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<document id="14D0BE52DCDE04C68448CABE0AF71646" ID-CLB-Dataset="25399" ID-DOI="10.11646/zootaxa.4648.1.4" ID-GBIF-Dataset="255c331c-4667-4bdd-8c3b-86eac44e7cd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3353927" ID-ZooBank="E1180B6D-4783-4858-8E6D-A72AE7078203" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564384110305" checkinUser="plazi" docAuthor="Bezark, Larry G. &amp; Santos-Silva, Antonio" docDate="2019" docId="03A887FA09610B6AFF25F89DFF3476E6" docLanguage="en" docName="zootaxa.4648.1.4.pdf" docOrigin="Zootaxa 4648 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tethystola alboangulata Bezark &amp; Santos-Silva 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="102" masterDocId="FF91FF8209660B60FFB2FFF3FF88744D" masterDocTitle="Synonymies and seven new species in Lamiinae from the Neotropical Region (Coleoptera, Cerambycidae)" masterLastPageNumber="110" masterPageNumber="92" pageNumber="99" updateTime="1698747737307" updateUser="ExternalLinkService">
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<mods:title id="DA791CE6C2B6E3214DD68B5CDFD22DBB">Synonymies and seven new species in Lamiinae from the Neotropical Region (Coleoptera, Cerambycidae)</mods:title>
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<mods:namePart id="105239EFC53CA5B6EBEB008CB9F1D025">Bezark, Larry G.</mods:namePart>
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<mods:namePart id="56F386DB449039AE7F2CF30A5D4C6711">Santos-Silva, Antonio</mods:namePart>
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<mods:date id="5C1946753EEBAA051E4F1BE4D844B66B">2019</mods:date>
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<mods:number id="421F29FF5732D210E1944E25305B241A">2019-07-29</mods:number>
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<mods:identifier id="43E8CC67928FA1472E4A3D884BC0FE82" type="DOI">10.11646/zootaxa.4648.1.4</mods:identifier>
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<treatment id="03A887FA09610B6AFF25F89DFF3476E6" ID-DOI="http://doi.org/10.5281/zenodo.5943471" ID-GBIF-Taxon="159398521" ID-Zenodo-Dep="5943471" LSID="urn:lsid:plazi:treatment:03A887FA09610B6AFF25F89DFF3476E6" httpUri="http://treatment.plazi.org/id/03A887FA09610B6AFF25F89DFF3476E6" lastPageId="10" lastPageNumber="102" pageId="7" pageNumber="99">
<subSubSection id="C31B656709610B67FF25F89DFD9673C5" box="[151,542,1902,1929]" pageId="7" pageNumber="99" type="nomenclature">
<paragraph id="8BBE36EC09610B67FF25F89DFD9673C5" blockId="7.[151,542,1902,1964]" box="[151,542,1902,1929]" pageId="7" pageNumber="99">
<heading id="D0F6818009610B67FF25F89DFD9673C5" bold="true" box="[151,542,1902,1929]" fontSize="11" level="1" pageId="7" pageNumber="99" reason="1">
<emphasis id="B975EAFE09610B67FF25F89DFD9673C5" bold="true" box="[151,542,1902,1929]" pageId="7" pageNumber="99">
<taxonomicName id="4C014D6F09610B67FF25F89DFE3173C4" ID-CoL="BLM2H" authority="Bezark &amp; Santos-Silva, 2019" authorityName="Bezark &amp; Santos-Silva" authorityYear="2019" box="[151,441,1902,1929]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="99" phylum="Arthropoda" rank="species" species="alboangulata" status="sp. nov.">
<emphasis id="B975EAFE09610B67FF25F89DFE3173C4" bold="true" box="[151,441,1902,1929]" italics="true" pageId="7" pageNumber="99">Tethystola alboangulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246578509610B67FE72F89DFD9673C5" box="[448,542,1902,1928]" pageId="7" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B656709610B67FF25F861FEC373E1" box="[151,331,1938,1964]" pageId="7" pageNumber="99" type="description">
<paragraph id="8BBE36EC09610B67FF25F861FEC373E1" blockId="7.[151,542,1902,1964]" box="[151,331,1938,1964]" pageId="7" pageNumber="99">
(
<figureCitation id="133A2A6909610B67FF2DF861FECA73E1" box="[159,322,1938,1964]" captionStart="FIGURES 19–27. 19–22" captionStartId="9.[151,264,1895,1920]" captionTargetBox="[208,1386,168,1876]" captionTargetId="figure@9.[195,1392,166,1876]" captionTargetPageId="9" captionText="FIGURES 19–27. 19–22) Tethystola alboangulata, holotype female: 19) Dorsal habitus; 20) Ventral habitus; 21) Lateral habitus; 22) Head, frontal view. 23) Tethystola flavoapicalis Breuning, 1942 (= T. brasiliensis), dorsal habitus, holotype. 24) Tethystola brasiliensis, dorsal habitus, holotype. 25–26) Tethystola minima, dorsal habitus: 25) Holotype male; 26) Male from Venezuela. 27) Tethystola inermis, dorsal habitus, male from Venezuela." httpUri="https://zenodo.org/record/3353935/files/figure.png" pageId="7" pageNumber="99">Figures 19–22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B656709610B6AFF25F82AFF3476E6" lastPageId="10" lastPageNumber="102" pageId="7" pageNumber="99" type="materials_examined">
<paragraph id="8BBE36EC09610B68FF25F82AFA1E74B7" blockId="7.[151,1436,2009,2036]" lastBlockId="8.[151,1437,152,2015]" lastPageId="8" lastPageNumber="100" pageId="7" pageNumber="99">
<emphasis id="B975EAFE09610B67FF25F82AFEA973B9" bold="true" box="[151,289,2009,2036]" pageId="7" pageNumber="99">Description</emphasis>
.
<emphasis id="B975EAFE09610B67FE83F829FE0073B9" bold="true" box="[305,392,2010,2036]" pageId="7" pageNumber="99">Female</emphasis>
. Integument mostly dark brown, almost black on some areas; mouthparts and distal area of labrum dark reddish-brown; antennomeres V–XI with basal reddish-brown ring; elytra slightly, gradually lighter toward apex, with narrow orange-brown band along suture; meso- and metatibiae, and meso- and metatarsi dark reddish-brown depending on intensity of light; central area of abdominal ventrites I–IV dark reddish-brown centrally.
</paragraph>
<paragraph id="8BBE36EC096E0B68FF75FEF6FA967712" blockId="8.[151,1437,152,2015]" pageId="8" pageNumber="100">
<emphasis id="B975EAFE096E0B68FF75FEF6FE8F7552" bold="true" box="[199,263,261,287]" pageId="8" pageNumber="100">Head</emphasis>
. Frons, vertex, and area behind eyes moderately coarsely, abundantly punctate; with yellowish-brown pubescence not obscuring integument, slightly denser laterally on frons and vertex, with a few long, erect, thick white setae interspersed, and long, erect yellowish-brown setae interspersed close to eyes. Antennal tubercles elevated, with sculpturing as on frons basally, smooth at apex; with yellowish-brown pubescence, as on frons basally, denser toward apex. Genae transversely, finely striate, except smooth apex; with yellowish-brown pubescence centrally, nearly glabrous close to eye, glabrous posteriorly. Postclypeus with sculpturing and pubescence as on frons on wide central area, with long, erect yellowish-brown setae interspersed, smooth and glabrous laterally. Labrum coplanar with anteclypeus on posterior 2/3, inclined on anterior third; finely, abundantly punctate on coplanar area; with yellowish-brown pubescence distinctly not obscuring integument on coplanar area, with long, erect setae of same color interspersed; anterior margin with fringe of golden setae. Distance between upper eye lobes 0.48 times length of scape; in frontal view, distance between lower eye lobes equal to length of scape. Antennae 1.8 times elytral length, reaching elytral apex at base of antennomere IX. Scape with dense yellowish-brown pubescence dorsally, gradually shorter, sparser toward ventral surface; with long, sparse, erect whitish and dark setae dorsally, whitish ventrally. Pedicel and antennomeres with yellowish-white pubescence partially obscuring integument (more yellowish-brown on basal antennomeres), with long, erect dark setae interspersed throughout, longer ventrally (basal antennomeres with a few white setae interspersed). Antennal formula (ratio) based on length of antennomere III: scape = 0.86; pedicel = 0.27; IV = 1.04; V = 0.90; VI = 0.86; VII = 0.81; VIII = 0.72; IX = 0.67; X = 0.58; XI = 0.65.
</paragraph>
<paragraph id="8BBE36EC096E0B68FF75FC9AFB9472F3" blockId="8.[151,1437,152,2015]" pageId="8" pageNumber="100">
<emphasis id="B975EAFE096E0B68FF75FC9AFEA877CE" bold="true" box="[199,288,873,899]" pageId="8" pageNumber="100">Thorax</emphasis>
. Prothorax almost as long as wide; sides sinuous, forming two rounded protuberances, one anteriorly, another about middle. Pronotum with one moderately large tubercle on each side of anterior half, another less elevated centrally on posterior half; coarsely, abundantly punctate; with two longitudinal yellowish-white pubescent centrally, one at anterior quarter, another at posterior half; remaining surface with yellowish-brown pubescence, not obscuring integument, more yellowish-white laterally on some areas; with long, sparse, erect white setae. Sides of prothorax coarsely, abundantly punctate; with yellowish-brown pubescence partially obscuring integument, with long, erect white setae interspersed. Prosternum and prosternal process coarsely, abundantly punctate; with yellowish-white pubescence not obscuring integument (whiter depending on light intensity), with long, erect white setae interspersed. Ventral surface of mesothorax with yellowish pubescence not obscuring integument (whiter depending on light intensity), distinctly shorter and sparser on central area of mesoventrite and mesoventral process, with long, erect white setae interspersed; mesoventral process, mesanepisternum and mesepimeron coarsely, moderately abundantly punctate; central area of mesoventrite coarsely, shallowly, sparsely punctate. Ventral surface of metathorax coarsely, moderately abundantly punctate laterally, punctures slightly finer and sparser on anterocentral area, distinctly finer and sparser on posterocentral area; with yellowish pubescence not obscuring integument (whiter depending on light intensity), with long, erect white setae interspersed. Scutellum with yellowish-brown pubescence not obscuring integument laterally, lighter, slightly denser along longitudinal central area.
<emphasis id="B975EAFE096E0B68FB34FA76FB5A71D2" bold="true" box="[1158,1234,1413,1439]" pageId="8" pageNumber="100">Elytra</emphasis>
. Parallel-sided on anterior 3/4, gradually narrowed toward widely truncate apex posteriorly; coarsely, moderately abundantly punctate on basal third, punctures gradually finer, sparser toward apex; with oblique, dense white pubescent band centrally, not reaching suture and epipleural margin; with another oblique white pubescent band in posterior third, wider than the former toward suture, with irregular maculae with yellowish pubescence interspersed dorsally; with two more white pubescent bands near apex, irregular and less conspicuous; remaining surface with light yellowish-brown pubescence interspersed with yellowish-white pubescence, not obscuring integument; with long, sparse, erect dark setae dorsally (and a few white setae), white laterally.
<emphasis id="B975EAFE096E0B68FD5FF972FCAD72D6" bold="true" box="[749,805,1665,1691]" pageId="8" pageNumber="100">Legs</emphasis>
. Femora and tibiae with abundant light yellowish-brown pubescence, not obscuring integument, with long, erect white setae interspersed.
</paragraph>
<paragraph id="8BBE36EC096E0B68FF75F93BFBFD734A" blockId="8.[151,1437,152,2015]" pageId="8" pageNumber="100">
<emphasis id="B975EAFE096E0B68FF75F93BFEB472AE" bold="true" box="[199,316,1736,1763]" pageId="8" pageNumber="100">Abdomen</emphasis>
. Ventrites with abundant yellowish-white pubescence not obscuring integument (whiter depending on light intensity), with long, erect white setae interspersed. Apex of ventrite V truncate.
</paragraph>
<paragraph id="8BBE36EC096E0B68FF75F8E3FC5073DA" blockId="8.[151,1437,152,2015]" pageId="8" pageNumber="100">
<emphasis id="B975EAFE096E0B68FF75F8E3FE127366" bold="true" box="[199,410,1808,1835]" pageId="8" pageNumber="100">Dimensions (mm)</emphasis>
.
<materialsCitation id="3B693CB1096E0B68FE15F8E3FC5073DA" ID-GBIF-Occurrence="2417946728" collectingDate="1981-06-02" collectingDateMax="1981-06-15" collectingDateMin="1981-06-02" collectionCode="LGBC, CASC" collectorName="Hanson &amp; Clemons" country="Trinidad and Tobago" county="Simla" location="Total" municipality="Trinidad" pageId="8" pageNumber="100" specimenCount="1" typeStatus="holotype">
<location id="8EDE6037096E0B68FE15F8E3FE687367" LSID="urn:lsid:plazi:treatment:03A887FA09610B6AFF25F89DFF3476E6:8EDE6037096E0B68FE15F8E3FE687367" box="[423,480,1808,1834]" country="Trinidad and Tobago" county="Simla" municipality="Trinidad" name="Total" pageId="8" pageNumber="100">Total</location>
length, 6.65; prothoracic length, 1.35; anterior prothoracic width, 1.15; posterior prothoracic width, 1.10; maximum prothoracic width, 1.35; humeral width, 1.70; elytral length, 4.60.
<emphasis id="B975EAFE096E0B68FB5FF8C6FA1D7302" bold="true" box="[1261,1429,1844,1871]" pageId="8" pageNumber="100">Type material</emphasis>
.
<typeStatus id="54BA884E096E0B68FF25F8ABFF77733F" box="[151,255,1880,1906]" pageId="8" pageNumber="100" type="holotype">Holotype</typeStatus>
female from
<collectingCountry id="F316767C096E0B68FE29F8AAFD6D733E" box="[411,741,1881,1907]" name="Trinidad and Tobago" pageId="8" pageNumber="100">TRINIDAD AND TOBAGO</collectingCountry>
,
<collectingMunicipality id="6BDAAC96096E0B68FD40F8ABFCDA733F" box="[754,850,1880,1906]" pageId="8" pageNumber="100">
<emphasis id="B975EAFE096E0B68FD40F8ABFCDA733F" box="[754,850,1880,1906]" italics="true" pageId="8" pageNumber="100">Trinidad</emphasis>
</collectingMunicipality>
:
<collectingCounty id="62DF4E60096E0B68FCD1F8ABFC2C733F" box="[867,932,1880,1906]" pageId="8" pageNumber="100">Simla</collectingCounty>
[William Beebe Tropical Research Station],
<collectingDate id="EFFBE9C4096E0B68FF25F88FFEB873DA" box="[151,304,1916,1943]" pageId="8" pageNumber="100" value="1981-06-02" valueMax="1981-06-15" valueMin="1981-06-02">
<date id="FFBF102C096E0B68FF25F88FFEB873DA" box="[151,304,1916,1943]" pageId="8" pageNumber="100" value="1981-06-02" valueMax="1981-06-15" valueMin="1981-06-02">2-15.VI.1981</date>
</collectingDate>
,
<collectorName id="26F4533A096E0B68FE8EF88EFE1A73DA" box="[316,402,1917,1943]" pageId="8" pageNumber="100">Hanson</collectorName>
&amp;
<collectorName id="26F4533A096E0B68FE07F88FFD9173DB" box="[437,537,1916,1942]" pageId="8" pageNumber="100">Clemons</collectorName>
col. (
<collectionCode id="ED10AE29096E0B68FDEAF88EFD2F73DA" box="[600,679,1917,1943]" pageId="8" pageNumber="100">LGBC</collectionCode>
, to be deposited at
<collectionCode id="ED10AE29096E0B68FCCCF88EFC4473DA" box="[894,972,1917,1943]" pageId="8" pageNumber="100">CASC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BBE36EC096E0B6AFF75F852FE5F762E" blockId="8.[151,1437,152,2015]" lastBlockId="10.[151,1437,153,683]" lastPageId="10" lastPageNumber="102" pageId="8" pageNumber="100">
<emphasis id="B975EAFE096E0B68FF75F852FEBC73F6" bold="true" box="[199,308,1953,1979]" pageId="8" pageNumber="100">Remarks</emphasis>
.
<taxonomicName id="4C014D6F096E0B68FEF7F853FDDE73F7" authority="Bezark &amp; Santos-Silva, 2019" authorityName="Bezark &amp; Santos-Silva" authorityYear="2019" box="[325,598,1952,1978]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="100" phylum="Arthropoda" rank="species" species="alboangulata" status="sp. nov.">
<emphasis id="B975EAFE096E0B68FEF7F853FDDE73F7" box="[325,598,1952,1978]" italics="true" pageId="8" pageNumber="100">Tethystola alboangulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2465785096E0B68FDD3F852FD3573F6" box="[609,701,1953,1979]" pageId="8" pageNumber="100" rank="species">
<emphasis id="B975EAFE096E0B68FDD3F852FD3573F6" bold="true" box="[609,701,1953,1979]" pageId="8" pageNumber="100">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the group of species with a truncate elytral apex. It differs from
<taxonomicName id="4C014D6F096E0B68FE94F836FDF37392" authority="Breuning, 1940" authorityName="Breuning" authorityYear="1940" box="[294,635,1988,2015]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="100" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B975EAFE096E0B68FE94F836FE4C7393" box="[294,452,1988,2014]" italics="true" pageId="8" pageNumber="100">T. brasiliensis</emphasis>
Breuning, 1940
</taxonomicName>
(
<figureCitation id="133A2A69096E0B68FD3EF837FC9E7392" box="[652,790,1988,2015]" captionStart="FIGURES 19–27. 19–22" captionStartId="9.[151,264,1895,1920]" captionTargetBox="[208,1386,168,1876]" captionTargetId="figure@9.[195,1392,166,1876]" captionTargetPageId="9" captionText="FIGURES 19–27. 19–22) Tethystola alboangulata, holotype female: 19) Dorsal habitus; 20) Ventral habitus; 21) Lateral habitus; 22) Head, frontal view. 23) Tethystola flavoapicalis Breuning, 1942 (= T. brasiliensis), dorsal habitus, holotype. 24) Tethystola brasiliensis, dorsal habitus, holotype. 25–26) Tethystola minima, dorsal habitus: 25) Holotype male; 26) Male from Venezuela. 27) Tethystola inermis, dorsal habitus, male from Venezuela." httpUri="https://zenodo.org/record/3353935/files/figure.png" pageId="8" pageNumber="100">Figs. 23–24</figureCitation>
), by the antennomeres IX–X or IX–XI being not entirely yellowish (yellowish in
<taxonomicName id="4C014D6F096C0B6AFE10FF69FDB674FE" authorityName="Breuning" authorityYear="1940" box="[418,574,153,179]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B975EAFE096C0B6AFE10FF69FDB674FE" box="[418,574,153,179]" italics="true" pageId="10" pageNumber="102">T. brasiliensis</emphasis>
</taxonomicName>
), the white pubescent band on the central area of the elytra is less oblique and not followed by a dark area (more oblique and followed by dark area in
<taxonomicName id="4C014D6F096C0B6AFC62FF4DFBE6749A" authorityName="Breuning" authorityYear="1940" box="[976,1134,189,215]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B975EAFE096C0B6AFC62FF4DFBE6749A" box="[976,1134,189,215]" italics="true" pageId="10" pageNumber="102">T. brasiliensis</emphasis>
</taxonomicName>
), and the posterior area of the elytra without a dark area (present in
<taxonomicName id="4C014D6F096C0B6AFDDBFF11FC8F74B6" authorityName="Breuning" authorityYear="1940" box="[617,775,225,251]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B975EAFE096C0B6AFDDBFF11FC8F74B6" box="[617,775,225,251]" italics="true" pageId="10" pageNumber="102">T. brasiliensis</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="4C014D6F096C0B6AFC74FF11FADC74B6" authority="Galileo &amp; Martins, 2001" authorityName="Galileo &amp; Martins" authorityYear="2001" box="[966,1364,225,251]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B975EAFE096C0B6AFC74FF11FBBF74B6" box="[966,1079,226,251]" italics="true" pageId="10" pageNumber="102">T. minima</emphasis>
Galileo &amp; Martins, 2001
</taxonomicName>
(
<figureCitation id="133A2A69096C0B6AFAD6FF12FF6A7552" captionStart="FIGURES 19–27. 19–22" captionStartId="9.[151,264,1895,1920]" captionTargetBox="[208,1386,168,1876]" captionTargetId="figure@9.[195,1392,166,1876]" captionTargetPageId="9" captionText="FIGURES 19–27. 19–22) Tethystola alboangulata, holotype female: 19) Dorsal habitus; 20) Ventral habitus; 21) Lateral habitus; 22) Head, frontal view. 23) Tethystola flavoapicalis Breuning, 1942 (= T. brasiliensis), dorsal habitus, holotype. 24) Tethystola brasiliensis, dorsal habitus, holotype. 25–26) Tethystola minima, dorsal habitus: 25) Holotype male; 26) Male from Venezuela. 27) Tethystola inermis, dorsal habitus, male from Venezuela." httpUri="https://zenodo.org/record/3353935/files/figure.png" pageId="10" pageNumber="102">Figs. 25–26</figureCitation>
), by the stouter body (more slender in
<taxonomicName id="4C014D6F096C0B6AFD12FEF5FC997552" authorityName="Galileo &amp; Martins" authorityYear="2001" box="[672,785,262,287]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B975EAFE096C0B6AFD12FEF5FC997552" box="[672,785,262,287]" italics="true" pageId="10" pageNumber="102">T. minima</emphasis>
</taxonomicName>
), the antennae mostly dark (mostly orangish-brown in
<taxonomicName id="4C014D6F096C0B6AFA3AFEF5FF63750E" authorityName="Galileo &amp; Martins" authorityYear="2001" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B975EAFE096C0B6AFA3AFEF5FF63750E" italics="true" pageId="10" pageNumber="102">T. minima</emphasis>
</taxonomicName>
), and different pubescent patterns on the pronotum and elytra.
<taxonomicName id="4C014D6F096C0B6AFC18FEDAFB30750E" authorityName="Bezark &amp; Santos-Silva" authorityYear="2019" box="[938,1208,297,323]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="alboangulata">
<emphasis id="B975EAFE096C0B6AFC18FEDAFB30750E" box="[938,1208,297,323]" italics="true" pageId="10" pageNumber="102">Tethystola alboangulata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C014D6F096C0B6AFAF9FED9FE56752A" authority="Galileo &amp; Martins, 2001" authorityName="Galileo &amp; Martins" authorityYear="2001" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EAFE096C0B6AFAF9FED9FF36752A" italics="true" pageId="10" pageNumber="102">T. inermis</emphasis>
Galileo &amp; Martins, 2001
</taxonomicName>
(
<figureCitation id="133A2A69096C0B6AFE5DFEBEFDCC752A" box="[495,580,333,359]" captionStart="FIGURES 19–27. 19–22" captionStartId="9.[151,264,1895,1920]" captionTargetBox="[208,1386,168,1876]" captionTargetId="figure@9.[195,1392,166,1876]" captionTargetPageId="9" captionText="FIGURES 19–27. 19–22) Tethystola alboangulata, holotype female: 19) Dorsal habitus; 20) Ventral habitus; 21) Lateral habitus; 22) Head, frontal view. 23) Tethystola flavoapicalis Breuning, 1942 (= T. brasiliensis), dorsal habitus, holotype. 24) Tethystola brasiliensis, dorsal habitus, holotype. 25–26) Tethystola minima, dorsal habitus: 25) Holotype male; 26) Male from Venezuela. 27) Tethystola inermis, dorsal habitus, male from Venezuela." httpUri="https://zenodo.org/record/3353935/files/figure.png" pageId="10" pageNumber="102">Fig. 27</figureCitation>
), especially by the noticeably different pubescent patterns on the pronotum and elytra, and darker legs (orangish-brown in
<taxonomicName id="4C014D6F096C0B6AFD1FFE81FC9575C6" authorityName="Galileo &amp; Martins" authorityYear="2001" box="[685,797,370,395]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EAFE096C0B6AFD1FFE81FC9575C6" box="[685,797,370,395]" italics="true" pageId="10" pageNumber="102">T. inermis</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="4C014D6F096C0B6AFC6CFE81FB5775C6" authority="Gahan, 1895" authorityName="Gahan" authorityYear="1895" box="[990,1247,368,395]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="mutica">
<emphasis id="B975EAFE096C0B6AFC6CFE81FBCF75C6" box="[990,1095,370,395]" italics="true" pageId="10" pageNumber="102">T. mutica</emphasis>
Gahan, 1895
</taxonomicName>
(see photograph of the
<typeStatus id="54BA884E096C0B6AFF50FE66FEC175E2" box="[226,329,405,431]" pageId="10" pageNumber="102" type="lectotype">lectotype</typeStatus>
female at
<bibRefCitation id="EF904B1D096C0B6AFE70FE66FDDE75E2" author="Bezark, L. G." box="[450,598,405,431]" pageId="10" pageNumber="102" refId="ref11405" refString="Bezark, L. G. (2019) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: https: // apps 2. cdfa. ca. gov / publicApps / plant / bycidDB / wsearch. asp? w = n (accessed 15 March 2019)" type="url" year="2019">Bezark 2019</bibRefCitation>
), by the oblique pubescent band of the central area of the elytra not sinuous (sinuous in
<taxonomicName id="4C014D6F096C0B6AFEFAFE49FE27759E" authorityName="Gahan" authorityYear="1895" box="[328,431,442,467]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="mutica">
<emphasis id="B975EAFE096C0B6AFEFAFE49FE27759E" box="[328,431,442,467]" italics="true" pageId="10" pageNumber="102">T. mutica</emphasis>
</taxonomicName>
), and the posterior oblique band distinctly wider and irregular (narrower and regular in
<taxonomicName id="4C014D6F096C0B6AFA3AFE49FF6B75BA" authorityName="Gahan" authorityYear="1895" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="mutica">
<emphasis id="B975EAFE096C0B6AFA3AFE49FF6B75BA" italics="true" pageId="10" pageNumber="102">T. mutica</emphasis>
</taxonomicName>
). Finally, it differs from
<taxonomicName id="4C014D6F096C0B6AFE4BFE2DFCFA75BA" authority="Martins &amp; Galileo, 2008" authorityName="Martins &amp; Galileo" authorityYear="2008" box="[505,882,477,503]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B975EAFE096C0B6AFE4BFE2DFDD075BA" box="[505,600,478,503]" italics="true" pageId="10" pageNumber="102">T. cincta</emphasis>
Martins &amp; Galileo, 2008
</taxonomicName>
(see photograph of the
<typeStatus id="54BA884E096C0B6AFBC8FE2EFB5475BA" box="[1146,1244,477,503]" pageId="10" pageNumber="102" type="holotype">holotype</typeStatus>
at
<bibRefCitation id="EF904B1D096C0B6AFB4CFE2EFA1875BA" author="Bezark, L. G." box="[1278,1424,477,503]" pageId="10" pageNumber="102" refId="ref11405" refString="Bezark, L. G. (2019) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: https: // apps 2. cdfa. ca. gov / publicApps / plant / bycidDB / wsearch. asp? w = n (accessed 15 March 2019)" type="url" year="2019">Bezark 2019</bibRefCitation>
), by the stouter body (slenderer in
<taxonomicName id="4C014D6F096C0B6AFDB7FDF1FDEC7656" authorityName="Martins &amp; Galileo" authorityYear="2008" box="[517,612,514,539]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B975EAFE096C0B6AFDB7FDF1FDEC7656" box="[517,612,514,539]" italics="true" pageId="10" pageNumber="102">T. cincta</emphasis>
</taxonomicName>
), elytra being proportionally shorter (3.4 times pronotal length) (longer in
<taxonomicName id="4C014D6F096C0B6AFF25FDD5FF7F7672" authorityName="Martins &amp; Galileo" authorityYear="2008" box="[151,247,550,575]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B975EAFE096C0B6AFF25FDD5FF7F7672" box="[151,247,550,575]" italics="true" pageId="10" pageNumber="102">T. cincta</emphasis>
</taxonomicName>
, about 3.7 times longer than pronotum), and the elytral apex is not distinctly oblique (oblique, with outer angle projected in
<taxonomicName id="4C014D6F096C0B6AFED5FDB9FE4E762E" authorityName="Martins &amp; Galileo" authorityYear="2008" box="[359,454,586,611]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B975EAFE096C0B6AFED5FDB9FE4E762E" box="[359,454,586,611]" italics="true" pageId="10" pageNumber="102">T. cincta</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF7E6664096F0B69FF25F894FCFB73A1" httpUri="https://zenodo.org/record/3353935/files/figure.png" pageId="9" pageNumber="101" startId="9.[151,264,1895,1920]" targetBox="[208,1386,168,1876]" targetPageId="9">
<paragraph id="8BBE36EC096F0B69FF25F894FCFB73A1" blockId="9.[151,1436,1895,2028]" pageId="9" pageNumber="101">
<emphasis id="B975EAFE096F0B69FF25F894FEDA73CD" bold="true" box="[151,338,1895,1920]" pageId="9" pageNumber="101">FIGURES 19–27</emphasis>
.
<emphasis id="B975EAFE096F0B69FED1F894FE2273CD" bold="true" box="[355,426,1895,1920]" pageId="9" pageNumber="101">19–22)</emphasis>
<taxonomicName id="4C014D6F096F0B69FE07F89BFD2673CD" authorityName="Bezark &amp; Santos-Silva" authorityYear="2019" box="[437,686,1896,1920]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="101" phylum="Arthropoda" rank="species" species="alboangulata">
<emphasis id="B975EAFE096F0B69FE07F89BFD2673CD" box="[437,686,1896,1920]" italics="true" pageId="9" pageNumber="101">Tethystola alboangulata</emphasis>
</taxonomicName>
, holotype female:
<emphasis id="B975EAFE096F0B69FCCAF894FC1373CD" bold="true" box="[888,923,1895,1920]" pageId="9" pageNumber="101">19)</emphasis>
Dorsal habitus;
<emphasis id="B975EAFE096F0B69FBFFF894FBE773CD" bold="true" box="[1101,1135,1895,1920]" pageId="9" pageNumber="101">20)</emphasis>
Ventral habitus;
<emphasis id="B975EAFE096F0B69FA9AF894FAC273CD" bold="true" box="[1320,1354,1895,1920]" pageId="9" pageNumber="101">21)</emphasis>
Lateral habitus;
<emphasis id="B975EAFE096F0B69FF5DF878FE9973E9" bold="true" box="[239,273,1931,1956]" pageId="9" pageNumber="101">22)</emphasis>
Head, frontal view.
<emphasis id="B975EAFE096F0B69FE54F878FD8F73E9" bold="true" box="[486,519,1931,1956]" pageId="9" pageNumber="101">23)</emphasis>
<taxonomicName id="4C014D6F096F0B69FDBDF87FFC2173E9" authority="Breuning, 1942" authorityName="Breuning" authorityYear="1942" box="[527,937,1931,1956]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="101" phylum="Arthropoda" rank="species" species="flavoapicalis">
<emphasis id="B975EAFE096F0B69FDBDF87FFC8973E9" box="[527,769,1932,1956]" italics="true" pageId="9" pageNumber="101">Tethystola flavoapicalis</emphasis>
Breuning, 1942
</taxonomicName>
(=
<taxonomicName id="4C014D6F096F0B69FC7DF87EFBE873E9" authorityName="Breuning" authorityYear="1940" box="[975,1120,1932,1956]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="101" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B975EAFE096F0B69FC7DF87EFC5573E9" box="[975,989,1933,1956]" italics="true" pageId="9" pageNumber="101">T</emphasis>
.
<emphasis id="B975EAFE096F0B69FC59F87FFBE873E9" box="[1003,1120,1932,1956]" italics="true" pageId="9" pageNumber="101">brasiliensis</emphasis>
</taxonomicName>
), dorsal habitus, holotype.
<emphasis id="B975EAFE096F0B69FAC8F878FA1473E9" bold="true" box="[1402,1436,1931,1956]" pageId="9" pageNumber="101">24)</emphasis>
<taxonomicName id="4C014D6F096F0B69FF25F843FEF27385" authorityName="Breuning" authorityYear="1940" box="[151,378,1968,1992]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="101" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B975EAFE096F0B69FF25F843FEF27385" box="[151,378,1968,1992]" italics="true" pageId="9" pageNumber="101">Tethystola brasiliensis</emphasis>
</taxonomicName>
, dorsal habitus, holotype.
<emphasis id="B975EAFE096F0B69FD3AF85CFD587385" bold="true" box="[648,720,1967,1992]" pageId="9" pageNumber="101">25–26)</emphasis>
<taxonomicName id="4C014D6F096F0B69FD65F843FC197385" authorityName="Galileo &amp; Martins" authorityYear="2001" box="[727,913,1968,1992]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="101" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B975EAFE096F0B69FD65F843FC197385" box="[727,913,1968,1992]" italics="true" pageId="9" pageNumber="101">Tethystola minima</emphasis>
</taxonomicName>
, dorsal habitus:
<emphasis id="B975EAFE096F0B69FB88F85CFBD47385" bold="true" box="[1082,1116,1967,1992]" pageId="9" pageNumber="101">25)</emphasis>
Holotype male;
<emphasis id="B975EAFE096F0B69FAB5F85CFAA17385" bold="true" box="[1287,1321,1967,1992]" pageId="9" pageNumber="101">26)</emphasis>
Male from Venezuela.
<emphasis id="B975EAFE096F0B69FEBEF820FEA573A1" bold="true" box="[268,301,2003,2028]" pageId="9" pageNumber="101">27)</emphasis>
<taxonomicName id="4C014D6F096F0B69FE86F827FE6673A1" authorityName="Galileo &amp; Martins" authorityYear="2001" box="[308,494,2004,2028]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="101" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EAFE096F0B69FE86F827FE6673A1" box="[308,494,2004,2028]" italics="true" pageId="9" pageNumber="101">Tethystola inermis</emphasis>
</taxonomicName>
, dorsal habitus, male from Venezuela.
</paragraph>
</caption>
<paragraph id="8BBE36EC096C0B6AFF75FD9EFF3476E6" blockId="10.[151,1437,153,683]" pageId="10" pageNumber="102">
<emphasis id="B975EAFE096C0B6AFF75FD9EFEC076CA" bold="true" box="[199,328,621,647]" pageId="10" pageNumber="102">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C014D6F096C0B6AFD88FD9EFD4776CA" authorityName="Bezark &amp; Santos-Silva" authorityYear="2019" box="[570,719,621,647]" class="Insecta" family="Cerambycidae" genus="Tethystola" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="102" phylum="Arthropoda" rank="species" species="alboangulata">
<emphasis id="B975EAFE096C0B6AFD88FD9EFD4776CA" box="[570,719,621,647]" italics="true" pageId="10" pageNumber="102">alboangulata</emphasis>
</taxonomicName>
refers to the two angulate white lines of pubescence on the elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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