281 lines
35 KiB
XML
281 lines
35 KiB
XML
<document id="30570302496E42215CD49AC977EB2662" ID-DOI="10.11646/zootaxa.5002.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5122655" ID-ZooBank="5DC2A66A-3F04-42D0-8A8C-F0686054E556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626888359662" checkinUser="plazi" docAuthor="Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott" docDate="2021" docId="03A887D05F70FFBBFF3EEA08FB594889" docLanguage="en" docName="zootaxa.5002.1.1.pdf" docOrigin="Zootaxa 5002 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomosvaryella oshidae Motamedinia & Skevington, sp. nov." docType="treatment" docVersion="4" lastPageNumber="51" masterDocId="FF91FFA85F42FF89FFA9EB42FFAF4F30" masterDocTitle="Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species" masterLastPageNumber="103" masterPageNumber="1" pageNumber="51" updateTime="1699012478263" updateUser="plazi">
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<mods:titleInfo id="81483C3B539FEB7793DFA85087BA0B10">
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<mods:title id="7C80D24EEF864E231B5D907429F98C66">Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species</mods:title>
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</mods:titleInfo>
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<mods:name id="49F129520099CD0CBCA294A131D5CC33" type="personal">
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<mods:roleTerm id="C46BF4364780D6F8192D2349FE15D52B">Author</mods:roleTerm>
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<mods:namePart id="6DA620025059A67920AE41A98214BC87">Motamedinia, Behnam</mods:namePart>
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<mods:roleTerm id="2A22E68C015F67A09E53B2ECCE34CA64">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="57AA9097EEC84D7EA28AE9E198373F83">Skevington, Jeffrey H.</mods:namePart>
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<mods:nameIdentifier id="15AEA2277619DEFC8150DEA8B0926914" type="ORCID">0000-0002-1445-9870</mods:nameIdentifier>
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<mods:nameIdentifier id="8981686DE86DE0E9D4E67C3B10CF104F" type="email">jhskevington@gmail.com</mods:nameIdentifier>
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<mods:name id="021F134A445962E409FECC0975DF5DA1" type="personal">
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<mods:roleTerm id="8B095468A4FF6EBF320901A11A327C8F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9DA9620D1D38223011858790F8D183E4">Kelso, Scott</mods:namePart>
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<mods:nameIdentifier id="62D325CF48326BAF128EE9D213C9F03E" type="ORCID">0000-0001-5559-9477</mods:nameIdentifier>
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<mods:nameIdentifier id="0265B02C4CD2C6282486658A06AC8401" type="email">scott.kelso@canada.ca</mods:nameIdentifier>
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<mods:typeOfResource id="FE6E2F9AA3CE317C7881C7CD25B0D5CD">text</mods:typeOfResource>
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<mods:title id="0C3284AD62DCAB0EA0E3C094101EDA3E">Zootaxa</mods:title>
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<mods:part id="958874F65CB90D3EE6AB9D50ED635F9B">
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<mods:date id="CAE18B3A846EB3B29DB229A7F1D331F4">2021</mods:date>
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<mods:number id="751E0B95279647491398AA948D4D6D30">2021-07-16</mods:number>
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<mods:detail id="BBD6A4A849DCC63595D2FDA51ADE31BC" type="volume">
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<mods:number id="C3E69A0B287D5D8A21783C396438E5BA">5002</mods:number>
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<mods:detail id="B3D37FFA94A9BC429902F430F045DA0A" type="issue">
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<mods:number id="08A8BC2D16AFE2FEB86F8B50C0643B94">1</mods:number>
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<mods:classification id="1B13931ACC86B9926A8A6B856267902A">journal article</mods:classification>
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<mods:identifier id="CB020B95CED209232A8BEB3135857357" type="DOI">10.11646/zootaxa.5002.1.1</mods:identifier>
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<mods:identifier id="138C2BED8D127EC8418C8A77DD032AAF" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A887D05F70FFBBFF3EEA08FB594889" ID-DOI="http://doi.org/10.5281/zenodo.5122591" ID-Zenodo-Dep="5122591" LSID="urn:lsid:plazi:treatment:03A887D05F70FFBBFF3EEA08FB594889" httpUri="http://treatment.plazi.org/id/03A887D05F70FFBBFF3EEA08FB594889" lastPageNumber="51" pageId="50" pageNumber="51">
