150 lines
19 KiB
XML
150 lines
19 KiB
XML
<document id="C4329CA7839C4941F8D7D18D09D6D908" ID-DOI="10.5281/zenodo.178667" ID-GBIF-Dataset="3911b39d-310e-4ad3-aa72-c01b81c0bd80" ID-ISSN="1175-5326" ID-Zenodo-Dep="178667" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459904496754" checkinUser="plazi" docAuthor="Altaba, Cristian R." docDate="2007" docId="003487CEFFCCFFACFF3B71B5FA75FE12" docLanguage="en" docName="zt01595p052.pdf" docOrigin="Zootaxa 1595" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Balearena gymnesica Altaba, 2007, sp. nov." docType="treatment" docVersion="10" lastPageNumber="46" masterDocId="FC0DFFB6FFCEFFAFFFAC735FFFE9FFB0" masterDocTitle="A new genus and species of Enidae (Gastopoda: Pulmonata) from the Quaternary of the Balearic Islands (Western Mediterranean)" masterLastPageNumber="52" masterPageNumber="43" pageNumber="45" updateTime="1698228927245" updateUser="plazi">
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<mods:mods id="62F179369E4A21238D34BC81EEF927A9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0C4170355EE7653C275C6F163DE6DFF9">
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<mods:title id="2EE5094F4B6F3D87BFA9E524C40CC761">A new genus and species of Enidae (Gastopoda: Pulmonata) from the Quaternary of the Balearic Islands (Western Mediterranean)</mods:title>
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</mods:titleInfo>
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<mods:name id="DCB0CC47DACAAA1F0399B5A9683F0731" type="personal">
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<mods:roleTerm id="EC4068C1B8B039BA504958123AE58D88">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AB7CEEAD7CA91003A9CA43F33E3AA557">Altaba, Cristian R.</mods:namePart>
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<mods:typeOfResource id="36730AACFBED03810B4AC5950F5F4356">text</mods:typeOfResource>
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<mods:title id="F9427CC041CD6A62B1AAD7E73DFD3803">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6676AB3487E71588E9F779FD13894EF8">
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<mods:date id="448894584960FF32BB0E84E84D0CAA33">2007</mods:date>
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<mods:detail id="57423DDBFFAA94D782A6701153246530" type="volume">
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<mods:number id="14922444FF78E7B2EA9251B9506004C9">1595</mods:number>
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<mods:start id="8AEB687CA05069605B144B94A4327955">43</mods:start>
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<mods:end id="EB0EDD57C32C3A94B9744E5EFD8E8E16">52</mods:end>
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<mods:classification id="57B12A2A39A8C41E92B205AF1A308634">journal article</mods:classification>
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<mods:identifier id="E015CA41A25DC0C87DDED497D16504DB" type="DOI">10.5281/zenodo.178667</mods:identifier>
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<mods:identifier id="DC8BEE3A52DBDC4E33D858200D794F21" type="GBIF-Dataset">3911b39d-310e-4ad3-aa72-c01b81c0bd80</mods:identifier>
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<mods:identifier id="9A31878E51E48F2B83CEF71535BF80E5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="28E624AA0066DD4C48DAE7CAA545907B" type="Zenodo-Dep">178667</mods:identifier>
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<treatment id="003487CEFFCCFFACFF3B71B5FA75FE12" ID-DOI="http://doi.org/10.5281/zenodo.5677295" ID-GBIF-Taxon="119373305" ID-Zenodo-Dep="5677295" LSID="urn:lsid:plazi:treatment:003487CEFFCCFFACFF3B71B5FA75FE12" httpUri="http://treatment.plazi.org/id/003487CEFFCCFFACFF3B71B5FA75FE12" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45">
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<subSubSection id="C0876553FFCCFFADFF3B71B5FEDFFC97" pageId="2" pageNumber="45" type="nomenclature">
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<paragraph id="882236D8FFCCFFADFF3B71B5FE16FCB4" blockId="2.[151,511,746,807]" box="[151,511,746,772]" pageId="2" pageNumber="45">
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<heading id="D36A81B4FFCCFFADFF3B71B5FE16FCB4" bold="true" box="[151,511,746,772]" fontSize="11" level="1" pageId="2" pageNumber="45" reason="1">
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<emphasis id="BAE9EACAFFCCFFADFF3B71B5FE16FCB4" bold="true" box="[151,511,746,772]" pageId="2" pageNumber="45">
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<taxonomicName id="4F9D4D5BFFCCFFADFF3B71B5FE72FCB4" box="[151,411,746,772]" class="Gastropoda" family="Enidae" genus="Balearena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="45" phylum="Mollusca" rank="species" species="gymnesica" status="sp. nov.">
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<emphasis id="BAE9EACAFFCCFFADFF3B71B5FE72FCB4" bold="true" box="[151,411,746,772]" italics="true" pageId="2" pageNumber="45">Balearena gymnesica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A1DA57B1FFCCFFADFE0E71B5FE16FCB4" box="[418,511,746,772]" pageId="2" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="882236D8FFCCFFADFF3B7052FEDFFC97" blockId="2.[151,511,746,807]" box="[151,310,781,807]" pageId="2" pageNumber="45">
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(
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<figureCitation id="10A62A5DFFCCFFADFF337052FEC5FC97" box="[159,300,781,807]" captionStart-0="FIGURE 1 – 2. 1" captionStart-1="FIGURE 3" captionStartId-0="3.[151,255,1458,1482]" captionStartId-1="4.[151,254,1263,1287]" captionTargetBox-0="[151,1411,432,1457]" captionTargetBox-1="[184,1404,188,1220]" captionTargetId-0="figure@3.[151,1436,432,1460]" captionTargetId-1="figure@4.[174,1428,167,1245]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1 – 2. 1. Balearena gymnesica gen. nov., spec. nov. Holotype, upper coastal cliff at eastern edge of Es Carnatge, Palma de Mallorca, Mallorca island, Balearic Islands, Western Mediterranean. Shell length 16.7 mm; scale bar 2.0 mm. Not shown are a thin encrustation over most of the first whorl and minimal fractures on the peristome. 2. Protoconch sculpture of Balearena gymnesica. This reconstruction is based on partial remains of these delicate features that are observable in various specimens. Scale bar 0.5 mm." captionText-1="FIGURE 3. Comparison of shells of Mastus pupa (top row; Korbous, Tunisia) and Balearena gymnesica (bottom row; Es Carnatge, Mallorca). Scale bar 10 mm." httpUri-0="https://zenodo.org/record/178668/files/figure.png" httpUri-1="https://zenodo.org/record/178669/files/figure.png" pageId="2" pageNumber="45">Figures 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C0876553FFCCFFADFF3B7002FC00FB3F" pageId="2" pageNumber="45" type="materials_examined">
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<paragraph id="882236D8FFCCFFADFF3B7002FF01FC77" blockId="2.[151,1437,861,2025]" pageId="2" pageNumber="45">
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<emphasis id="BAE9EACAFFCCFFADFF3B7002FEB8FCC7" bold="true" box="[151,337,861,887]" pageId="2" pageNumber="45">
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<typeStatus id="5726887AFFCCFFADFF3B7002FF3BFCC7" box="[151,210,861,887]" pageId="2" pageNumber="45">Type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="5726887AFFCCFFADFEF77002FE22FCC7" box="[347,459,861,887]" pageId="2" pageNumber="45" type="holotype">Holotype</typeStatus>
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,
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<collectionCode id="EE8CAE1DFFCCFFADFE747002FDF1FCC7" box="[472,536,861,887]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="45">MZB</collectionCode>
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2005-1022;
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<typeStatus id="5726887AFFCCFFADFD1E7002FCCBFCC7" box="[690,802,861,887]" pageId="2" pageNumber="45" type="paratype">paratypes</typeStatus>
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(three complete, four incomplete),
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<collectionCode id="EE8CAE1DFFCCFFADFB617002FAE4FCC7" box="[1229,1293,861,887]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="45">MZB</collectionCode>
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2005-1023. Additional
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<typeStatus id="5726887AFFCCFFADFEB070DAFE65FC2F" box="[284,396,901,927]" pageId="2" pageNumber="45" type="paratype">paratypes</typeStatus>
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(five each: one complete and four incomplete) deposited in
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<collectionCode id="EE8CAE1DFFCCFFADFBFE70DAFB45FC2F" box="[1106,1196,901,927]" pageId="2" pageNumber="45">MBCN</collectionCode>
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, CRA-10862, CRA- 10907.
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</paragraph>
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<paragraph id="882236D8FFCCFFADFF6A708AFC00FB3F" blockId="2.[151,1437,861,2025]" pageId="2" pageNumber="45">
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<emphasis id="BAE9EACAFFCCFFADFF6A708AFE86FC5F" bold="true" box="[198,367,981,1007]" pageId="2" pageNumber="45">
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<typeStatus id="5726887AFFCCFFADFF6A708AFEEAFC5F" box="[198,259,981,1007]" pageId="2" pageNumber="45">Type</typeStatus>
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locality:
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</emphasis>
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Top level of the low (
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<quantity id="4F659B3DFFCCFFADFDDC708AFD77FC5F" box="[624,670,981,1007]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="45" unit="m" value="2.0">2 m</quantity>
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) coastal cliff at Cala Pudent, in Camp de Tir, the eastern sector of Es Carnatge, in Palma de Mallorca, Mallorca Island, Balearic Islands (
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<geoCoordinate id="EDA9501FFFCCFFADFC7470A2FBA7FBA7" box="[984,1102,1021,1047]" direction="north" orientation="latitude" pageId="2" pageNumber="45" precision="925" value="39.533333">39º 32’ N</geoCoordinate>
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,
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<geoCoordinate id="EDA9501FFFCCFFADFBF770A2FB56FBA7" box="[1115,1215,1021,1047]" direction="east" orientation="longitude" pageId="2" pageNumber="45" precision="925" value="2.7">2º 42’ E</geoCoordinate>
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; UTM coordinates 31S DD 7477). This terrace, overlying various Upper Pleistocene strata, is formed by terrigenous clay mixed with marine biogenetic sand, marking at its top the latest marine regression at the end of the Neotyrrhenan, the last interglacial transgressive sequence recorded in Mediterranean strata.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0876553FFCCFFADFF6A77C2FC95FACF" pageId="2" pageNumber="45" type="distribution">
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<paragraph id="882236D8FFCCFFADFF6A77C2FC95FACF" blockId="2.[151,1437,861,2025]" pageId="2" pageNumber="45">
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<emphasis id="BAE9EACAFFCCFFADFF6A77C2FE80FB07" bold="true" box="[198,361,1181,1207]" pageId="2" pageNumber="45">Distribution:</emphasis>
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Known only from the northern Balearic Islands, or Gymnesics. The
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<typeStatus id="5726887AFFCCFFADFB6777C2FB17FB07" box="[1227,1278,1181,1207]" pageId="2" pageNumber="45">type</typeStatus>
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stratum was formed at the onset of the Würm II glacial cycle, with an age of ca. 80,000 years (
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<bibRefCitation id="EC0C4B29FFCCFFADFBFA779AFB1BFB6F" author="Cuerda" box="[1110,1266,1221,1247]" pageId="2" pageNumber="51" refString="Cuerda, J. (1989) Los tiempos cuaternarios en Baleares. 2 nd ed. Conselleria de Cultura, Educacio i Esports, Govern Balear, Palma de Mallorca, 310 pp. + 20 pls." type="book" year="1989">Cuerda 1989</bibRefCitation>
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;
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<bibRefCitation id="EC0C4B29FFCCFFADFB52779AFEAAFAB7" author="Hillaire-Marcel" pageId="2" pageNumber="52" refString="Hillaire-Marcel, C., Gariepy, C., Ghaleb, B., Goy, J. - L., Zazo, C. & Cuerda Barcelo, J. (1996) U-series measurements in Tyrrhenian deposits from Mallorca - Further evidence for two last-interglacial high sea levels in the Balearic Islands. Quaternary Science Reviews, 15, 53 - 62." type="journal article" year="1996">
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Hillaire-Marcel
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<emphasis id="BAE9EACAFFCCFFADFF6D77B2FF14FAB6" box="[193,253,1261,1286]" italics="true" pageId="2" pageNumber="45">et al.</emphasis>
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1996
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</bibRefCitation>
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;
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<bibRefCitation id="EC0C4B29FFCCFFADFEFC77B2FD3AFAB7" author="Gonzalez-Hernandez" box="[336,723,1261,1287]" pageId="2" pageNumber="51" refString="Gonzalez-Hernandez, F. M., Goy, J. L., Zazo, C. & Silva, P. G. (2001) Actividad eolica - cambios en el nivel del mar durante los ultimos 170.000 anos (litoral de Mallorca, Islas Baleares). Revista de Cuaternario y Geologia, 15, 67 - 75." type="journal article" year="2001">
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González-Hernández
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<emphasis id="BAE9EACAFFCCFFADFDFC77B2FD62FAB6" box="[592,651,1261,1286]" italics="true" pageId="2" pageNumber="45">et al.</emphasis>
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2001
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</bibRefCitation>
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). Moderately common in late Pleistocene coastal paleosoils, fossil beaches and eolianites in Mallorca,
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<collectingCountry id="F08A7648FFCCFFADFD2B764AFD06FA9F" box="[647,751,1301,1327]" name="Spain" pageId="2" pageNumber="45">Menorca</collectingCountry>
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and Cabrera (
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<bibRefCitation id="EC0C4B29FFCCFFADFC3B764AFBDAFA9F" author="Cuerda" box="[919,1075,1301,1327]" pageId="2" pageNumber="51" refString="Cuerda, J. (1989) Los tiempos cuaternarios en Baleares. 2 nd ed. Conselleria de Cultura, Educacio i Esports, Govern Balear, Palma de Mallorca, 310 pp. + 20 pls." type="book" year="1989">Cuerda 1989</bibRefCitation>
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,
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<bibRefCitation id="EC0C4B29FFCCFFADFBEC764AFB97FA9F" author="Cuerda" box="[1088,1150,1301,1327]" pageId="2" pageNumber="51" refString="Cuerda, J. (1993) Nota sobre el Quaternari. In: Alcover, J. A., Fornos, J. & Ballesteros, E. (Eds.), Historia natural de l'Arxipelag de Cabrera. Ed. Moll & C. S. I. C, Palma de Mallorca, 117 - 130." type="book chapter" year="1993">1993</bibRefCitation>
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;
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<bibRefCitation id="EC0C4B29FFCCFFADFB27764AFAC9FA9F" author="Altaba" box="[1163,1312,1301,1327]" pageId="2" pageNumber="50" refString="Altaba, C. R. (1993) Els caragols i llimacs terrestres (Mollusca: Gastropoda). In: Alcover, J. A., Fornos, J. & Ballesteros, E. (Eds.), Historia natural de l'Arxipelag de Cabrera. Ed. Moll & C. S. I. C, Palma de Mallorca, 409 - 426." type="book chapter" year="1993">Altaba 1993</bibRefCitation>
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). The species is also reported from older cave fillings in coastal cliffs in
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<collectingCountry id="F08A7648FFCCFFADFC227662FC10FAE7" box="[910,1017,1341,1367]" name="Spain" pageId="2" pageNumber="45">Menorca</collectingCountry>
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, putatively dated as latest Miocene (
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<bibRefCitation id="EC0C4B29FFCCFFADFF33763AFEA6FACF" author="Quintana" box="[159,335,1381,1407]" pageId="2" pageNumber="52" refString="Quintana, J. (1995) Fauna malacologica asociada a Cheirogaster gymnesica (Bate, 1914). Implicaciones biogeograficas. Bolleti de la Societat d'Historia Natural de les Balears, 38, 95 - 119." type="journal article" year="1995">Quintana 1995</bibRefCitation>
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) but likely being Plio-Pleistocene cave fillings.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0876553FFCCFFACFF6A76D2FF25FEB2" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45" type="description">
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<paragraph id="882236D8FFCCFFACFF6A76D2FF25FEB2" blockId="2.[151,1437,861,2025]" lastBlockId="3.[151,1436,152,418]" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45">
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<emphasis id="BAE9EACAFFCCFFADFF6A76D2FE88FA17" bold="true" box="[198,353,1421,1447]" pageId="2" pageNumber="45">Description:</emphasis>
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Based on
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<specimenCount id="9E9BFD51FFCCFFADFE4F76D2FD6EFA17" box="[483,647,1421,1447]" pageId="2" pageNumber="45" type="generic">17 specimens</specimenCount>
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, plus numerous fragments, all from the
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<typeStatus id="5726887AFFCCFFADFBC776D2FB74FA17" box="[1131,1181,1421,1447]" pageId="2" pageNumber="45">type</typeStatus>
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locality and stratum. Shell typically enid, with spire conspicuously slender and aperture gently expanded basally, rather thin, fragile. Spire acute elongated, whorls 7 1/8–7 1/4, spire angle approximately 30o. Profile of spire slightly concave from whorls 2 to 5, becoming convex only in later whorls; general shape thus lightly fusiform. Suture very deep in early whorls, gradually becoming less pronounced but always incised. Whorls rotund initially, then less so; penultimate whorl compressed near suture, giving sinuous profile; last 1 1/2 whorls uniformly flattened laterally; last whorl slightly descending. Aperture 1/3 of shell height, suboval, slightly oblique, gently expanded at rounded lower half, with columellar, palatal margins nearly straight, parallel. Peristome sharp, somewhat reflexed along palatal margin more reflexed along columellar margin, gently flaring, its parietal insertions widely distant but joined by distinctive, shallow callus. In full-grown specimens one blunt angular tooth often present. Columella broadened internally, forming narrow flat area dominated by shallow vertical fold. Umbilicus very narrow slit, partially occluded by reflected columellar edge. Embryonic whorls 1 1/4, distinct, having sculpture (discernible only in best preserved specimens) consisting of regular, shallow radial grooves running from suture on upper side, thin spiral lines on bottom half of whorls (
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<figureCitation id="10A62A5DFFCCFFADFB1E74F8FB15F871" box="[1202,1276,1959,1985]" captionStart="FIGURE 1 – 2. 1" captionStartId="3.[151,255,1458,1482]" captionTargetBox="[151,1411,432,1457]" captionTargetId="figure@3.[151,1436,432,1460]" captionTargetPageId="3" captionText="FIGURE 1 – 2. 1. Balearena gymnesica gen. nov., spec. nov. Holotype, upper coastal cliff at eastern edge of Es Carnatge, Palma de Mallorca, Mallorca island, Balearic Islands, Western Mediterranean. Shell length 16.7 mm; scale bar 2.0 mm. Not shown are a thin encrustation over most of the first whorl and minimal fractures on the peristome. 2. Protoconch sculpture of Balearena gymnesica. This reconstruction is based on partial remains of these delicate features that are observable in various specimens. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/178668/files/figure.png" pageId="2" pageNumber="45">Fig. 2</figureCitation>
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||
). Teleoconch sculpture formed by low but rough, uneven, shallow radial wrinkles, giving dull shine. In approximately first two teleoconch whorls, delicate, interrupted spiral sculpture, subjugated to radial wrinkles. Color apparently uniform olive-brown, except for white peristome, without any markings. Height 13.5–16.7 mm, width 5.6–6.6 mm.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C0876553FFCDFFACFF6A7250FEDEFEE2" pageId="3" pageNumber="46" type="etymology">
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||
<paragraph id="882236D8FFCDFFACFF6A7250FEDEFEE2" blockId="3.[151,1436,152,418]" pageId="3" pageNumber="46">
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||
<emphasis id="BAE9EACAFFCDFFACFF6A7250FEBEFE99" bold="true" box="[198,343,271,297]" pageId="3" pageNumber="46">Etymology:</emphasis>
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The specific epithet refers to the Gymnesics, or northern Balearic Islands, where the species was endemic.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0876553FFCDFFACFF6A7200FA75FE12" pageId="3" pageNumber="46" type="discussion">
|
||
<paragraph id="882236D8FFCDFFACFF6A7200FA75FE12" blockId="3.[151,1436,152,418]" pageId="3" pageNumber="46">
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||
<emphasis id="BAE9EACAFFCDFFACFF6A7200FEABFEC9" bold="true" box="[198,322,351,377]" pageId="3" pageNumber="46">Remarks:</emphasis>
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||
The series of nine specimens discussed and figured by
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||
<bibRefCitation id="EC0C4B29FFCDFFACFC72723FFB6DFECA" author="Cuerda" box="[990,1156,352,378]" pageId="3" pageNumber="51" refString="Cuerda Barcelo, J. (1959) Presencia de Mastus pupa, Bruguiere en el tirreniense de las Baleares orientales. Boletin de la Sociedad de Historia Natural de Baleares, 5, 45 - 50, lam. iv." type="journal article" year="1959">Cuerda (1959</bibRefCitation>
|
||
,
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||
<bibRefCitation id="EC0C4B29FFCDFFACFB3D723FFB39FECA" author="Cuerda" box="[1169,1232,352,378]" pageId="3" pageNumber="51" refString="Cuerda, J. (1989) Los tiempos cuaternarios en Baleares. 2 nd ed. Conselleria de Cultura, Educacio i Esports, Govern Balear, Palma de Mallorca, 310 pp. + 20 pls." type="book" year="1989">1989</bibRefCitation>
|
||
), collected in the same stratum as the
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<typeStatus id="5726887AFFCDFFACFE2B72D7FE18FE12" box="[391,497,392,418]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
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, and kept in his collection now in SHNB (no catalog number), are also referred
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||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |