357 lines
32 KiB
XML
357 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. & Dowling, Ashley P. G." docDate="2017" docId="06B3F8E2AF9BFE1D2CAF842BAA8B417B" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola pacificensis Fisher & Dowling, sp. n." docType="treatment" docUuid="CEB459C5-08DD-4A1B-A84F-0B2113AAEC2D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, J. Ray</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Danielle M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skvarla, Michael J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nelson, Whitney A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>701</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>496</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
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<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="134839651" LSID="urn:lsid:zoobank.org:act:CEB459C5-08DD-4A1B-A84F-0B2113AAEC2D" httpUri="http://treatment.plazi.org/id/06B3F8E2AF9BFE1D2CAF842BAA8B417B" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/CEB459C5-08DD-4A1B-A84F-0B2113AAEC2D" authority="Fisher & Dowling" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">Torrenticola pacificensis Fisher & Dowling</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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HOLOTYPE (♀): from USA, Washington, Clallam County, Green Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="48.17917">48°10'45"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-124.20583">124°12'21"W</geoCoordinate>
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), 24 Jul 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0724-005.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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PARATYPES (34 ♀; 31 ♂): British Columbia, Canada: 2 ♀ and 1 ♂ from Vancouver Island, Caycuse, Nixon Creek, 8 Jul 1976, by IM Smith, IMS760197 & IMS760198
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Vancouver Island, beside Highway 4, 35.6 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS760206
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Vancouver Island, beside Highway 4, 1.3 kilometers east of Pacific Rim Road, Lost Shoe Creek, 9 Jul 1976, by IM Smith, IMS760202
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<normalizedToken originalValue="•">*</normalizedToken>
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2 ♀ and 2 ♂ from Vancouver Island, Lake Cowichan, Skutz Falls, Skutz Creek, near Cowichan River, 9 Jul 1979, by IM Smith, IMS790036A & IMS790036B
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Vancouver Island, Lake Cowichan, Cowichan River, above Skutz Falls, 9 Jul 1979, by IM Smith, IMS790035
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, Robertson River, 4 Jul 1976, by IM Smith, IMS760183A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, tributary of Robertson River, 10 Jul 1976, by IM Smith, IMS760183B
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, tributary of Robertson River, 4-10 Jul 1976, by IM Smith, IMS760182
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, Goldstream River, 26 Jun 1979, by IM Smith, IMS790028A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, near Mt. Finlayson Trail, 26 Jun 1979, by IM Smith, IMS790027A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Vancouver Island, beside South Shore Road, 2.3 kilometers south of Caycuse, 26 Jul 1979, by IM Smith, IMS790052
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Vancouver Island, beside North Shore Road, 1.7 kilometers north of Lake Cowichan, 7 Jun 1979, by IM Smith, IMS790008C
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Vancouver Island, North Shore Road, 3.2 kilometers south of Youbou, 4 Jul 1976, by IM Smith, IMS760190
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Vancouver Island, Youbou, North Shore Road, Shaw Creek, 4.3 kilometers south of north end of Cowichan Lake, by IM Smith, IMS760196
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<normalizedToken originalValue="•">*</normalizedToken>
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Oregon, USA: 1 ♀ and 1 ♂ from Benton County, Marys Peak near Philomath, Parker Creek, 27-28 Jun 1983, by IM Smith & AB Smith, IMS830006
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Benton County, Marys Peak near Philomath, Parker Creek, 28 Jun 1983, by IM Smith, IMS830007
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Benton County, Philomath, Marys Peak campground, Park Creek, 23 Jun 1976, by IM Smith, IMS760146
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Coos County, Siskiyou National Forest, road 33 between Powers and Agness, Coal Creek, 2 Jul 1983, by IM Smith, IMS830015
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Coos County, Siskiyou National Forest, road 33 between Powers and Agness, 0.5 kilometers east of Daphne Grove campground, 2 Jul 1983, by IM Smith, IMS830017
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<normalizedToken originalValue="•">*</normalizedToken>
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3 ♀ and 2 ♂ from Curry County, Port Orford, small spring run beside road from Humbug Mountain State Park to McGribble campground, 25 Jun 1976, IMS760161
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, Brush Creek, beside Route 1, 3 Jul 1983, by IM Smith, IMS830020A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Curry County, Siskiyou National Forest, road 33 between Powers and Agness, North Fork of Foster Creek, 2 Jul 1983, by IM Smith, IMS830019
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Lane County, Gate Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="44.146667">44°8'48"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-122.57222">122°34'20"W</geoCoordinate>
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), 11 Aug 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0811-001
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Lincoln County, Blackberry campground near Tidewater, Alsea River, 28 Jun 1983, by IM Smith & AB Smith, IMS830009
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<normalizedToken originalValue="•">*</normalizedToken>
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4 ♀ and 2 ♂ from Marion County, Marion Forks Riverside campground, North Fork of Santiam River, 22 Jun 1976, by IM Smith, IMS760145
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<normalizedToken originalValue="•">*</normalizedToken>
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2 ♀ and 2 ♂ from Multnomah County, Columbia River Scenic Highway, stream below Horsetail Falls, 27 Jun 1983, by IM Smith, IMS830005
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Tillamook County, Siuslaw National Forest, Alder Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="45.1575">45°9'27"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-123.8">123°47'60"W</geoCoordinate>
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), 6 Aug 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0806-002
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<normalizedToken originalValue="•">*</normalizedToken>
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Washington, USA: 1 ♂ (ALLOTYPE) from Clallam County, Green Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="48.17917">48°10'45"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-124.20583">124°12'21"W</geoCoordinate>
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), 24 Jul 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0724-005
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<normalizedToken originalValue="•">*</normalizedToken>
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2 ♀ and 1 ♂ from Clallam County, Green Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="48.17917">48°10'45"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-124.20583">124°12'21"W</geoCoordinate>
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), 24 Jul 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0724-005
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<normalizedToken originalValue="•">*</normalizedToken>
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2 ♀ and 1 ♂ from Lewis County, Gifford Pinchot National Forest, Snake Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="46.64778">46°38'52"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-121.71889">121°43'8"W</geoCoordinate>
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), 23 Jul 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0723-006
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 3 ♂ from Snohomish County, Mount Baker National Forest, Marten River (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="48.071945">48°4'19"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-121.60667">121°36'24"W</geoCoordinate>
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), 28 Jul 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0728-002.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type deposition">
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<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
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<paragraph pageId="0" pageNumber="1">Holotype (♀), allotype (♂), and most paratypes (29 ♀; 325♂) deposited in the CNC; other paratypes (5 ♀; 5 ♂) deposited in ACUA.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">Torrenticola pacificensis</taxonomicName>
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are similar to members of the Miniforma group (
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<taxonomicName lsidName="T. copipalpa" pageId="0" pageNumber="1" rank="species" species="copipalpa">T. copipalpa</taxonomicName>
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,
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<taxonomicName lsidName="T. manni" pageId="0" pageNumber="1" rank="species" species="manni">T. manni</taxonomicName>
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,
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<taxonomicName lsidName="T. miniforma" pageId="0" pageNumber="1" rank="species" species="miniforma">T. miniforma</taxonomicName>
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,
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<taxonomicName lsidName="T. rockyensis" pageId="0" pageNumber="1" rank="species" species="rockyensis">T. rockyensis</taxonomicName>
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,
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<taxonomicName lsidName="T. oliveri" pageId="0" pageNumber="1" rank="species" species="oliveri">T. oliveri</taxonomicName>
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, and
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<taxonomicName lsidName="T. pinocchio" pageId="0" pageNumber="1" rank="species" species="pinocchio">T. pinocchio</taxonomicName>
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) in having short, stocky pedipalps (except
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<taxonomicName lsidName="T. oliveri" pageId="0" pageNumber="1" rank="species" species="oliveri">T. oliveri</taxonomicName>
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and
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<taxonomicName lsidName="T. pinocchio" pageId="0" pageNumber="1" rank="species" species="pinocchio">T. pinocchio</taxonomicName>
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); similar pedipalpal extensions (unique to members of this group); and being among the smallest
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Torrenticola</taxonomicName>
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in the west (dorsum 500-625 long) (except
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<taxonomicName lsidName="T. oliveri" pageId="0" pageNumber="1" rank="species" species="oliveri">T. oliveri</taxonomicName>
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).
