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<document id="5BD95191DE163E168717B12656640F24" ID-DOI="10.1649/0010-065X-75.2.279" ID-ISSN="1938-4394" ID-Zenodo-Dep="13251790" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723001616113" checkinUser="felipe" docAuthor="Ratcliffe, Brett C., Cave, Ronald D. &amp; Mondaca, José" docDate="2021" docId="03A887D8FFB07D5C6E63FA06068740D3" docLanguage="en" docName="TheColeopBull.75.2.279-309.pdf" docOrigin="The Coleopterists Bulletin 75 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-75.2.279" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Golofa inermis Thomson 1859" docType="treatment" docVersion="2" lastPageNumber="298" masterDocId="FF91FFA0FFA27D486D66FF9A06674147" masterDocTitle="The Dynastine Scarab Beetles (Coleoptera: Scarabaeidae: Dynastinae) of Chile" masterLastPageNumber="309" masterPageNumber="279" pageNumber="296" updateTime="1723003482374" updateUser="ExternalLinkService">
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<mods:title id="F38ABD753627ABC6A37E6594BEFEBFFB">The Dynastine Scarab Beetles (Coleoptera: Scarabaeidae: Dynastinae) of Chile</mods:title>
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<mods:namePart id="CE0C24816E9875045B2B091F8FDB9778">Ratcliffe, Brett C.</mods:namePart>
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<mods:namePart id="6ACA63325A2C1FDD7AFE706EA1BE0605">Cave, Ronald D.</mods:namePart>
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<mods:namePart id="34FC7EA88D678A1AEFE215F483C84534">Mondaca, José</mods:namePart>
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<mods:title id="001BDFB57408FB42C2A5B2DF61F016CD">The Coleopterists Bulletin</mods:title>
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<mods:date id="D06ADADC2DE750837D97D9B291BC852E">2021</mods:date>
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<mods:classification id="70F22BC96F6B5738548F50FAE9A85A11">journal article</mods:classification>
<mods:identifier id="CF84C82C184B80578BD6E29B7586F38B" type="DOI">10.1649/0010-065X-75.2.279</mods:identifier>
<mods:identifier id="42635508F8044304EC9A85DF2A098606" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03A887D8FFB07D5C6E63FA06068740D3" LSID="urn:lsid:plazi:treatment:03A887D8FFB07D5C6E63FA06068740D3" httpUri="http://treatment.plazi.org/id/03A887D8FFB07D5C6E63FA06068740D3" lastPageId="20" lastPageNumber="298" pageId="18" pageNumber="296">
<subSubSection id="C31B6545FFB07D5A6E63FA06025C44F6" box="[773,1083,1436,1458]" pageId="18" pageNumber="296" type="nomenclature">
<paragraph id="8BBE36CEFFB07D5A6E63FA06025C44F6" blockId="18.[773,1083,1436,1485]" box="[773,1083,1436,1458]" pageId="18" pageNumber="296">
<heading id="D0F681A2FFB07D5A6E63FA06025C44F6" box="[773,1083,1436,1458]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="296" reason="2">
<taxonomicName id="4C014D4DFFB07D5A6E63FA06025C44F6" ID-CoL="3GRWJ" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" box="[773,1083,1436,1458]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="296" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB07D5A6E63FA06025C44F6" bold="true" box="[773,1083,1436,1458]" pageId="18" pageNumber="296">
<emphasis id="B975EADCFFB07D5A6E63FA0605FD44F6" bold="true" box="[773,922,1436,1458]" italics="true" pageId="18" pageNumber="296">Golofa inermis</emphasis>
<bibRefCitation id="EF904B3FFFB07D5A6EF9FA06025C44F6" author="Thomson, J." box="[927,1083,1436,1457]" pageId="18" pageNumber="296" pagination="3 - 22" refId="ref18285" refString="Thomson, J. 1859. Essai synoptique sur la sous-tribu des scarabaeitae vrais. Arcana Naturae 1: 3 - 22." type="journal article" year="1859">Thomson, 1859</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B6545FFB07D5A6E39FA2D0586448B" box="[863,993,1463,1485]" pageId="18" pageNumber="296" type="description">
<paragraph id="8BBE36CEFFB07D5A6E39FA2D0586448B" blockId="18.[773,1083,1436,1485]" box="[863,993,1463,1485]" pageId="18" pageNumber="296">
(
<figureCitation id="133A2A4BFFB07D5A6E00FA2D05BD448B" box="[870,986,1463,1485]" captionStart="Figs" captionStartId="19.[198,237,1108,1127]" captionTargetBox="[175,1187,193,1082]" captionTargetId="figure-285@19.[173,1187,193,1082]" captionTargetPageId="19" captionText="Figs. 3437. Golofa inermis. 34) Lectotype, MNHN-EC-2378; 35) Lectotype labels; 36) Parameres; 37) Distribution. Images in Figs. 3435 courtesy of Antoine Mantilleri (Muséum National dHistoire Naturelle, Paris, France)." figureDoi="http://doi.org/10.5281/zenodo.13251827" httpUri="https://zenodo.org/record/13251827/files/figure.png" pageId="18" pageNumber="296">Figs. 3437</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B6545FFB07D5A6FCBFA78052A4755" pageId="18" pageNumber="296" type="reference_group">
<paragraph id="8BBE36CEFFB07D5A6FCBFA78052A4755" blockId="18.[685,1170,1505,1554]" pageId="18" pageNumber="296">
<treatmentCitationGroup id="AB1111E0FFB07D5A6FCBFA78052A4755" pageId="18" pageNumber="296">
<treatmentCitation id="0AA010DFFFB07D5A6FCBFA78024A44B0" author="Thomson, J." box="[685,1069,1505,1528]" page="11" pageId="18" pageNumber="296" year="1859">
<taxonomicName id="4C014D4DFFB07D5A6FCBFA78024A44B0" ID-CoL="3GRWJ" authority="Thomson 1859: 11" authorityName="Thomson" authorityPageNumber="11" authorityYear="1859" box="[685,1069,1505,1528]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="296" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB07D5A6FCBFA78052C44B0" box="[685,843,1506,1527]" italics="true" pageId="18" pageNumber="296">Golofa inermis</emphasis>
<bibRefCitation id="EF904B3FFFB07D5A6E38FA78024A44B0" author="Thomson, J." box="[862,1069,1505,1528]" pageId="18" pageNumber="296" pagination="3 - 22" refId="ref18285" refString="Thomson, J. 1859. Essai synoptique sur la sous-tribu des scarabaeitae vrais. Arcana Naturae 1: 3 - 22." type="journal article" year="1859">Thomson 1859: 11</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31B6545FFB07D5B6FA3F9BD04794763" lastPageId="19" lastPageNumber="297" pageId="18" pageNumber="296" type="description">
<paragraph id="8BBE36CEFFB07D5B6FA3F9BD04794763" blockId="18.[684,1171,1575,1730]" lastBlockId="19.[172,660,1230,1732]" lastPageId="19" lastPageNumber="297" pageId="18" pageNumber="296">
<emphasis id="B975EADCFFB07D5A6FA3F9BD053D477A" bold="true" box="[709,858,1575,1597]" pageId="18" pageNumber="296">Redescription.</emphasis>
Length 20.025.0 mm; width 8.613.0 mm. Color uniformly dark yellowish brown or reddish brown except for black head, scutellum, sutural striae, tibiae, tarsi, pygidium, and venter.
<emphasis id="B975EADCFFB07D5A6F93F908055447E0" bold="true" box="[757,819,1682,1703]" pageId="18" pageNumber="296">Head:</emphasis>
Frons with moderately large, sparse punctures. Clypeus with erect, sharply acuminate tubercle; apex strongly narrowed, emarginate. Interocular width equals 2.7 transverse eye diameters. Antenna with 10 antennomeres, club slightly longer than antennomeres 27.
<emphasis id="B975EADCFFB17D5B6CB1FA8404214474" bold="true" box="[471,582,1310,1331]" pageId="19" pageNumber="297">Pronotum:</emphasis>
Surface with punctures moderate in size and density.A shallow, short, longitudinal sulcus present just behind anterior margin. Base with marginal bead.
<emphasis id="B975EADCFFB17D5B6F2FFAF404F544C4" bold="true" box="[585,658,1390,1411]" pageId="19" pageNumber="297">Elytra:</emphasis>
Surface with punctures moderate in size and density, shallow. Sutural stria a completely impressed line.
<emphasis id="B975EADCFFB17D5B6DC8FA2407704493" bold="true" box="[174,279,1470,1492]" pageId="19" pageNumber="297">Pygidium:</emphasis>
Surface with punctures moderate in size and density. Propygidium lacking long, dense setae.
<emphasis id="B975EADCFFB17D5B6DCAFA6E0683474E" bold="true" box="[172,228,1524,1545]" pageId="19" pageNumber="297">Legs:</emphasis>
Protibia tridentate, basal tooth removed from other teeth.
<emphasis id="B975EADCFFB17D5B6C79F99407F24764" bold="true" box="[287,405,1550,1571]" pageId="19" pageNumber="297">Parameres:</emphasis>
As in
<figureCitation id="133A2A4BFFB17D5B6CB5F994047D4763" box="[467,538,1550,1572]" captionStart="Figs" captionStartId="19.[198,237,1108,1127]" captionTargetBox="[175,1187,193,1082]" captionTargetId="figure-285@19.[173,1187,193,1082]" captionTargetPageId="19" captionText="Figs. 3437. Golofa inermis. 34) Lectotype, MNHN-EC-2378; 35) Lectotype labels; 36) Parameres; 37) Distribution. Images in Figs. 3435 courtesy of Antoine Mantilleri (Muséum National dHistoire Naturelle, Paris, France)." figureDoi="http://doi.org/10.5281/zenodo.13251827" httpUri="https://zenodo.org/record/13251827/files/figure.png" pageId="19" pageNumber="297">Fig. 36</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF7E6646FFB17D5B6DA0FBCE074C45DF" ID-DOI="http://doi.org/10.5281/zenodo.13251827" ID-Zenodo-Dep="13251827" httpUri="https://zenodo.org/record/13251827/files/figure.png" pageId="19" pageNumber="297" startId="19.[198,237,1108,1127]" targetBox="[175,1187,193,1082]" targetPageId="19" targetType="figure">
<paragraph id="8BBE36CEFFB17D5B6DA0FBCE074C45DF" blockId="19.[173,1193,1108,1176]" pageId="19" pageNumber="297">
<emphasis id="B975EADCFFB17D5B6DA0FBCE0759452F" bold="true" box="[198,318,1108,1128]" pageId="19" pageNumber="297">Figs. 3437.</emphasis>
<taxonomicName id="4C014D4DFFB17D5B6C35FBCF07874520" authorityName="Thomson" authorityYear="1859" box="[339,480,1109,1128]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="297" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB17D5B6C35FBCF07874520" box="[339,480,1109,1128]" italics="true" pageId="19" pageNumber="297">Golofa inermis</emphasis>
</taxonomicName>
.
<emphasis id="B975EADCFFB17D5B6C90FBCE04754520" bold="true" box="[502,530,1108,1127]" pageId="19" pageNumber="297">34)</emphasis>
Lectotype, MNHN-EC-2378;
<emphasis id="B975EADCFFB17D5B6E5BFBCE053E4520" bold="true" box="[829,857,1108,1127]" pageId="19" pageNumber="297">35)</emphasis>
Lectotype labels;
<emphasis id="B975EADCFFB17D5B697DFBCE025F4520" bold="true" box="[1051,1080,1108,1127]" pageId="19" pageNumber="297">36)</emphasis>
Parameres;
<emphasis id="B975EADCFFB17D5B6DCBFBF606AD4538" bold="true" box="[173,202,1132,1151]" pageId="19" pageNumber="297">37)</emphasis>
Distribution. Images in Figs. 3435 courtesy of Antoine Mantilleri (Muséum National dHistoire Naturelle, Paris, France).
</paragraph>
</caption>
<subSubSection id="C31B6545FFB17D5B6DA3F9B302474473" pageId="19" pageNumber="297" type="distribution">
<paragraph id="8BBE36CEFFB17D5B6DA3F9B304494734" blockId="19.[172,660,1230,1732]" pageId="19" pageNumber="297">
<emphasis id="B975EADCFFB17D5B6DA3F9B307214779" bold="true" box="[197,326,1577,1599]" pageId="19" pageNumber="297">Distribution.</emphasis>
<taxonomicName id="4C014D4DFFB17D5B6C2DF9B307BF4778" authorityName="Thomson" authorityYear="1859" box="[331,472,1577,1599]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="297" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB17D5B6C2DF9B307BF4778" box="[331,472,1577,1599]" italics="true" pageId="19" pageNumber="297">Golofa inermis</emphasis>
</taxonomicName>
occurs in
<collectingCountry id="F316765EFFB17D5B6F50F9B3040E4778" box="[566,617,1577,1599]" name="Chile" pageId="19" pageNumber="297">Chile</collectingCountry>
(the
<typeStatus id="54BA886CFFB17D5B6DCAF9DE0763471D" box="[172,260,1604,1626]" pageId="19" pageNumber="297" type="lectotype">lectotype</typeStatus>
and
<specimenCount id="9D07FD47FFB17D5B6C53F9DE0791471D" box="[309,502,1604,1626]" count="3" pageId="19" pageNumber="297" type="generic" typeStatus="paralectotype">three paralectotypes</specimenCount>
in MNHN) and
<collectingCountry id="F316765EFFB17D5B6DCBF9C406934733" box="[173,244,1630,1652]" name="Bolivia" pageId="19" pageNumber="297">Bolivia</collectingCountry>
(
<specimenCount id="9D07FD47FFB17D5B6C67F9C407D04733" box="[257,439,1630,1652]" count="2" pageId="19" pageNumber="297" type="generic" typeStatus="paralectotype">two paralectotypes</specimenCount>
in MNHN).
</paragraph>
<paragraph id="8BBE36CEFFB17D5B6DA3F9E307244784" blockId="19.[172,660,1230,1732]" pageId="19" pageNumber="297">
<emphasis id="B975EADCFFB17D5B6DA3F9E3071147C9" bold="true" box="[197,374,1657,1679]" pageId="19" pageNumber="297">Locality Records</emphasis>
(
<figureCitation id="133A2A4BFFB17D5B6CE3F9E307B747C9" box="[389,464,1657,1679]" captionStart="Figs" captionStartId="19.[198,237,1108,1127]" captionTargetBox="[175,1187,193,1082]" captionTargetId="figure-285@19.[173,1187,193,1082]" captionTargetPageId="19" captionText="Figs. 3437. Golofa inermis. 34) Lectotype, MNHN-EC-2378; 35) Lectotype labels; 36) Parameres; 37) Distribution. Images in Figs. 3435 courtesy of Antoine Mantilleri (Muséum National dHistoire Naturelle, Paris, France)." figureDoi="http://doi.org/10.5281/zenodo.13251827" httpUri="https://zenodo.org/record/13251827/files/figure.png" pageId="19" pageNumber="297">Fig. 37</figureCitation>
).
<materialsCitation id="3B693C93FFB17D5B6C85F9E307584784" collectionCode="MNHN" pageId="19" pageNumber="297" specimenCount="5">
<specimenCount id="9D07FD47FFB17D5B6C85F9E3043B47C8" box="[483,604,1657,1679]" count="5" pageId="19" pageNumber="297" type="generic">5 specimens</specimenCount>
from
<collectionCode id="ED10AE0BFFB17D5B6DCBF90E069E47EE" box="[173,249,1684,1705]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="297">MNHN</collectionCode>
,
<bibRefCitation id="EF904B3FFFB17D5B6C73F90E07AC47EE" author="Gutierrez, R." box="[277,459,1683,1706]" pageId="19" pageNumber="297" pagination="52 - 75" refId="ref16281" refString="Gutierrez, R. 1950. Scarabaeidae del norte de Chile (Coleopt. Lamellic.). Anales de la Sociedad Cientifica Argentina 149 (2): 52 - 75." type="journal article" year="1950">Gutiérrez (1950)</bibRefCitation>
, and Ferrú and Elgueta (2011)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBE36CEFFB17D5B6FBDFB550248445E" blockId="19.[705,1193,1230,1732]" pageId="19" pageNumber="297">
<collectingRegion id="49C5F82CFFB17D5B6FBDFB55021C45A4" box="[731,1147,1230,1254]" country="Chile" name="Arica y Parinacota" pageId="19" pageNumber="297">REGIÓN DE ARICA Y PARINACOTA</collectingRegion>
(1): ARICA (1): Arica.
<materialsCitation id="3B693C93FFB17D5B6E1DFB73024C445E" collectorName="Minita. NO" location="REGION DE TARAPACA" pageId="19" pageNumber="297" specimenCount="1" stateProvince="Tarapaca">
<collectingRegion id="49C5F82CFFB17D5B6E1DFB7302184447" box="[891,1151,1257,1280]" country="Chile" name="Tarapaca" pageId="19" pageNumber="297">REGIÓN DE TARAPACÁ</collectingRegion>
(1): IQUIQUE (1):
<collectorName id="26F45318FFB17D5B6E33FA9905A3445E" box="[853,964,1283,1305]" pageId="19" pageNumber="297">Miñita. NO</collectorName>
DATA (3)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBE36CEFFB17D5B6FBDFA8402474473" blockId="19.[705,1193,1230,1732]" box="[731,1056,1310,1332]" pageId="19" pageNumber="297">
<emphasis id="B975EADCFFB17D5B6FBDFA8405A04473" bold="true" box="[731,967,1310,1332]" pageId="19" pageNumber="297">Temporal Distribution.</emphasis>
No data.
</paragraph>
</subSubSection>
<subSubSection id="C31B6545FFB17D5C6FBDFAA3068740D3" lastPageId="20" lastPageNumber="298" pageId="19" pageNumber="297" type="diagnosis">
<paragraph id="8BBE36CEFFB17D5B6FBDFAA302CE471E" blockId="19.[705,1193,1230,1732]" pageId="19" pageNumber="297">
<emphasis id="B975EADCFFB17D5B6FBDFAA305264408" bold="true" box="[731,833,1337,1359]" pageId="19" pageNumber="297">Diagnosis.</emphasis>
The pronotum and elytra of
<taxonomicName id="4C014D4DFFB17D5B6921FAA302CF4408" authorityName="Thomson" authorityYear="1859" box="[1095,1192,1337,1359]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="297" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB17D5B6921FAA302CF4408" box="[1095,1192,1337,1359]" italics="true" pageId="19" pageNumber="297">G. inermis</emphasis>
</taxonomicName>
are monochromatic dark yellowish brown or reddish brown (except for the black elytral suture); the elytral suture is a complete, impressed stria; and the propygidium lacks long, dense setae. Conversely,
<taxonomicName id="4C014D4DFFB17D5B6FA5FA25054C4493" authorityName="Sternberg" authorityYear="1910" box="[707,811,1471,1492]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="297" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B975EADCFFB17D5B6FA5FA25054C4493" box="[707,811,1471,1492]" italics="true" pageId="19" pageNumber="297">G. minutus</emphasis>
</taxonomicName>
has dark areas on the pronotum, and the elytral margins are black; the elytral sutural stria is a row of closely spaced punctures; and the propygidium has long, dense setae. The parameres of the two species differ, with those of
<taxonomicName id="4C014D4DFFB17D5B6962F9B3020C4779" authorityName="Thomson" authorityYear="1859" box="[1028,1131,1577,1598]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="297" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB17D5B6962F9B3020C4779" box="[1028,1131,1577,1598]" italics="true" pageId="19" pageNumber="297">G. inermis</emphasis>
</taxonomicName>
being thicker and more robust (compare
<figureCitation id="133A2A4BFFB17D5B696FF9D90232471E" box="[1033,1109,1603,1625]" captionStart="Figs" captionStartId="19.[198,237,1108,1127]" captionTargetBox="[175,1187,193,1082]" captionTargetId="figure-285@19.[173,1187,193,1082]" captionTargetPageId="19" captionText="Figs. 3437. Golofa inermis. 34) Lectotype, MNHN-EC-2378; 35) Lectotype labels; 36) Parameres; 37) Distribution. Images in Figs. 3435 courtesy of Antoine Mantilleri (Muséum National dHistoire Naturelle, Paris, France)." figureDoi="http://doi.org/10.5281/zenodo.13251827" httpUri="https://zenodo.org/record/13251827/files/figure.png" pageId="19" pageNumber="297">Figs. 36</figureCitation>
and
<figureCitation id="133A2A4BFFB17D5B69E4F9DE02F9471E" box="[1154,1182,1604,1625]" captionStart="Figs" captionStartId="21.[197,236,1666,1685]" captionTargetBox="[182,1182,193,1637]" captionTargetId="figure-6@21.[173,1182,193,1640]" captionTargetPageId="21" captionText="Figs. 3842. Golofa minutus. 38) Male, dorsal view; 39) Male, lateral view; 40) Parameres; 41) Female, dorsal view; 42) Female, lateral view. Photographs by Marcelo Guerrero (Santiago, Chile)." figureDoi="http://doi.org/10.5281/zenodo.13251831" httpUri="https://zenodo.org/record/13251831/files/figure.png" pageId="19" pageNumber="297">40</figureCitation>
).
</paragraph>
<paragraph id="8BBE36CEFFB17D5C6FBDF9C4044F4004" blockId="19.[705,1193,1230,1732]" lastBlockId="20.[152,638,193,404]" lastPageId="20" lastPageNumber="298" pageId="19" pageNumber="297">
<emphasis id="B975EADCFFB17D5B6FBDF9C405174734" bold="true" box="[731,880,1630,1651]" pageId="19" pageNumber="297">Nomenclature.</emphasis>
<bibRefCitation id="EF904B3FFFB17D5B6E13F9C402744733" author="Thomson, J." box="[885,1043,1630,1652]" pageId="19" pageNumber="297" pagination="3 - 22" refId="ref18285" refString="Thomson, J. 1859. Essai synoptique sur la sous-tribu des scarabaeitae vrais. Arcana Naturae 1: 3 - 22." type="journal article" year="1859">Thomson (1859)</bibRefCitation>
indicated a size range when he described
<taxonomicName id="4C014D4DFFB17D5B6ED3F9E3027C47C8" authorityName="Thomson" authorityYear="1859" box="[949,1051,1657,1679]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="297" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB17D5B6ED3F9E3027C47C8" box="[949,1051,1657,1679]" italics="true" pageId="19" pageNumber="297">G. inermis</emphasis>
</taxonomicName>
, which clearly indicates he had at least
<specimenCount id="9D07FD47FFB17D5B6EDDF90E022847ED" box="[955,1103,1684,1706]" count="2" pageId="19" pageNumber="297" type="generic">two specimens</specimenCount>
.
<bibRefCitation id="EF904B3FFFB17D5B693DF90E05684783" author="Endrodi, S." pageId="19" pageNumber="297" pagination="37 - 86" refId="ref15954" refString="Endrodi, S. 1977 b. Monographie der Dynastinae (Coleoptera) 6. Tribus Dynastini. II. Acta Zoologica Academiae Scientiarum Hungaricae 23: 37 - 86." type="journal article" year="1977">Endrödi (1977b)</bibRefCitation>
said the type of
<taxonomicName id="4C014D4DFFB17D5B6ED2F935027C4783" authorityName="Thomson" authorityYear="1859" box="[948,1051,1711,1732]" class="Insecta" family="Scarabaeidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="297" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B975EADCFFB17D5B6ED2F935027C4783" box="[948,1051,1711,1732]" italics="true" pageId="19" pageNumber="297">G. inermis</emphasis>
</taxonomicName>
was unknown to him, but
<bibRefCitation id="EF904B3FFFB67D5C6C79FF5807BB4190" author="Dechambre, R. - P." box="[287,476,193,216]" pageId="20" pageNumber="298" pagination="84 - 89" refId="ref15543" refString="Dechambre, R. - P. 1975. Designation de types et note synonymique sur des Dynastinae [Col. Scarabaeidae]. Bulletin de la Societe Entomologique de France (N. S.) 80: 84 - 89." type="journal article" year="1975">Dechambre (1975)</bibRefCitation>
discovered the Thomson specimens in the MNHN and designated a
<typeStatus id="54BA886CFFB67D5C6DCDFF6D0764404A" box="[171,259,247,269]" pageId="20" pageNumber="298" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="133A2A4BFFB67D5C6C75FF6D07ED404A" box="[275,394,247,269]" captionStart="Figs" captionStartId="19.[198,237,1108,1127]" captionTargetBox="[175,1187,193,1082]" captionTargetId="figure-285@19.[173,1187,193,1082]" captionTargetPageId="19" captionText="Figs. 3437. Golofa inermis. 34) Lectotype, MNHN-EC-2378; 35) Lectotype labels; 36) Parameres; 37) Distribution. Images in Figs. 3435 courtesy of Antoine Mantilleri (Muséum National dHistoire Naturelle, Paris, France)." figureDoi="http://doi.org/10.5281/zenodo.13251827" httpUri="https://zenodo.org/record/13251827/files/figure.png" pageId="20" pageNumber="298">Figs. 3435</figureCitation>
) and
<specimenCount id="9D07FD47FFB67D5C6CA3FF6D0419404A" box="[453,638,247,269]" count="5" pageId="20" pageNumber="298" type="generic" typeStatus="paralectotype">five paralectotypes</specimenCount>
(three from
<collectingCountry id="F316765EFFB67D5C6C6DFE880726406F" box="[267,321,274,296]" name="Chile" pageId="20" pageNumber="298">Chile</collectingCountry>
, two from
<collectingCountry id="F316765EFFB67D5C6CCFFE880793406F" box="[425,500,274,296]" name="Bolivia" pageId="20" pageNumber="298">Bolivia</collectingCountry>
). We were unable to directly examine these specimens.
</paragraph>
<paragraph id="8BBE36CEFFB67D5C6DD6FED2068740D3" blockId="20.[152,638,193,404]" pageId="20" pageNumber="298">
<emphasis id="B975EADCFFB67D5C6DD6FED2073E4019" bold="true" box="[176,345,328,350]" pageId="20" pageNumber="298">Natural History.</emphasis>
Adults are attracted to lights, but we know nothing else of the species natural history.
</paragraph>
</subSubSection>
</treatment>
</document>