144 lines
18 KiB
XML
144 lines
18 KiB
XML
<document id="130A0F807F638DE44F8FADE3450C34A3" ENCODING="UTF-8" ID-GBIF-Dataset="446a0cdc-e5c9-4b1e-b677-254175699a77" ModsDocID="z00809p001" checkinTime="1247125880944" checkinUser="thomas" docAuthor="M. Eric Anderson" docDate="2005" docId="C5EDA0E3F24652AA8C04303F0C7D3D1C" docLanguage="en" docName="2005_Anderson2_gg2.xml" docOrigin="Zootaxa 809" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:8D4D671D-D408-4DEE-8550-BB0D16BE61B8" docTitle="Echiodon atopus Anderson, 2005, new species" docType="treatment" docUuid="48A1FBE4-167D-49F4-B78F-4F5AA97725AC" docUuidSource="ZooBank" docVersion="13" lastPageNumber="4" masterDocId="22B1FF708DBFCCAF40B4480E9DAB82B4" masterDocTitle="Description of a new species of Echiodon (Teleostei: Carapidae) from the South Atlantic Ocean." masterLastPageNumber="5" masterPageNumber="1" pageNumber="2" updateTime="1701314085112" updateUser="plazi">
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<mods:title id="74E5B9741634AB3C04234B40BFA8842A">Description of a new species of Echiodon (Teleostei: Carapidae) from the South Atlantic Ocean.</mods:title>
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<mods:namePart id="D70ABF6C0AA4FE59A9DB5F1B6FDE0D2E">M. Eric Anderson</mods:namePart>
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<mods:title id="B835A9F8BA8CD69E43711E7D678C905B">Zootaxa</mods:title>
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<mods:part id="6C404E4FDD482A69A4F3BC8D70520D26">
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<mods:date id="6504C821B68558C336A88D1E866A32B5">2005</mods:date>
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<mods:number id="4A79F217BF408579531C8E324304BC78">809</mods:number>
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<mods:classification id="01EEFECFA142F78C6BE8C467EC49A0C6">journal article</mods:classification>
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<treatment id="C5EDA0E3F24652AA8C04303F0C7D3D1C" ID-DOI="http://doi.org/10.5281/zenodo.6265015" ID-GBIF-Taxon="100124146" ID-Zenodo-Dep="6265015" LSID="urn:lsid:zoobank.org:act:48A1FBE4-167D-49F4-B78F-4F5AA97725AC" httpUri="http://treatment.plazi.org/id/C5EDA0E3F24652AA8C04303F0C7D3D1C" lastPageNumber="4" pageNumber="2">
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<subSubSection id="7F6C2FE38C196A5FE6B2A2274B2845B6" type="nomenclature">
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<paragraph id="040C33E642E88E8FB34C94EED7D0137E" pageNumber="2">
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<taxonomicName id="EF882598847195AAE743F30BF4029551" ID-CoL="38LN3" LSID-ZBK="urn:lsid:zoobank.org:act:48A1FBE4-167D-49F4-B78F-4F5AA97725AC" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon atopus Anderson 2005:2" order="Ophidiiformes" pageNumber="2" phylum="Chordata" rank="species" species="atopus">Echiodon atopus</taxonomicName>
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,
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<taxonomicNameLabel id="9C15DC1A17D06C4DE90D60AB44DB07E3" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="315B0E17F15CC731E6FFB3073BFB800C" type="description">
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<paragraph id="8F7D859E00F0701FEAD82F08E35CEC9F" pageNumber="2">(Figs. 1-2)</paragraph>
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</subSubSection>
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<subSubSection id="675C3FC25B90F6E81AECE8218D0E78C6" type="nomenclature">
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<paragraph id="10ECF0FB617BCF5FD9EED87527D577DB" pageNumber="2">
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<materialsCitation id="F0B9A74FC197255AB6658B4139AB43CD" ID-GBIF-Occurrence="923892894" collectingDate="2004-07-09" collectingMethod="10 m otter trawl; 0- 139 m" collectionCode="USNM" collectorName="RVIB NATHANIEL B. PALMER, cruise ICEFISH-04" latitude="-37.2725" location="north of Inaccessible Island, sta. 91/OT 60" longitude="-12.71083" specimenCode="USNM 380406" stateProvince="South Atlantic near Tristan da Cunha group" typeStatus="Holotype">
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<typeStatus id="7D8783397B48EFB75AC07288C129B704">Holotype</typeStatus>
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:
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<specimenCode id="371CC05A3FAD0B60EF1341027ED32ACC">
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<collectionCode id="1F2B86EDD33467599D73E15467442341" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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380406
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</specimenCode>
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(male, 176 mm TL),
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<location id="85673EF950A58ACE29F7E3843F4311C1" LSID="urn:lsid:plazi:treatment:C5EDA0E3F24652AA8C04303F0C7D3D1C:85673EF950A58ACE29F7E3843F4311C1" country="Great Britain" latitude="-37.27667" longitude="-12.718333" name="north of Inaccessible Island">north of Inaccessible Island</location>
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;
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<collectorName id="37B35E506E80FE29CF6179BBA2B8D6E2">RVIB NATHANIEL B. PALMER</collectorName>
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<location id="677DEA4C27CA6027D1655D3FF78BECEC" LSID="urn:lsid:plazi:treatment:C5EDA0E3F24652AA8C04303F0C7D3D1C:677DEA4C27CA6027D1655D3FF78BECEC" country="Great Britain" latitude="-37.27667" longitude="-12.718333" name="sta. 91/OT 60">sta. 91/OT 60</location>
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;
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<geoCoordinate id="D538264E69392FF5759C34BE0D5F63D4" direction="south" orientation="latitude" value="-37.268333">37° 16.1'S</geoCoordinate>
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,
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<geoCoordinate id="12FF280DA3C2954E540E54D26B3917C3" direction="west" orientation="longitude" value="-12.703333">12° 42.2'W</geoCoordinate>
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to
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<geoCoordinate id="EF564A986595AFA2306FF3D18C02CB13" direction="south" orientation="latitude" value="-37.27667">37° 16.6'S</geoCoordinate>
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,
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<geoCoordinate id="5913B9295A6F353ACCF829D98AF70CE2" direction="west" orientation="longitude" value="-12.718333">12° 43.1'W</geoCoordinate>
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;
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<collectingMethod id="4D6A8AB7545A6B6567E1562A00F41519">10 m otter trawl; 0- 139 m</collectingMethod>
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; 19:49- 20:06 hr;
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<collectingDate id="A11F274B3DCFC0A6D489C2BB13ADFF00" day="09" month="07">
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9
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<collectingDate id="B5D6F01C163A91ED61369F18F7BC0B54" month="07" value="2004-07" year="2004">July 2004</collectingDate>
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</collectingDate>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6C90636F59E3E3C5FA85AB8166A4D2DF" type="diagnosis">
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<paragraph id="E6443535763A189A3D712D4FC9AE9296" pageNumber="2">
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Diagnosis. A species of
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<taxonomicName id="E39D15782B8CECF0690E05A11AA9FA3D" LSID-ZBK="urn:lsid:zoobank.org:act:3582CA89-8236-4B5C-8098-37084743EA3D" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon Thompson 1837:55" order="Ophidiiformes" pageNumber="2" phylum="Chordata" rank="genus">Echiodon</taxonomicName>
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as defined by Markle and Olney (1990), with the following combination of characters: PCV 38; D30 46; A30 46; ARDO 7; P1 18; posterior swim bladder tunic ridges ventral.
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</paragraph>
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</subSubSection>
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<subSubSection id="D4E51D7FFD1242AE60D70340BB942688" type="description">
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<paragraph id="EB051CBF1070845F30ACC8D7C62AAFC9" pageNumber="2">Description. Meristics: PCV 38; D30 46; A30 46; P1 18; ARDO 7; DRAO 0; MAO 7; MDO 8; VAO 6; VDO 8; VPB 6; IO 8; SO 5; LT 3; MD 5; POP 2; ST 2; branchiostegal rays 7. Proportions as percent HL: body depth 48.8; bony interorbital width 13.7; eye diameter 20.3; head depth 48.8; head width 34.3; lower jaw length 52.0; upper-jaw length 46.4; prepectoral length 96.4; preanal length 118.1; predorsal length 129.3; pectoral-fin length 65.2; snout length 17.0; distance snout to anus 111.8.</paragraph>
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<paragraph id="BAB73E921D8FA980C370E064944FF61A" lastPageNumber="4" pageNumber="3">
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Head elongate-ovoid; eye circular, entering dorsal profile of head when viewed laterally. Cephalic pores small, rounded, difficult to observe, pored lateral line mediolateral, extending posteriorly to anterior quarter of tail; pores becoming more widely spaced posteriad. No sensory papillae on head or along seismosensory canals. First gill arch with three elongate, tooth pads on upper limb, three slender, developed rakers and four rounded, toothed pads on lower limb. Posterior portion of maxilla unsheathed. Dorsal portion of gill opening extending forward about 3/4 eye diameter. Branchiostegal membrane free of isthmus, joined on vertical about 1 1/2 eye diameters posterior to rear margin of eye. Single enlarged caniniform tooth near symphysis in premaxilla, dentary with two. Vomer with three smaller canines in a row surrounded by 25 tiny denticles. Palatine teeth in four irregular rows anteriorly blending into single, posterior row. Outer flap of upper lip at symphysis with single, furcate, ventrally-directed lobe fused to a broad, inner flange with a larger, conical lobe on either side. Dorsal-fin origin one eye diameter posterior to vertical through anal-fin origin. Posterior swim bladder constricted, with ventral tunic ridges as illustrated for
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<taxonomicName id="42D3C1A64FDC24D2BD49D4F382611411" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophidiiformes" pageNumber="3" phylum="Chordata" rank="species" species="dentatus">E. dentatus (Cuvier)</taxonomicName>
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by Markle and Olney (1990, fig. 35).
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</paragraph>
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<paragraph id="66C5BA3C39F445AAAA896B972C7420F5" pageNumber="4">Fresh coloration silvery blue, with fine, scattered melanophores. These are denser on dorsum than ventrally, except on tail, and form chevron patterns outlining myomeres on body posteriorly to middle of tail. Abdomen and eye dark blue. Peritoneum, stomach, lining of orobranchial chamber, including tongue and gill bars, black.</paragraph>
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</subSubSection>
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<subSubSection id="2E2118273CA4D54BF43654ADA76BAB78" type="etymology">
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<paragraph id="5A65698D025EF93F4B38EE6176F595F3" pageNumber="4">
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Etymology. From the Greek atopos, (out of place) in reference to my original expectation that the specimen was
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<taxonomicName id="23D6136CF006A3BB7B2CF1754173BD27" LSID-ZBK="urn:lsid:zoobank.org:act:2EE173D4-A7C6-4A62-B531-A7B18313F5AD" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon cryomargarites Markle, Williams & Olney 1983:647" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="cryomargarites">Echiodon cryomargarites</taxonomicName>
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and its subsequent identification.
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</paragraph>
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</subSubSection>
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<subSubSection id="2E84F8E9AC271AFED5DA214E82BE1287" type="discussion">
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<paragraph id="086FCA5A1D8D60B9A84F5845BED112FC" pageNumber="4">
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Comparisons. Markle and Olney’s (1990) zoogeographic analysis of
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<taxonomicName id="C1DC751D8243827D6A9A893DF55AC337" LSID-ZBK="urn:lsid:zoobank.org:act:3582CA89-8236-4B5C-8098-37084743EA3D" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon Thompson 1837:55" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="genus">Echiodon</taxonomicName>
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revealed three distributional patterns that they hypothesized resulted from species radiation of an outer-shelf Indo-West Pacific ancestor. These are: 1) North Atlantic; 2) austral; and 3) a largely boreal temperate/subtropical group that includes the austral
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<taxonomicName id="70BA6D2A7FF11C2C9B2F8F77572A9A80" LSID-ZBK="urn:lsid:zoobank.org:act:2EE173D4-A7C6-4A62-B531-A7B18313F5AD" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon cryomargarites Markle, Williams & Olney 1983:647" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="cryomargarites">E. cryomargarites</taxonomicName>
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.
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<taxonomicName id="B040C9714EA6956576EEE5B47E90050B" LSID-ZBK="urn:lsid:zoobank.org:act:48A1FBE4-167D-49F4-B78F-4F5AA97725AC" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon atopus Anderson 2005:2" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="atopus">Echiodon atopus</taxonomicName>
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is close to the austral
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<taxonomicName id="6D9C6BF55EFCE305FECA9D91AB9366D6" LSID-ZBK="urn:lsid:zoobank.org:act:9040DE1C-5C52-4575-8CB6-ACC8442FC8C5" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon neotes Markle & Olney 1990:351" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="neotes">E. neotes Markle and Olney</taxonomicName>
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1990 and
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<taxonomicName id="92D8AEB51CA8D59BFC1C3030C50EC7FF" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="rendahli">E. rendahli (Whitley)</taxonomicName>
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and seems to fit in that clade, although its larvae are unknown (Williams and Machida 1992). It differs from
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<taxonomicName id="60D24E4AAE683DEDF490011C1FB793AF" LSID-ZBK="urn:lsid:zoobank.org:act:9040DE1C-5C52-4575-8CB6-ACC8442FC8C5" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon neotes Markle & Olney 1990:351" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="neotes">E. neotes</taxonomicName>
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in its significantly higher D30 count (38 in
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<taxonomicName id="89658B58D45F70773EBEAC794C021276" LSID-ZBK="urn:lsid:zoobank.org:act:9040DE1C-5C52-4575-8CB6-ACC8442FC8C5" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon neotes Markle & Olney 1990:351" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="neotes">E. neotes</taxonomicName>
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vs 46 in
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<taxonomicName id="5AB1BBBC45C5F0F7A86279736BF2893B" LSID-ZBK="urn:lsid:zoobank.org:act:48A1FBE4-167D-49F4-B78F-4F5AA97725AC" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon atopus Anderson 2005:2" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="atopus">E. atopus</taxonomicName>
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), more precaudal vertebrae (35 in
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<taxonomicName id="8C84C47B2F0B4EF55F3D89F3D1C025A9" LSID-ZBK="urn:lsid:zoobank.org:act:9040DE1C-5C52-4575-8CB6-ACC8442FC8C5" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon neotes Markle & Olney 1990:351" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="neotes">E. neotes</taxonomicName>
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vs 38), ventral swim bladder tunic ridges (vs. lateral ridges in
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<taxonomicName id="80ED357BB1587C550D95A096E878211D" LSID-ZBK="urn:lsid:zoobank.org:act:9040DE1C-5C52-4575-8CB6-ACC8442FC8C5" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon neotes Markle & Olney 1990:351" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="neotes">E. neotes</taxonomicName>
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) and lack of the larval vexillum, although the latter character is difficult to evaluate in the one known specimen of
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<taxonomicName id="08EEFFFD847026E75B5B95EEA765C31E" LSID-ZBK="urn:lsid:zoobank.org:act:9040DE1C-5C52-4575-8CB6-ACC8442FC8C5" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon neotes Markle & Olney 1990:351" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="neotes">E. neotes</taxonomicName>
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.
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<taxonomicName id="FE5500DD1BFEDE0544F6D5FD11762C58" LSID-ZBK="urn:lsid:zoobank.org:act:48A1FBE4-167D-49F4-B78F-4F5AA97725AC" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon atopus Anderson 2005:2" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="atopus">Echiodon atopus</taxonomicName>
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differs from
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<taxonomicName id="43BAD792F13D0D26F8B07EFD81771316" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="rendahli">E. rendahli</taxonomicName>
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in its A30 count (37-38 in
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<taxonomicName id="FB9AC2F6D20E14872BC8FA68C434DA47" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="rendahli">E. rendahli</taxonomicName>
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vs 46 in
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<taxonomicName id="07A17CD3566B08C5182C29434186AC00" LSID-ZBK="urn:lsid:zoobank.org:act:48A1FBE4-167D-49F4-B78F-4F5AA97725AC" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon atopus Anderson 2005:2" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="atopus">E. atopus</taxonomicName>
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), greater number of anal-fin rays anterior to the dorsal-fin origin (0-4 in
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<taxonomicName id="210E4D2BDD3EE28E93CC7CB2B536808A" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="rendahli">E. rendahli</taxonomicName>
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vs 7) and greater number of precaudal vertebrae (31-35 in
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<taxonomicName id="1B40CE51B1A02744F29D93C838D23AEC" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="rendahli">E. rendahli</taxonomicName>
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vs 38). Finally,
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<taxonomicName id="21671ABE397D53B1A43B37A49F5E43FD" LSID-ZBK="urn:lsid:zoobank.org:act:48A1FBE4-167D-49F4-B78F-4F5AA97725AC" family="Carapidae" genus="Echiodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Echiodon atopus Anderson 2005:2" order="Ophidiiformes" pageNumber="4" phylum="Chordata" rank="species" species="atopus">E. atopus</taxonomicName>
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differs from all other congeners in its equal D30 and A30 counts, although, owing to variability in fin-ray anatomy in ophidiiform fishes, this should not be expected to endure as a diagnostic character for the species when more specimens are found.
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</paragraph>
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</subSubSection>
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</treatment>
|
||
</document> |