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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are best differentiated from
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<taxonomicName lsidName="T. rockyensis" pageId="0" pageNumber="1" rank="species" species="rockyensis">T. rockyensis</taxonomicName>
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by females having a stockier rostrum (length/width ♀ = 2.59-2.68 in
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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, 2.72-2.91 in
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<taxonomicName lsidName="T. rockyensis" pageId="0" pageNumber="1" rank="species" species="rockyensis">T. rockyensis</taxonomicName>
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); and by being distributed in the Pacific Coast Ranges of Washington and Oregon instead of the Rocky Mountains (Idaho & Montana).
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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are best differentiated from
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<taxonomicName lsidName="T. copipalpa" pageId="0" pageNumber="1" rank="species" species="copipalpa">T. copipalpa</taxonomicName>
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by having tuberculate pedipalp femoral extensions (broad and flat in
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<taxonomicName lsidName="T. copipalpa" pageId="0" pageNumber="1" rank="species" species="copipalpa">T. copipalpa</taxonomicName>
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) and by being distributed in the Pacific Coast Ranges of Washington and Oregon, whereas although
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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overlap, they are more southern (southwest Oregon & California).
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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are best differentiated from
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<taxonomicName lsidName="T. manni" pageId="0" pageNumber="1" rank="species" species="manni">T. manni</taxonomicName>
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by having stockier tibiae (length/width = 2.67-3.00 in
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<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">pacificensis</taxonomicName>
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, 3.13-3.38 in
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<taxonomicName lsidName="T. manni" pageId="0" pageNumber="1" rank="species" species="manni">T. manni</taxonomicName>
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); having a stockier rostrum (length/width ♀ = 2.59-2.68 in
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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, 3.00-3.13 in
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<taxonomicName lsidName="T. manni" pageId="0" pageNumber="1" rank="species" species="manni">T. manni</taxonomicName>
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; ♂ = 2.76-3.07 in
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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, 3.13-3.20 in
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<taxonomicName lsidName="T. manni" pageId="0" pageNumber="1" rank="species" species="manni">T. manni</taxonomicName>
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); and by being distributed in the Pacific Coast Ranges of Washington and Oregon, whereas
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<taxonomicName lsidName="T. manni" pageId="0" pageNumber="1" rank="species" species="manni">T. manni</taxonomicName>
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is only known from Catron County, New Mexico.
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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are best differentiated from
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<taxonomicName lsidName="T. miniforma" pageId="0" pageNumber="1" rank="species" species="miniforma">T. miniforma</taxonomicName>
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by having stockier anterio-medial platelets (length/width = 2.00-2.16 in
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<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
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, 2.44-2.65 in
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<taxonomicName lsidName="T. miniforma" pageId="0" pageNumber="1" rank="species" species="miniforma">T. miniforma</taxonomicName>
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); stockier subcapitular rostra (length/width = 2.59-3.07 in
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
; 3.13-3.19 in
|
||
<taxonomicName lsidName="T. miniforma" pageId="0" pageNumber="1" rank="species" species="miniforma">T. miniforma</taxonomicName>
|
||
); and by being distributed in the Pacific Coast Ranges of Washington and Oregon, whereas
|
||
<taxonomicName lsidName="T. miniforma" pageId="0" pageNumber="1" rank="species" species="miniforma">T. miniforma</taxonomicName>
|
||
is only known from Humboldt County, California.
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
can be differentiated from
|
||
<taxonomicName lsidName="T. oliveri" pageId="0" pageNumber="1" rank="species" species="oliveri">T. oliveri</taxonomicName>
|
||
by having a shorter anterior venter (177-235 in
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
, 250-310 in
|
||
<taxonomicName lsidName="T. oliveri" pageId="0" pageNumber="1" rank="species" species="oliveri">T. oliveri</taxonomicName>
|
||
) and less elongate pedipalpal tibiae (length/width = 2.6-3.0 in
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
, 3.6-4.2 in
|
||
<taxonomicName lsidName="T. oliveri" pageId="0" pageNumber="1" rank="species" species="oliveri">T. oliveri</taxonomicName>
|
||
).
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
can be differentiated from
|
||
<taxonomicName lsidName="T. pinocchio" pageId="0" pageNumber="1" rank="species" species="pinocchio">T. pinocchio</taxonomicName>
|
||
by having a less elongate rostrum (length/width = 2.5-3.1 in
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
, 4.5-4.9 in
|
||
<taxonomicName lsidName="T. pinocchio" pageId="0" pageNumber="1" rank="species" species="pinocchio">T. pinocchio</taxonomicName>
|
||
) and less elongate pedipalpal tibiae (length/width = 2.67-3.00 in
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
, 3.13-3.50 in
|
||
<taxonomicName lsidName="T. pinocchio" pageId="0" pageNumber="1" rank="species" species="pinocchio">T. pinocchio</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Female (Figure 182) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Dorsum - (530-625 (580) long; 335-430 (400) wide) ovoid with purple coloration restricted posteriorly. Anterio-medial platelets (100-117.5 (110) long; 47.5-55 (52.5) wide). Anterio-lateral platelets (145-170 (162.5) long; 52.5-63.75 (57.5) wide) free from dorsal plate. Dgl-4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl-4 270-345 (300)). Dorsal plate proportions: dorsum length/width 1.42-1.58 (1.45); dorsal width/distance between Dgl-4 1.24-1.36 (1.33); anterio-medial platelet length/width 2.05-2.14 (2.10); anterio-lateral platelet length/width 2.63-2.95 (2.83); anterio-lateral/anterio-medial length 1.43-1.48 (1.48).</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (287.5-325 (300) long (ventral); 210-236 (225) long (dorsal); 110-125 (115) tall) colorless. Rostrum (110-127.5 (120) long; 41.25-47.5 (45) wide). Chelicerae (265-331 (305) long) with curved fangs (50-62 (52.5) long). Subcapitular proportions: ventral length/height 2.52-2.65 (2.61); rostrum length/width 2.59-2.68 (2.67). Pedipalps short and stocky (especially tibiae) with nearly tuberculate, dentate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30-35 (32.5) long); femur (85-95 (95) long); genu (57.5-67.5 (65) long); tibia (55-67.5 (62.5) long; 20-23.75 (22.5) wide); tarsus (12.5-17.5 (15) long). Palpomere proportions: femur/genu 1.40-1.48 (1.46); tibia/femur 0.65-0.71 (0.66); tibia length/width 2.71-3.00 (2.78).</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Venter - (650-770 (720) long; 434-521 (440) wide) colorless or with faint purple coloration. Gnathosomal bay (122.5-142.5 (140) long; 66.25-87.5 (67.5) wide). Cxgl-4 subapical. Medial suture (47.5-55 (50) long). Genital plates (162.5-185 (180) long; 143.75-172.5 (160) wide). Additional measurements: Cx-1 (230-289 (250) long (total); 110-142 (120) long (medial)); Cx-3 (280-331 (280) wide); anterior venter (177.5-205 (182.5) long). Ventral proportions: gnathosomal bay length/width 1.49-2.07 (2.07); anterior venter/genital field length 1.01-1.11 (1.01); anterior venter length/genital field width 1.07-1.23 (1.14); anterior venter/medial suture 3.38-3.89 (3.65).</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Male (Figure 183) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Dorsum - (525-590 (550) long; 355-375 (370) wide) ovoid with purple coloration restricted posteriorly. Anterio-medial platelets (97.5-115 (102.5) long; 47.5-55 (47.5) wide). Anterio-lateral platelets (147.5-162.5 (147.5) long; 52.5-57.5 52.5) wide) free from dorsal plate. Dgl-4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl-4 280-290 (290)). Dorsal plate proportions: dorsum length/width 1.48-1.57 (1.49); dorsal width/distance between Dgl-4 1.28-1.30 (1.28); anterio-medial platelet length/width 2.00-2.16 (2.16); anterio-lateral platelet length/width 2.73-2.83 (2.81); anterio-lateral/anterio-medial length 1.41-1.54 (1.44).</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (275-287.5 (285) long (ventral); 195-220 (212) long (dorsal); 103.75-115 (107.5) tall) colorless. Rostrum (107.5-117.5 (113.75) long; 35-42.5 (38.75) wide). Chelicerae (251-281 (273) long) with curved fangs (46-57 (47) long). Subcapitular proportions: ventral length/height 2.50-2.65 (2.65); rostrum length/width 2.76-3.07 (2.94). Pedipalps short and stocky (especially tibiae) with nearly tuberculate, dentate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30-32.5 (32.5) long); femur (82.5-90 (85) long); genu (57.5-65 (62.5) long); tibia (55-60 (57.5) long; 20-22.5 (21.25) wide); tarsus (15-16.25 (16.25) long). Palpomere proportions: femur/genu 1.36-1.50 (1.36); tibia/femur 0.65-0.68 (0.68); tibia length/width 2.67-2.94 (2.71).