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<subSubSection id="C31B654D5F70FFBBFF3EEA08FCDD4E55" box="[151,882,330,357]" pageId="50" pageNumber="51" type="nomenclature">
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<paragraph id="8BBE36C65F70FFBBFF3EEA08FCDD4E55" blockId="50.[151,882,330,393]" box="[151,882,330,357]" pageId="50" pageNumber="51">
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<heading id="D0F681AA5F70FFBBFF3EEA08FCDD4E55" bold="true" box="[151,882,330,357]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="3">
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||
<emphasis id="B975EAD45F70FFBBFF3EEA08FCDD4E55" bold="true" box="[151,882,330,357]" pageId="50" pageNumber="51">
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<taxonomicName id="4C014D455F70FFBBFF3EEA08FCA24E54" ID-CoL="9HKN8" authority="Motamedinia & Skevington" authorityName="Motamedinia & Skevington" box="[151,781,330,357]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oshidae" status="sp. nov.">
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<emphasis id="B975EAD45F70FFBBFF3EEA08FE054E55" bold="true" box="[151,426,330,357]" italics="true" pageId="50" pageNumber="51">Tomosvaryella oshidae</emphasis>
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Motamedinia & Skevington
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</taxonomicName>
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<taxonomicNameLabel id="A24657AF5F70FFBBFCBDEA09FCDD4E55" box="[788,882,331,357]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31B654D5F70FFBBFF3EEA2DFE4A4EB9" box="[151,485,366,393]" pageId="50" pageNumber="51" type="description">
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||
<paragraph id="8BBE36C65F70FFBBFF3EEA2DFE4A4EB9" blockId="50.[151,882,330,393]" box="[151,485,366,393]" pageId="50" pageNumber="51">
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||
<figureCitation id="133A2A435F70FFBBFF3EEA2DFEAD4EB9" box="[151,258,367,393]" captionStart="FIGURE 27" captionStartId="51.[151,250,1677,1701]" captionTargetBox="[158,1430,212,1618]" captionTargetId="figure-21@51.[158,1430,211,1620]" captionTargetPageId="51" captionText="FIGURE 27. Ovipositors in lateral view A) Tomosvaryella ampliasa sp. nov., paratype (JSS43959, TAU), B) Tomosvaryella pilosiventris (JSS51961, CNC), C) Tomosvaryella pruinosa (CNC470591, CNC), D) Tomosvaryella spinula sp. nov., paratype (JSS51820, CNC), E) Tomosvaryella emaratensis sp. nov., paratype (CNCD137556, CNC), F) Tomosvaryella parakuthyi (CNC470514, CNC), G) Tomosvaryella oshidae sp. nov., paratype (JSS51976, CNC), H) Tomosvaryella soziana sp. nov., paratype (JSS52205, CNC), I) Tomosvaryella nodosa (CNC470648, CNC), J) T. hamounensis sp. nov., paratype (JSS52178, CNC)." figureDoi="http://doi.org/10.5281/zenodo.5122727" httpUri="https://zenodo.org/record/5122727/files/figure.png" pageId="50" pageNumber="51">Figs 27G</figureCitation>
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,
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<figureCitation id="133A2A435F70FFBBFEA4EA2DFEF04EB9" box="[269,351,367,393]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">28A–E</figureCitation>
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,
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<figureCitation id="133A2A435F70FFBBFEC3EA2CFE274EB9" box="[362,392,366,393]" captionStart="FIGURE 59" captionStartId="90.[151,250,847,871]" captionTargetBox="[151,1436,181,823]" captionTargetId="figure-31@90.[151,1436,181,823]" captionTargetPageId="90" captionText="FIGURE 59. Distribution of Tomosvaryella nodosa, T. oshidae sp. nov., T. pistacia in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122789" httpUri="https://zenodo.org/record/5122789/files/figure.png" pageId="50" pageNumber="51">59</figureCitation>
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,
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<figureCitation id="133A2A435F70FFBBFE3DEA2DFE4A4EB9" box="[404,485,367,393]" captionStart="FIGURE 68" captionStartId="96.[151,250,1518,1542]" captionTargetBox="[151,1436,182,1494]" captionTargetId="figure-31@96.[151,1436,181,1494]" captionTargetPageId="96" captionText="FIGURE 68. Male habitus of Tomosvaryella species A–B) T. nimroozensis sp. nov., holotype (JSS51819, CNC) A) in dorsal view B) in lateral view, C–D) T. oshidae sp. nov., holotype (JSS51818, CNC) C) in dorsal view D) in lateral view, E–F) T. osteodes sp. nov., holotype (CNCD175298, CNC) E) in dorsal view F) in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122809" httpUri="https://zenodo.org/record/5122809/files/figure.png" pageId="50" pageNumber="51">68C–D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B654D5F70FFBBFF3EEAF4FDEC4D0D" pageId="50" pageNumber="51" type="diagnosis">
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<paragraph id="8BBE36C65F70FFBBFF3EEAF4FDEC4D0D" blockId="50.[151,1437,438,1977]" pageId="50" pageNumber="51">
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<emphasis id="B975EAD45F70FFBBFF3EEAF4FEBB4EE0" bold="true" box="[151,276,438,464]" pageId="50" pageNumber="51">Diagnosis:</emphasis>
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The male of this species can be recognized by the shape of surstyli in dorsal view, broad basically, constricted in apical third (
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<figureCitation id="133A2A435F70FFBBFE33EA99FDAC4EC5" box="[410,515,475,501]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">Fig. 28A</figureCitation>
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); large gonopods in ventral view (
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<figureCitation id="133A2A435F70FFBBFCD7EA99FC4B4EC5" box="[894,996,475,501]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">Fig. 28B</figureCitation>
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); broad phallic guide; ejaculatory ducts strictly bent into phallic guide in lateral view (
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<figureCitation id="133A2A435F70FFBBFD08EABDFCA74D29" box="[673,776,511,537]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">Fig. 28C</figureCitation>
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). The female can be recognized by long ovipositor, curved and reaching the mid coxa (
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<figureCitation id="133A2A435F70FFBBFE67E961FD984D0D" box="[462,567,547,573]" captionStart="FIGURE 27" captionStartId="51.[151,250,1677,1701]" captionTargetBox="[158,1430,212,1618]" captionTargetId="figure-21@51.[158,1430,211,1620]" captionTargetPageId="51" captionText="FIGURE 27. Ovipositors in lateral view A) Tomosvaryella ampliasa sp. nov., paratype (JSS43959, TAU), B) Tomosvaryella pilosiventris (JSS51961, CNC), C) Tomosvaryella pruinosa (CNC470591, CNC), D) Tomosvaryella spinula sp. nov., paratype (JSS51820, CNC), E) Tomosvaryella emaratensis sp. nov., paratype (CNCD137556, CNC), F) Tomosvaryella parakuthyi (CNC470514, CNC), G) Tomosvaryella oshidae sp. nov., paratype (JSS51976, CNC), H) Tomosvaryella soziana sp. nov., paratype (JSS52205, CNC), I) Tomosvaryella nodosa (CNC470648, CNC), J) T. hamounensis sp. nov., paratype (JSS52178, CNC)." figureDoi="http://doi.org/10.5281/zenodo.5122727" httpUri="https://zenodo.org/record/5122727/files/figure.png" pageId="50" pageNumber="51">Fig. 27G</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B654D5F70FFBBFF6EE904FE1149A9" pageId="50" pageNumber="51" type="description">
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<paragraph id="8BBE36C65F70FFBBFF6EE904FA374A3D" blockId="50.[151,1437,438,1977]" pageId="50" pageNumber="51">
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<emphasis id="B975EAD45F70FFBBFF6EE904FE184D50" bold="true" box="[199,439,582,608]" pageId="50" pageNumber="51">Description: MALE</emphasis>
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: Body length (excluding antennae):
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<quantity id="4CF99B235F70FFBBFCFFE904FC1E4D51" box="[854,945,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="50" pageNumber="51" unit="mm" value="4.5">4.5 mm</quantity>
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.
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<emphasis id="B975EAD45F70FFBBFC15E904FBAD4D50" box="[956,1026,582,609]" italics="true" pageId="50" pageNumber="51">
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<emphasis id="B975EAD45F70FFBBFC15E904FC554D51" bold="true" box="[956,1018,582,609]" italics="true" pageId="50" pageNumber="51">Head</emphasis>
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.
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</emphasis>
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Scape, pedicel, flagellum and arista dark. Pedicel with three upper dark setae; flagellum light brown, gray pollinose and tapering (LF:WF = 1.7). Eyes meeting for a distance of 13‒14 facets. Frons silver-gray pollinose; vertex shining; occiput gray pollinose with scattered light brown setae.
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<emphasis id="B975EAD45F70FFBBFE7FE9F0FD834DFD" bold="true" box="[470,556,690,717]" italics="true" pageId="50" pageNumber="51">Thorax</emphasis>
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. Postpronotal lobe light brown with pale setae along the upper margin (up to
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<quantity id="4CF99B235F70FFBBFF3EE995FEAF4DC1" box="[151,256,727,753]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="51" unit="mm" value="0.01">0.01 mm</quantity>
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). Prescutum and scutum black, light brown pollinose; Scutum with two dark uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with 12 setae along posterior margin. Subscutellum black, pleura dark brown.
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<emphasis id="B975EAD45F70FFBBFF45E85DFE9F4C08" box="[236,304,799,825]" italics="true" pageId="50" pageNumber="51">
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<emphasis id="B975EAD45F70FFBBFF45E85DFE864C09" bold="true" box="[236,297,799,825]" italics="true" pageId="50" pageNumber="51">Wing</emphasis>
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.
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</emphasis>
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Wing length:
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<quantity id="4CF99B235F70FFBBFE67E85DFD884C09" box="[462,551,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="50" pageNumber="51" unit="mm" value="2.4">2.4 mm</quantity>
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. LW:MWW = 3.4. Wing almost entirely covered with microtrichia. M
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<subScript id="178534835F70FFBBFA9BE873FA954C0F" attach="left" box="[1330,1338,817,831]" fontSize="6" pageId="50" pageNumber="51">1</subScript>
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strongly undulating in middle. Halter length:
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<quantity id="4CF99B235F70FFBBFD85E800FD2A4C6D" box="[556,645,834,861]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="mm" value="0.5">0.5 mm</quantity>
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. Brown to yellowish, narrowly black at base.
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<emphasis id="B975EAD45F70FFBBFBD6E801FB1A4C6D" bold="true" box="[1151,1205,835,861]" italics="true" pageId="50" pageNumber="51">Legs</emphasis>
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. Coxae and trochanters black, gray pollinose, mid coxa with three black anterior setae; hind trochanter with scattered short pale bristle. Femora dark brown with narrow yellow apices, silver pollinose and 5‒8 ventral spines towards apex. All femora with one wrinkled indentation in basal. Tibiae light brown at base, gray pollinose, with two rows of short black setae on anterior and three rows on posterior side. Hind tibia with one wrinkled indentation in middle. Tibiae without distal spines. Tarsi light brown, distitarsi dark, claws light brown with black tips.
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<emphasis id="B975EAD45F70FFBBFB98E8B4FB304B21" bold="true" box="[1073,1183,1014,1041]" italics="true" pageId="50" pageNumber="51">Abdomen</emphasis>
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. Ground color brown, tergite 1 covered by densely silver-gray pollinose, with eight short dark lateral setae, tergite 2‒5 with weak silvergray pollinose and with scattered brown setae. Sternites white-yellow laterally with dark mid-line centrally, gray pollinose.
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<emphasis id="B975EAD45F70FFBBFEA5EF20FED14B4C" box="[268,382,1122,1149]" italics="true" pageId="50" pageNumber="51">
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<emphasis id="B975EAD45F70FFBBFEA5EF20FED84B4D" bold="true" box="[268,375,1122,1149]" italics="true" pageId="50" pageNumber="51">Genitalia</emphasis>
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.
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</emphasis>
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Genital capsule in dorsal view: surstyli asymmetrical, both with broad base and constricted in apical third but base of the right boarder than left one, left surstyli knobbed apically, right surstyli curved toward left one apically (
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<figureCitation id="133A2A435F70FFBBFE98EFE9FE344BF5" box="[305,411,1195,1221]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">Fig. 28A</figureCitation>
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), epandrium wider than long. Genital capsule in ventral view: gonopods large, broadened, equal in height (
|
||
<figureCitation id="133A2A435F70FFBBFEFBEF8DFE144BD9" box="[338,443,1231,1257]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">Fig. 28B</figureCitation>
|
||
). Genital capsule in lateral view: both surstyli broadened at base, strictly bent in apical third towards sternite (
|
||
<figureCitation id="133A2A435F70FFBBFE3DEFB1FDB34A3D" box="[404,540,1267,1293]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">Fig. 28D‒E</figureCitation>
|
||
); phallic guide broad, pointed; phallus wavy, bent into phallic guide (
|
||
<figureCitation id="133A2A435F70FFBBFA8DEFB1FA244A3D" box="[1316,1419,1267,1293]" captionStart="FIGURE 28" captionStartId="52.[151,250,1816,1840]" captionTargetBox="[164,1423,188,1783]" captionTargetId="figure-31@52.[151,1436,181,1792]" captionTargetPageId="52" captionText="FIGURE 28. Male genitalia of Tomosvaryella oshidae sp. nov., holotype (JSS51818, CNC) A) dorsal view, B) ventral view, C) phallic guide and phallus in lateral view, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122729" httpUri="https://zenodo.org/record/5122729/files/figure.png" pageId="50" pageNumber="51">Fig. 28C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F70FFBBFF6EEE54FE1149A9" blockId="50.[151,1437,438,1977]" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFF6EEE54FE904A00" bold="true" box="[199,319,1302,1328]" pageId="50" pageNumber="51">FEMALE</emphasis>
|
||
: Body length (excluding antennae):
|
||
<quantity id="4CF99B235F70FFBBFD43EE55FCE84A01" box="[746,839,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="50" pageNumber="51" unit="mm" value="3.6">3.6 mm</quantity>
|
||
.
|
||
<emphasis id="B975EAD45F70FFBBFCFCEE54FC3C4A01" bold="true" box="[853,915,1302,1329]" italics="true" pageId="50" pageNumber="51">Head</emphasis>
|
||
. Eyes separated. Frons silver-gray pollinose, concaved shortly after ocelli to lower third of frons. Occiput gray pollinose. Pedicel with four upper dark setae and one lower setae.
|
||
<emphasis id="B975EAD45F70FFBBFEFEEE1CFE1C4A48" box="[343,435,1374,1401]" italics="true" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFEFEEE1CFE024A49" bold="true" box="[343,429,1374,1401]" italics="true" pageId="50" pageNumber="51">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Postpronotal lobe light yellow, gray pollinose, with 4–5 light brown setae along the upper margin (up to
|
||
<quantity id="4CF99B235F70FFBBFEC1EEC1FE7D4AAD" box="[360,466,1411,1437]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="51" unit="mm" value="0.01">0.01 mm</quantity>
|
||
). Prescutum and scutum, scutellum black, light brown pollinose in dorsocentral area.
|
||
<emphasis id="B975EAD45F70FFBBFF3EEEE5FF744AF0" box="[151,219,1447,1473]" italics="true" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFF3EEEE5FF7B4AF1" bold="true" box="[151,212,1447,1473]" italics="true" pageId="50" pageNumber="51">Wing</emphasis>
|
||
.
|
||
</emphasis>
|
||
Length:
|
||
<quantity id="4CF99B235F70FFBBFEECEEE4FE0F4AF1" box="[325,416,1446,1473]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="50" pageNumber="51" unit="mm" value="3.5">3.5 mm</quantity>
|
||
. LW:MWW = 3.8.
|
||
<emphasis id="B975EAD45F70FFBBFDD6EEE5FD1A4AF1" bold="true" box="[639,693,1447,1473]" italics="true" pageId="50" pageNumber="51">Legs</emphasis>
|
||
. Hind trochanter with 6–8 light brown setae. All femora bearing one wrinkled indentation in basal. Fore femur more widened. Hind trochanter with two wrinkled indentations in middle. Distitarsi with long brown setae at anterior margin. Pulvilli broadened and leaf-shaped; claws long, twice the length of distitarsi.
|
||
<emphasis id="B975EAD45F70FFBBFE0BED50FDB7491C" box="[418,536,1554,1581]" italics="true" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFE0BED50FDBF491D" bold="true" box="[418,528,1554,1581]" italics="true" pageId="50" pageNumber="51">Abdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
Tergites 1–6 gray pollinose with scattered short light brown setae.
|
||
<emphasis id="B975EAD45F70FFBBFAB4ED51FA3A491D" bold="true" box="[1309,1429,1555,1581]" italics="true" pageId="50" pageNumber="51">Ovipositor</emphasis>
|
||
. Base of ovipositor dark brown, gray pollinose dorsally, with some scattered short pale setae. Viewed laterally (
|
||
<figureCitation id="133A2A435F70FFBBFAC6ED75FF644945" captionStart="FIGURE 27" captionStartId="51.[151,250,1677,1701]" captionTargetBox="[158,1430,212,1618]" captionTargetId="figure-21@51.[158,1430,211,1620]" captionTargetPageId="51" captionText="FIGURE 27. Ovipositors in lateral view A) Tomosvaryella ampliasa sp. nov., paratype (JSS43959, TAU), B) Tomosvaryella pilosiventris (JSS51961, CNC), C) Tomosvaryella pruinosa (CNC470591, CNC), D) Tomosvaryella spinula sp. nov., paratype (JSS51820, CNC), E) Tomosvaryella emaratensis sp. nov., paratype (CNCD137556, CNC), F) Tomosvaryella parakuthyi (CNC470514, CNC), G) Tomosvaryella oshidae sp. nov., paratype (JSS51976, CNC), H) Tomosvaryella soziana sp. nov., paratype (JSS52205, CNC), I) Tomosvaryella nodosa (CNC470648, CNC), J) T. hamounensis sp. nov., paratype (JSS52178, CNC)." figureDoi="http://doi.org/10.5281/zenodo.5122727" httpUri="https://zenodo.org/record/5122727/files/figure.png" pageId="50" pageNumber="51">Fig. 27G</figureCitation>
|
||
), base of piercer curved, piercer long and angled between proximal and distal part and reached mid coxa. LP: LB = 2.5. LDP:LPP = 4.0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F70FFBBFF6EEDE1FACE487D" pageId="50" pageNumber="51" type="etymology">
|
||
<paragraph id="8BBE36C65F70FFBBFF6EEDE1FC8B49D1" blockId="50.[151,1437,438,1977]" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFF6EEDE1FEFD498D" bold="true" box="[199,338,1699,1725]" pageId="50" pageNumber="51">Etymology:</emphasis>
|
||
The specific epithet refers to Oshida, the old name of Khaje mountain and a female name in mythology, from where the
|
||
<typeStatus id="54BA88645F70FFBBFE02ED85FDA249D1" box="[427,525,1735,1761]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
|
||
specimen was collected.
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F70FFBBFF6EEDA8FACE487D" blockId="50.[151,1437,438,1977]" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFF6EEDA8FE644834" bold="true" box="[199,459,1770,1796]" pageId="50" pageNumber="51">Specimens examined:</emphasis>
|
||
<materialsCitation id="3B693C9B5F70FFBBFE7DEDA8FC884819" accessionNumber="JSS51818" collectingDate="2016-06-06" collectionCode="CNC" collectorName="Sistan & Baluchestan & H. Derafshan" country="Iran" elevation="481" latitude="31.116667" location="Zabol" longLatPrecision="1218" longitude="61.466667" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B975EAD45F70FFBBFE7DEDA8FDD34834" bold="true" box="[468,636,1770,1796]" pageId="50" pageNumber="51">
|
||
<typeStatus id="54BA88645F70FFBBFE7DEDA8FDD84834" box="[468,631,1770,1796]" pageId="50" pageNumber="51" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectingCountry id="F31676565F70FFBBFD2CEDA9FD614835" box="[645,718,1771,1797]" name="Iran" pageId="50" pageNumber="51">IRAN</collectingCountry>
|
||
:
|
||
<collectorName id="26F453105F70FFBBFD73EDA9FCB14835" box="[730,798,1771,1797]" pageId="50" pageNumber="51">Sistan</collectorName>
|
||
&
|
||
<collectorName id="26F453105F70FFBBFCEFEDA9FC7C4835" box="[838,979,1771,1797]" pageId="50" pageNumber="51">Baluchestan</collectorName>
|
||
:
|
||
<location id="8EDE601D5F70FFBBFC49EDA9FB8B4835" LSID="urn:lsid:plazi:treatment:03A887D05F70FFBBFF3EEA08FB594889:8EDE601D5F70FFBBFC49EDA9FB8B4835" box="[992,1060,1771,1797]" country="Iran" latitude="31.116667" longLatPrecision="1218" longitude="61.466667" name="Zabol" pageId="50" pageNumber="51">Zabol</location>
|
||
,
|
||
<geoCoordinate id="EE3550015F70FFBBFB98EDA9FB3B4835" box="[1073,1172,1771,1797]" degrees="31" direction="north" minutes="07" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="31.116667">31°07’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3550015F70FFBBFB0BEDA9FAAE4835" box="[1186,1281,1771,1797]" degrees="61" direction="east" minutes="28" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="61.466667">61°28’E</geoCoordinate>
|
||
,
|
||
<elevation id="002CD1F55F70FFBBFAA6EDA9FA384835" box="[1295,1431,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="4.81" pageId="50" pageNumber="51" unit="m" value="481.0">
|
||
<quantity id="4CF99B235F70FFBBFAA6EDA9FAF74835" box="[1295,1368,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="4.81" pageId="50" pageNumber="51" unit="m" value="481.0">481 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFBF10065F70FFBBFF3EEC4DFEAC4819" box="[151,259,1807,1833]" pageId="50" pageNumber="51" value="2016-06-06">
|
||
<collectingDate id="EFFBE9EE5F70FFBBFF3EEC4DFEAC4819" box="[151,259,1807,1833]" pageId="50" pageNumber="51" value="2016-06-06">6.vi.2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F453105F70FFBBFEA6EC4DFE084819" box="[271,423,1807,1833]" pageId="50" pageNumber="51">H. Derafshan</collectorName>
|
||
,
|
||
<collectingMethod id="52404ED15F70FFBBFE1BEC4DFD8B4819" box="[434,548,1807,1833]" pageId="50" pageNumber="51">sweep net</collectingMethod>
|
||
, JSS51818 (
|
||
<specimenCount id="9D07FD4F5F70FFBBFD19EC4DFD7B4819" box="[688,724,1807,1833]" pageId="50" pageNumber="51" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED10AE035F70FFBBFD49EC4DFCB14819" box="[736,798,1807,1833]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="50" pageNumber="51">CNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B693C9B5F70FFBBFC9BEC4CFAF2487D" accessionNumber="JSS51976" collectingDate="2016-07-03" collectingDateMax="2016-07-17" collectingDateMin="2016-07-03" collectionCode="CNC" collectorName="B. Motamedinia" country="Iran" elevation="1419" latitude="32.866665" location="Mohammadieh" longLatPrecision="1208" longitude="59.016666" pageId="50" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Khorasan-e Jonubi" typeStatus="paratype">
|
||
<emphasis id="B975EAD45F70FFBBFC9BEC4CFC7F4818" bold="true" box="[818,976,1806,1832]" pageId="50" pageNumber="51">
|
||
<typeStatus id="54BA88645F70FFBBFC9BEC4CFC644818" box="[818,971,1806,1832]" pageId="50" pageNumber="51" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectingCountry id="F31676565F70FFBBFC7FEC4DFBB04819" box="[982,1055,1807,1833]" name="Iran" pageId="50" pageNumber="51">IRAN</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C5F8245F70FFBBFB80EC4DFB4D4819" box="[1065,1250,1807,1833]" country="Iran" name="Khorasan-e Jonubi" pageId="50" pageNumber="51">South Khorasan</collectingRegion>
|
||
:
|
||
<location id="8EDE601D5F70FFBBFB45EC4DFA364819" LSID="urn:lsid:plazi:treatment:03A887D05F70FFBBFF3EEA08FB594889:8EDE601D5F70FFBBFB45EC4DFA364819" box="[1260,1433,1807,1833]" country="Iran" latitude="32.866665" longLatPrecision="1208" longitude="59.016666" name="Mohammadieh" pageId="50" pageNumber="51" stateProvince="Khorasan-e Jonubi">Mohammadieh</location>
|
||
,
|
||
<geoCoordinate id="EE3550015F70FFBBFF3EEC70FF55487D" box="[151,250,1842,1869]" degrees="32" direction="north" minutes="52" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="32.866665">32°52’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3550015F70FFBBFEAFEC70FEC9487D" box="[262,358,1842,1869]" degrees="59" direction="east" minutes="01" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="59.016666">59°01’E</geoCoordinate>
|
||
,
|
||
<elevation id="002CD1F55F70FFBBFEDBEC71FDAC487D" box="[370,515,1843,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.419" pageId="50" pageNumber="51" unit="m" value="1419.0">
|
||
<quantity id="4CF99B235F70FFBBFEDBEC71FE68487D" box="[370,455,1843,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.419" pageId="50" pageNumber="51" unit="m" value="1419.0">1419 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFBF10065F70FFBBFDA6EC71FD01487D" box="[527,686,1843,1869]" pageId="50" pageNumber="51" value="2016-07-03" valueMax="2016-07-17" valueMin="2016-07-03">
|
||
<collectingDate id="EFFBE9EE5F70FFBBFDA6EC71FD01487D" box="[527,686,1843,1869]" pageId="50" pageNumber="51" value="2016-07-03" valueMax="2016-07-17" valueMin="2016-07-03">3–17.vii.2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F453105F70FFBBFD10EC71FCDD487D" box="[697,882,1843,1869]" pageId="50" pageNumber="51">B. Motamedinia</collectorName>
|
||
,
|
||
<collectingMethod id="52404ED15F70FFBBFCD4EC71FBF6487D" box="[893,1113,1843,1869]" pageId="50" pageNumber="51">funnel Malaise trap</collectingMethod>
|
||
, JSS51976 (
|
||
<specimenCount id="9D07FD4F5F70FFBBFB41EC71FAA6487D" box="[1256,1289,1843,1869]" pageId="50" pageNumber="51" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED10AE035F70FFBBFABFEC71FAFB487D" box="[1302,1364,1843,1869]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="50" pageNumber="51">CNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F70FFBBFF6EEC14FB594889" pageId="50" pageNumber="51" type="distribution">
|
||
<paragraph id="8BBE36C65F70FFBBFF6EEC14FDA84841" blockId="50.[151,1437,438,1977]" box="[199,519,1878,1905]" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFF6EEC14FECC4840" bold="true" box="[199,355,1878,1904]" pageId="50" pageNumber="51">Distribution:</emphasis>
|
||
<collectingCountry id="F31676565F70FFBBFEC3EC15FE384841" box="[362,407,1879,1905]" name="Iran" pageId="50" pageNumber="51">Iran</collectingCountry>
|
||
(
|
||
<figureCitation id="133A2A435F70FFBBFE0FEC15FE544841" box="[422,507,1878,1905]" captionStart="FIGURE 59" captionStartId="90.[151,250,847,871]" captionTargetBox="[151,1436,181,823]" captionTargetId="figure-31@90.[151,1436,181,823]" captionTargetPageId="90" captionText="FIGURE 59. Distribution of Tomosvaryella nodosa, T. oshidae sp. nov., T. pistacia in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122789" httpUri="https://zenodo.org/record/5122789/files/figure.png" pageId="50" pageNumber="51">Fig. 59</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F70FFBBFF6EEC39FB594889" blockId="50.[151,1437,438,1977]" pageId="50" pageNumber="51">
|
||
<emphasis id="B975EAD45F70FFBBFF6EEC39FEA748A5" bold="true" box="[199,264,1915,1941]" pageId="50" pageNumber="51">Note:</emphasis>
|
||
Based on DNA barcodes,
|
||
<taxonomicName id="4C014D455F70FFBBFD98EC39FD0C48A5" box="[561,675,1915,1941]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oshidae" status="sp. nov.">
|
||
<emphasis id="B975EAD45F70FFBBFD98EC39FD0C48A5" box="[561,675,1915,1941]" italics="true" pageId="50" pageNumber="51">T. oshidae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B975EAD45F70FFBBFD03EC39FCAB48A5" bold="true" box="[682,772,1915,1941]" pageId="50" pageNumber="51">
|
||
<taxonomicNameLabel id="A24657AF5F70FFBBFD03EC39FCAB48A5" box="[682,772,1915,1941]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is genetically most similar to
|
||
<taxonomicName id="4C014D455F70FFBBFBFDEC39FAA048A5" baseAuthorityName="Perkins" baseAuthorityYear="1905" box="[1108,1295,1915,1941]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pseudophanes">
|
||
<emphasis id="B975EAD45F70FFBBFBFDEC39FAA048A5" box="[1108,1295,1915,1941]" italics="true" pageId="50" pageNumber="51">T. pseudophanes</emphasis>
|
||
</taxonomicName>
|
||
, with 11.1% pairwise difference. The male and female sequenced differ by only 0.17% (Supplementary
|
||
<tableCitation id="C683037D5F70FFBBFB3AECDDFB4A4889" box="[1171,1253,1951,1977]" captionStart="TABLE 1" captionStartId="5.[151,239,810,836]" captionText="TABLE 1. Cytochrome c oxidase subunit I (COI) and internal transcribed spacer (ITS) primers." pageId="50" pageNumber="51">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |