. E–F, M. mandalae. G–H, M. alvaroi sp. n. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088165" httpUri="https://zenodo.org/record/10088165/files/figure.png" pageId="5" pageNumber="6">Fig. 2G, H and A, B</figureCitation>
): smaller dimensions (diameter without processes 53.3–58.7 and 62.2–70.0 including them in
<taxonomicName id="5AEB4D57FF8EFFDCFC639953C20AD106" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[1013,1129,1571,1596]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFC639953C20AD106" box="[1013,1129,1571,1596]" italics="true" pageId="5" pageNumber="6">M.alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFBE49952C2D6D106" bold="true" box="[1138,1205,1570,1596]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFBE49952C2D6D106" box="[1138,1205,1570,1596]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, about 80 and 94.0 respectively in
<taxonomicName id="5AEB4D57FF8EFFDCFEDF993BC7B5D15E" authorityName="Pilato" authorityYear="2006" box="[329,470,1611,1636]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="AF9FEAC6FF8EFFDCFEDF993BC7B5D15E" box="[329,470,1611,1636]" italics="true" pageId="5" pageNumber="6">M. insularis</emphasis>
</taxonomicName>
), smaller processes (3.6–6.1 high in
<taxonomicName id="5AEB4D57FF8EFFDCFC13993BC263D15E" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[901,1024,1610,1636]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFC13993BC5FED15E" box="[901,925,1611,1636]" italics="true" pageId="5" pageNumber="6">M</emphasis>
.
<emphasis id="AF9FEAC6FF8EFFDCFC3A993BC263D15E" box="[940,1024,1611,1636]" italics="true" pageId="5" pageNumber="6">alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFB90993AC22BD15E" bold="true" box="[1030,1096,1610,1636]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFB90993AC22BD15E" box="[1030,1096,1610,1636]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<quantity id="5A139B31FF8EFFDCFB16993AC295D15E" box="[1152,1270,1610,1636]" metricMagnitude="-1" metricUnit="m" metricValue="2.0447" metricValueMax="2.2352" metricValueMin="1.8541999999999998" pageId="5" pageNumber="6" unit="in" value="8.05" valueMax="8.8" valueMin="7.3">7.3–8.8 in</quantity>
<taxonomicName id="5AEB4D57FF8EFFDCFB6B993BC3E9D15E" authorityName="Pilato" authorityYear="2006" box="[1277,1418,1611,1636]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="AF9FEAC6FF8EFFDCFB6B993BC3E9D15E" box="[1277,1418,1611,1636]" italics="true" pageId="5" pageNumber="6">M. insularis</emphasis>
</taxonomicName>
), more numerous processes (
<quantity id="5A139B31FF8EFFDCFE4F9902C420D1B6" box="[473,579,1650,1676]" metricMagnitude="-1" metricUnit="m" metricValue="6.223000000000001" metricValueMax="6.604000000000001" metricValueMin="5.8420000000000005" pageId="5" pageNumber="6" unit="in" value="24.5" valueMax="26.0" valueMin="23.0">23–26 in</quantity>
the circumference and about
<quantity id="5A139B31FF8EFFDCFC329902C58DD1B6" box="[932,1006,1650,1676]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="6" unit="in" value="100.0">100 in</quantity>
the hemisphere of
<taxonomicName id="5AEB4D57FF8EFFDCFB429903C333D1B6" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[1236,1360,1651,1676]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFB429903C333D1B6" box="[1236,1360,1651,1676]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFACF9902C3FFD1B6" bold="true" box="[1369,1436,1650,1676]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFACF9902C3FFD1B6" box="[1369,1436,1650,1676]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and 13–14 and 35 respectively in
<taxonomicName id="5AEB4D57FF8EFFDCFD8899EBC4C8D18E" authorityName="Pilato" authorityYear="2006" box="[542,683,1691,1716]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="AF9FEAC6FF8EFFDCFD8899EBC4C8D18E" box="[542,683,1691,1716]" italics="true" pageId="5" pageNumber="6">M. insularis</emphasis>
</taxonomicName>
), processes different in the shape, egg shell with reticular design and without dots.
</paragraph>
<paragraph id="9D5436D4FF8EFFDCFF50999AC7F3D0CE" blockId="5.[151,1437,1450,2036]" pageId="5" pageNumber="6">
Besides, the new species differs from
<taxonomicName id="5AEB4D57FF8EFFDCFD12999BC57AD03E" authorityName="Pilato" authorityYear="2006" box="[644,793,1771,1796]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="AF9FEAC6FF8EFFDCFD12999BC57AD03E" box="[644,793,1771,1796]" italics="true" pageId="5" pageNumber="6">M. ocotensis</emphasis>
</taxonomicName>
in lacking granulation on the legs and in some characters of the eggs (
<tableCitation id="D069036FFF8EFFDCFEC09862C7D3D016" box="[342,432,1810,1836]" captionStart="TABLE 3" captionStartId="5.[151,241,900,924]" captionText="TABLE 3. Measurements [in µm] of selected morphological structures of the eggs of species of the polyopus group." pageId="5" pageNumber="6">Table 3</tableCitation>
;
<figureCitation id="05D02A51FF8EFFDCFE2B9862C4C2D016" box="[445,673,1810,1836]" captionStart="FIGURE 2" captionStartId="4.[151,255,1978,2002]" captionTargetBox="[233,1364,184,1960]" captionTargetId="figure-47@4.[225,1372,178,1977]" captionTargetPageId="4" captionText="FIGURE 2. Eggs, and their details, of: A–B, M. insularis. C–D, M. ocotensis. E–F, M. mandalae. G–H, M. alvaroi sp. n. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10088165" httpUri="https://zenodo.org/record/10088165/files/figure.png" pageId="5" pageNumber="6">Fig 2G, H and C, D</figureCitation>
): slightly smaller dimensions (diameter 53.3–58.7 excluding processes and 62.2–70.0 including them in
<taxonomicName id="5AEB4D57FF8EFFDCFD1F984BC56AD06E" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[649,777,1851,1876]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFD1F984BC56AD06E" box="[649,777,1851,1876]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFC83984AC53FD06E" bold="true" box="[789,860,1850,1876]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFC83984AC53FD06E" box="[789,860,1850,1876]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, about 63.0 and about 80.0 respectively in
<taxonomicName id="5AEB4D57FF8EFFDCFAEA984BC767D046" authorityName="Pilato" authorityYear="2006" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="AF9FEAC6FF8EFFDCFAEA984BC767D046" italics="true" pageId="5" pageNumber="6">M. ocotensis</emphasis>
</taxonomicName>
); processes different in the shape (basal portion in shape of a hexagonal prism and conical distal portion with an apical, sharpened point with faint annulations in
<taxonomicName id="5AEB4D57FF8EFFDCFCFE98FBC581D09E" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[872,994,1931,1956]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="alvaroi" status="sp. nov.">
<emphasis id="AF9FEAC6FF8EFFDCFCFE98FBC581D09E" box="[872,994,1931,1956]" italics="true" pageId="5" pageNumber="6">M. alvaroi</emphasis>
</taxonomicName>
<emphasis id="AF9FEAC6FF8EFFDCFC7C98FAC250D09E" bold="true" box="[1002,1075,1930,1956]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="B4AC57BDFF8EFFDCFC7C98FAC24ED09E" box="[1002,1069,1930,1956]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
and in
<taxonomicName id="5AEB4D57FF8EFFDCFB1D98FBC37CD09E" authorityName="Pilato" authorityYear="2006" box="[1163,1311,1931,1956]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="AF9FEAC6FF8EFFDCFB1D98FBC37CD09E" box="[1163,1311,1931,1956]" italics="true" pageId="5" pageNumber="6">M. ocotensis</emphasis>
</taxonomicName>
basal portion in the shape of a hexagonal prism and a breast-shaped distal portion with an apical, sharpened point without annulations).
</paragraph>
</subSubSection>
</treatment>
</document>
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