</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Venter - (645-740 (740) long; 388-460 (431) wide) colorless or with faint purple coloration. Gnathosomal bay (120-140 (135) long; 60-75 (60) wide). Cxgl-4 subapical. Medial suture (82.5-87.5 (82.5) long). Genital plates (140-147.5 (140) long; 110-120 (113.75) wide). Additional measurements: Cx-1 (243-277 (277) long (total); 120-146 (125) long (medial)); Cx-3 (245-310 (300) wide); anterior venter (216.25-235 (220) long). Ventral proportions: gnathosomal bay length/width 1.60-2.25 (2.25); anterior venter/genital field length 1.50-1.59 (1.57); anterior venter length/genital field width 1.89-1.98 (1.93); anterior venter/medial suture 2.49-2.69 (2.67).</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Specific epithet (
|
||
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">pacificensis</taxonomicName>
|
||
) refers to the distribution of this species in the Pacific Ranges of the Pacific Northwest. This location-based name reflects that locality is the best way to differentiate this species from others in the Miniforma Group, particularly
|
||
<taxonomicName lsidName="T. rockyensis" pageId="0" pageNumber="1" rank="species" species="rockyensis">T. rockyensis</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Western Oregon and western Washington (Figure 181). It seems reasonable to suspect
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
also occurs in coastal British Columbia and northwestern California.
|
||
</paragraph>
|
||
<caption pageId="0" pageNumber="1">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Figure 181.
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">Torrenticola pacificensis</taxonomicName>
|
||
sp. n. distribution.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="1">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Figure 182.
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">Torrenticola pacificensis</taxonomicName>
|
||
sp. n. female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="1">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Figure 183.
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">Torrenticola pacificensis</taxonomicName>
|
||
sp. n. male: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola pacificensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pacificensis">Torrenticola pacificensis</taxonomicName>
|
||
groups with other members of the Miniforma Complex with high support and specimens of this species are less than 1% different in COI sequence from each other. In all analyses,
|
||
<taxonomicName lsidName="T. pacificensis" pageId="0" pageNumber="1" rank="species" species="pacificensis">T. pacificensis</taxonomicName>
|
||
groups with three other morphologically similar species:
|
||
<taxonomicName lsidName="T. rockyensis" pageId="0" pageNumber="1" rank="species" species="rockyensis">T. rockyensis</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="T. manni" pageId="0" pageNumber="1" rank="species" species="manni">T. manni</taxonomicName>
|
||
, and
|
||
<taxonomicName lsidName="T. copipalpa" pageId="0" pageNumber="1" rank="species" species="copipalpa">T. copipalpa</taxonomicName>
|
||
. These three species are greater than 4% different from each other in COI sequence. This species overlaps with
|
||
<taxonomicName lsidName="T. copipalpa" pageId="0" pageNumber="1" rank="species" species="copipalpa">T. copipalpa</taxonomicName>
|
||
in west-central Oregon, but is the only member of the Miniforma group that occurs north of that point of overlap.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Based upon overall similarity, the pedipalp genu extensions, and western distribution, we were able to place this species in the Miniforma Identification Group.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">This species hypothesis is supported by biogeography, low COI divergence within the species (0-2%) and high divergence between species (3-15%), and by the morphological characters outlined in the diagnosis.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |