270 lines
33 KiB
XML
270 lines
33 KiB
XML
<document id="9C01278BE81AF57B7C833A4BAD567406" ID-DOI="10.11646/phytotaxa.484.1.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="5421338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630379153666" checkinUser="marcus" docAuthor="Goula, Katerina & Constantinidis, Theophanis" docDate="2021" docId="03A887C3FFD6FFA6FF00F950FB61FEC2" docLanguage="en" docName="phytotaxa.484.1.7.pdf" docOrigin="Phytotaxa 484 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.484.1.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Anthemis rigida subsp. runemarkii Biel & Kit Tan 2020" docType="treatment" docVersion="4" lastPageNumber="140" masterDocId="FF91FFBBFFDEFFADFF88FFBCFF8EFFC2" masterDocTitle="Taxonomic diversity and karyology of Anthemis rigida (Anthemideae, Asteraceae) in the Aegean, Greece" masterLastPageNumber="143" masterPageNumber="129" pageNumber="137" updateTime="1699252987701" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E1FF65B5D3B778F3F4486C1CBCF32FB1">Taxonomic diversity and karyology of Anthemis rigida (Anthemideae, Asteraceae) in the Aegean, Greece</mods:title>
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<mods:name id="7C86D1A3F7E73EE62548346823B45AF7" type="personal">
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<mods:namePart id="3CCBE2B98CAA3FBC776606DC3096E8C2">Goula, Katerina</mods:namePart>
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<mods:affiliation id="75463B11FE89F2FC22EAA95A818C1084">agoula @ biol. uoa. gr; https: // orcid. org / 0000 - 0001 - 9207 - 3570</mods:affiliation>
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<mods:nameIdentifier id="AD9B7EFB51CA709FC220FEB947C01C33" type="email">agoula@biol.uoa.gr</mods:nameIdentifier>
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<mods:namePart id="0111B6922C2FF31A5D25BD188568C076">Constantinidis, Theophanis</mods:namePart>
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<mods:affiliation id="442239DE9343F6EC45189590478FB916">constgr @ biol. uoa. gr; https: // orcid. org / 0000 - 0001 - 9704 - 3864</mods:affiliation>
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<mods:nameIdentifier id="47027624AFE9C7BA4AD044FF6DF7259D" type="email">constgr@biol.uoa.gr</mods:nameIdentifier>
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<mods:title id="4DD483911D500684626C7727817BF12C">Phytotaxa</mods:title>
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<mods:date id="96CB79A341EB561A6A977F5611217762">2021</mods:date>
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<mods:number id="57A6E3615D98D16EC51CBF40E9D514AC">2021-02-15</mods:number>
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<mods:number id="4A2E05459B86370BBBD89254EBC5D617">484</mods:number>
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<subSubSection id="C31B655EFFD6FFA5FF00F950FC59F8C5" box="[136,983,1772,1799]" pageId="8" pageNumber="137" type="nomenclature">
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<paragraph id="8BBE36D5FFD6FFA5FF00F950FC59F8C5" blockId="8.[136,983,1772,1799]" box="[136,983,1772,1799]" pageId="8" pageNumber="137">
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<heading id="D0F681B9FFD6FFA5FF00F950FC59F8C5" box="[136,983,1772,1799]" fontSize="11" level="2" pageId="8" pageNumber="137" reason="3">
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<taxonomicName id="4C014D56FFD6FFA5FF00F950FCDFF8C4" ID-CoL="7ZB49" authority="Biel & Kit Tan (2020: 190)" authorityName="Biel & Kit Tan" authorityPageNumber="190" authorityYear="2020" box="[136,849,1772,1799]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="137" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="runemarkii">
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<emphasis id="B975EAC7FFD6FFA5FF00F950FECCF8C5" bold="true" box="[136,322,1772,1799]" italics="true" pageId="8" pageNumber="137">Anthemis rigida</emphasis>
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subsp.
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<emphasis id="B975EAC7FFD6FFA5FE1FF950FD94F8C5" bold="true" box="[407,538,1772,1799]" italics="true" pageId="8" pageNumber="137">runemarkii</emphasis>
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Biel & Kit Tan (2020: 190)
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</taxonomicName>
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||
(
|
||
<figureCitation id="133A2A50FFD6FFA5FCE9F950FC28F8C5" box="[865,934,1772,1799]" captionStart="FIGURE 5" captionStartId="9.[136,229,1943,1965]" captionTargetBox="[152,1435,191,1917]" captionTargetId="figure-17@9.[151,1436,190,1918]" captionTargetPageId="9" captionText="FIGURE 5. Photographs of Anthemis rigida in the field (a, c, e, g) and in cultivation (b, d, f, h): A. rigida subsp. rigida from Kriti (a & b); A. rigida subsp. liguliflora from Kithira (c & d); A. rigida subsp. runemarkii from Milos (e & f); A. rigida subsp. antri-neptuni from Karpathos (g & h) (all photos taken by the first author except c, by Stavroula Fatsea)." figureDoi="http://doi.org/10.5281/zenodo.5421348" httpUri="https://zenodo.org/record/5421348/files/figure.png" pageId="8" pageNumber="137">Fig. 5</figureCitation>
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: e,f)
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31B655EFFD6FFA7FF00F884FB09F9C4" lastPageId="10" lastPageNumber="139" pageId="8" pageNumber="137" type="materials_examined">
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<paragraph id="8BBE36D5FFD6FFA5FF00F884FCA1F8B0" blockId="8.[136,1451,1848,1906]" pageId="8" pageNumber="137">
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<emphasis id="B975EAC7FFD6FFA5FF00F884FF34F88F" bold="true" box="[136,186,1848,1869]" pageId="8" pageNumber="137">Type</emphasis>
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:―
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<materialsCitation id="3B693C88FFD6FFA5FF5FF884FCA5F8B0" collectingDate="2020-02-25" collectionCode="C" country="Greece" elevation="10" latitude="36.683334" location="Kiklades" longLatPrecision="1186" longitude="24.416666" municipality="Milos" pageId="8" pageNumber="137" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="F3167645FFD6FFA5FF5FF884FEB4F88F" box="[215,314,1848,1869]" name="Greece" pageId="8" pageNumber="137">GREECE</collectingCountry>
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||
.
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||
<location id="8EDE600EFFD6FFA5FECEF884FE11F88C" LSID="urn:lsid:plazi:treatment:03A887C3FFD6FFA6FF00F950FB61FEC2:8EDE600EFFD6FFA5FECEF884FE11F88C" box="[326,415,1848,1870]" country="Greece" latitude="36.683334" longLatPrecision="1186" longitude="24.416666" municipality="Milos" name="Kiklades" pageId="8" pageNumber="137">Kiklades</location>
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||
: island of
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<collectingMunicipality id="6BDAACAFFFD6FFA5FD83F884FDCBF88C" box="[523,581,1848,1870]" pageId="8" pageNumber="137">Milos</collectingMunicipality>
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||
,
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||
<locationDeviation id="9A0C2FF7FFD6FFA5FDDBF884FC9BF88F" box="[595,789,1848,1870]" location="Ag. Mamas" pageId="8" pageNumber="137">NW of Ag. Mamas</locationDeviation>
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, rocky phrygana slopes above coast,
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<quantity id="4CF99B30FFD6FFA5FB06F884FB4AF88F" box="[1166,1220,1848,1869]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="137" unit="m" value="10.0">
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||
<elevation id="002CD1E6FFD6FFA5FB06F884FB4AF88F" box="[1166,1220,1848,1869]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="137" unit="m" value="10.0">10 m</elevation>
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||
</quantity>
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||
,
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||
<geoCoordinate id="EE355012FFD6FFA5FB59F884FAA8F88C" box="[1233,1318,1848,1870]" degrees="36" direction="north" minutes="41" orientation="latitude" pageId="8" pageNumber="137" precision="925" value="36.683334">36°41’N</geoCoordinate>
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,
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||
<geoCoordinate id="EE355012FFD6FFA5FABCF884FA08F88C" box="[1332,1414,1848,1870]" degrees="24" direction="east" minutes="25" orientation="longitude" pageId="8" pageNumber="137" precision="925" value="24.416666">24°25’E</geoCoordinate>
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,
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||
<date id="FFBF1015FFD6FFA5FA1BF884FEC2F8B3" pageId="8" pageNumber="137" value="2020-02-25">
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<collectingDate id="EFFBE9FDFFD6FFA5FA1BF884FEC2F8B3" pageId="8" pageNumber="137" value="2020-02-25">25 February 2020</collectingDate>
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||
</date>
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,
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<emphasis id="B975EAC7FFD6FFA5FEDEF8E0FE5FF8B3" box="[342,465,1884,1905]" italics="true" pageId="8" pageNumber="137">Biel 2020.48</emphasis>
|
||
(
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||
<typeStatus id="54BA8877FFD6FFA5FE55F8E0FDBDF8B0" box="[477,563,1884,1906]" pageId="8" pageNumber="137" type="holotype">holotype</typeStatus>
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||
<collectionCode id="ED10AE10FFD6FFA5FDB1F8E0FDC5F8B3" box="[569,587,1884,1905]" country="Denmark" name="University of Copenhagen" pageId="8" pageNumber="137" type="Herbarium">C</collectionCode>
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||
; see
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<bibRefCitation id="EF904B24FFD6FFA5FDF2F8E0FCAAF8B0" author="Biel, B. & Tan, K." box="[634,804,1884,1906]" pageId="8" pageNumber="137" pagination="187 - 216" refId="ref7018" refString="Biel, B. & Tan, K. (2020) Reports 12 - 15. In: Vladimirov, V., Aybeke, M. & Tan, K. (Eds.) New floristic records in the Balkans: 42. Phytologia Balcanica 26: 187 - 216." type="journal article" year="2020">Biel & Tan, 2020</bibRefCitation>
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||
)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BBE36D5FFD6FFA7FF00F81CFB09F9C4" blockId="8.[136,1452,1952,2087]" lastBlockId="10.[136,1452,1228,2083]" lastPageId="10" lastPageNumber="139" pageId="8" pageNumber="137">
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Small annual with prostrate to ascending, simple or sparingly branched stems,
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<quantity id="4CF99B30FFD6FFA5FC7EF81CFBD6F879" box="[1014,1112,1952,1979]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="2.0" metricValueMin="0.1" pageId="8" pageNumber="137" unit="cm" value="10.5" valueMax="20.0" valueMin="1.0">1−20 cm</quantity>
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long. Stems and leaves tinged with purple, densely tomentose with medifixed hairs when young, laxly tomentose to glabrescent later. Lower leaves long-petiolate, 1-pinnatisect with segments pinnatifid to pinnatisect, glandular-punctate; ultimate lobes at least 2 times longer than wide, oblong-obovate to oblong-ovate, acute to shortly mucronate; upper leaves similar but smaller. Peduncles thickened, becoming rigid and usually recurved in fruit. Capitula solitary, radiate. Involucre 5−8 ×
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<quantity id="4CF99B30FFD4FFA7FACFFB70FA26FB25" box="[1351,1448,1228,1255]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="10" pageNumber="139" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5−7 mm</quantity>
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, almost cylindrical, bracts usually in 2−3 rows, imbricate, unequal in length, tomentose when young, thick, rigid, and glabrescent in fruit, often tinged with purple. Outer involucral bracts narrowly lanceolate to ovate-lanceolate, 2.5−4 × ca.
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<quantity id="4CF99B30FFD4FFA7FF00FA85FF45FA91" box="[136,203,1337,1363]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="139" unit="mm" value="1.0">1 mm</quantity>
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, usually not more than 1/2−2/3 as long as inner bracts, with scarious margins up to
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<quantity id="4CF99B30FFD4FFA7FBEAFA85FB37FA91" box="[1122,1209,1337,1363]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="139" unit="mm" value="0.4">0.4 mm</quantity>
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. Inner bracts obovate, 5−6 × ca.
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<quantity id="4CF99B30FFD4FFA7FF7DFAE0FEC5FAB5" box="[245,331,1372,1399]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="139" unit="mm" value="1.5">1.5 mm</quantity>
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, with broad scarious margins up to
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<quantity id="4CF99B30FFD4FFA7FD45FAE0FCADFAB5" box="[717,803,1372,1399]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="139" unit="mm" value="0.5">0.5 mm</quantity>
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||
. Ligulate florets with ligules oblong-ovate, toothed at apex, 5−8 ×
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<quantity id="4CF99B30FFD4FFA7FF46FA3CFEDBFA59" box="[206,341,1408,1435]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="10" pageNumber="139" unit="mm" value="2.1" valueMax="2.5" valueMin="1.7">1.7−2.5 mm</quantity>
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||
, dark to pale pink, white tinged pink or white, glandular punctate, sterile; disc florets ca.
|
||
<quantity id="4CF99B30FFD4FFA7FA99FA3DFADCFA59" box="[1297,1362,1409,1435]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="139" unit="mm" value="3.0">3 mm</quantity>
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||
, yellow, glandular punctate. Anthers, style, and pollen yellow. Receptacular scales hyaline, obovate to oblanceolate, mucronate and becoming lacerate at apex. Achenes obconical, faintly ribbed; outer achenes ca. 2 ×
|
||
<quantity id="4CF99B30FFD4FFA7FBC9FA75FB0DFA21" box="[1089,1155,1481,1507]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="139" unit="mm" value="1.0">1 mm</quantity>
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||
, with a corona
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||
<quantity id="4CF99B30FFD4FFA7FAAFFA74FA25FA21" box="[1319,1451,1480,1507]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="10" pageNumber="139" unit="mm" value="0.7" valueMax="0.9" valueMin="0.5">0.5−0.9 mm</quantity>
|
||
long; inner achenes longer and thinner, 2−2.8 ×
|
||
<quantity id="4CF99B30FFD4FFA7FD1CFA50FC93F9C5" box="[660,797,1516,1543]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="10" pageNumber="139" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6−0.8 mm</quantity>
|
||
, with a corona
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||
<quantity id="4CF99B30FFD4FFA7FC4BFA51FBC6F9C5" box="[963,1096,1517,1543]" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="8.0" metricValueMin="4.0" pageId="10" pageNumber="139" unit="mm" value="0.6" valueMax="0.8" valueMin="0.4">0.4−0.8 mm</quantity>
|
||
long.
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||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7E665DFFD7FFA4FF00F82BFC32F837" ID-DOI="http://doi.org/10.5281/zenodo.5421348" ID-Zenodo-Dep="5421348" httpUri="https://zenodo.org/record/5421348/files/figure.png" pageId="9" pageNumber="138" startId="9.[136,229,1943,1965]" targetBox="[152,1435,191,1917]" targetPageId="9">
|
||
<paragraph id="8BBE36D5FFD7FFA4FF00F82BFC32F837" blockId="9.[136,1452,1943,2037]" pageId="9" pageNumber="138">
|
||
<emphasis id="B975EAC7FFD7FFA4FF00F82BFF70F86F" bold="true" box="[136,254,1943,1965]" pageId="9" pageNumber="138">FIGURE 5.</emphasis>
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||
Photographs of
|
||
<taxonomicName id="4C014D56FFD7FFA4FE16F82BFDB7F86E" box="[414,569,1943,1964]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="138" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B975EAC7FFD7FFA4FE16F82BFDB7F86E" box="[414,569,1943,1964]" italics="true" pageId="9" pageNumber="138">Anthemis rigida</emphasis>
|
||
</taxonomicName>
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||
in the field (a, c, e, g) and in cultivation (b, d, f, h):
|
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<taxonomicName id="4C014D56FFD7FFA4FBBDF824FA9EF86E" authority="subsp. rigida" box="[1077,1296,1943,1965]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="138" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="rigida">
|
||
<emphasis id="B975EAC7FFD7FFA4FBBDF824FB02F86E" box="[1077,1164,1943,1965]" italics="true" pageId="9" pageNumber="138">A. rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD7FFA4FB5DF82BFA9EF86E" box="[1237,1296,1943,1964]" italics="true" pageId="9" pageNumber="138">rigida</emphasis>
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||
</taxonomicName>
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from Kriti (a & b);
|
||
<taxonomicName id="4C014D56FFD7FFA4FF21F800FE24F812" authorityName="Greuter" authorityYear="1968" baseAuthorityName="Halacsy" baseAuthorityYear="1908" box="[169,426,1979,2001]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="138" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="liguliflora">
|
||
<emphasis id="B975EAC7FFD7FFA4FF21F800FF71F812" box="[169,255,1979,2001]" italics="true" pageId="9" pageNumber="138">A. rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD7FFA4FEC1F807FE24F812" box="[329,426,1979,2000]" italics="true" pageId="9" pageNumber="138">liguliflora</emphasis>
|
||
</taxonomicName>
|
||
from Kithira (c & d);
|
||
<taxonomicName id="4C014D56FFD7FFA4FD0CF800FC01F812" authorityName="Biel & Kit Tan" authorityYear="2020" box="[644,911,1979,2001]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="138" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="runemarkii">
|
||
<emphasis id="B975EAC7FFD7FFA4FD0CF800FD54F812" box="[644,730,1979,2001]" italics="true" pageId="9" pageNumber="138">A. rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD7FFA4FCACF807FC01F812" box="[804,911,1979,2000]" italics="true" pageId="9" pageNumber="138">runemarkii</emphasis>
|
||
</taxonomicName>
|
||
from Milos (e & f);
|
||
<taxonomicName id="4C014D56FFD7FFA4FBD0F800FB20F812" box="[1112,1198,1979,2001]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="138" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B975EAC7FFD7FFA4FBD0F800FB20F812" box="[1112,1198,1979,2001]" italics="true" pageId="9" pageNumber="138">A. rigida</emphasis>
|
||
</taxonomicName>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD7FFA4FB70F800FAF9F813" box="[1272,1399,1980,2001]" italics="true" pageId="9" pageNumber="138">antri-neptuni</emphasis>
|
||
from Karpathos (g & h) (all photos taken by the first author except c, by Stavroula Fatsea).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7E665DFFD4FFA7FF00FBFCFD68FB5C" ID-DOI="http://doi.org/10.5281/zenodo.5421350" ID-Zenodo-Dep="5421350" httpUri="https://zenodo.org/record/5421350/files/figure.png" pageId="10" pageNumber="139" startId="10.[136,229,1088,1110]" targetBox="[152,1435,191,1063]" targetPageId="10">
|
||
<paragraph id="8BBE36D5FFD4FFA7FF00FBFCFD68FB5C" blockId="10.[136,1452,1088,1182]" pageId="10" pageNumber="139">
|
||
<emphasis id="B975EAC7FFD4FFA7FF00FBFCFE8FFB97" bold="true" box="[136,257,1088,1110]" pageId="10" pageNumber="139">FIGURE 6.</emphasis>
|
||
Current known distribution of
|
||
<taxonomicName id="4C014D56FFD4FFA7FDB5FBFCFC49FB97" authorityName="Greuter" authorityYear="1968" baseAuthorityName="Greuter & Rechinger" box="[573,967,1088,1110]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="ammanthiformis">
|
||
<emphasis id="B975EAC7FFD4FFA7FDB5FBFCFD55FB97" box="[573,731,1088,1109]" italics="true" pageId="10" pageNumber="139">Anthemis rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD4FFA7FCA2FBFCFC49FB97" box="[810,967,1088,1109]" italics="true" pageId="10" pageNumber="139">ammanthiformis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C014D56FFD4FFA7FC5EFBFCFBA1FB97" box="[982,1071,1088,1109]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="139" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B975EAC7FFD4FFA7FC5EFBFCFBA1FB97" box="[982,1071,1088,1109]" italics="true" pageId="10" pageNumber="139">A. rigida</emphasis>
|
||
</taxonomicName>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD4FFA7FBF7FBFDFB70FB94" box="[1151,1278,1089,1110]" italics="true" pageId="10" pageNumber="139">antri-neptuni</emphasis>
|
||
,
|
||
<taxonomicName id="4C014D56FFD4FFA7FA85FBFCFF67FBBB" authorityName="Greuter" authorityYear="1968" baseAuthorityName="Halacsy" baseAuthorityYear="1908" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="liguliflora">
|
||
<emphasis id="B975EAC7FFD4FFA7FA85FBFCFAE8FB97" box="[1293,1382,1088,1109]" italics="true" pageId="10" pageNumber="139">A. rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD4FFA7FF00FBD8FF67FBBB" box="[136,233,1124,1145]" italics="true" pageId="10" pageNumber="139">liguliflora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C014D56FFD4FFA7FE95FBD8FDA1FBBB" authorityName="Biel & Kit Tan" authorityYear="2020" box="[285,559,1124,1146]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="runemarkii">
|
||
<emphasis id="B975EAC7FFD4FFA7FE95FBD8FEF8FBBB" box="[285,374,1124,1145]" italics="true" pageId="10" pageNumber="139">A. rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD4FFA7FE4CFBD8FDA1FBBB" box="[452,559,1124,1145]" italics="true" pageId="10" pageNumber="139">runemarkii</emphasis>
|
||
</taxonomicName>
|
||
in the three phytogeographical regions of south Greece. For a detailed distribution of the widespread
|
||
<taxonomicName id="4C014D56FFD4FFA7FF73FB34FE5AFB5F" authority="subsp. rigida" box="[251,468,1160,1182]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="rigida">
|
||
<emphasis id="B975EAC7FFD4FFA7FF73FB34FEDFFB5F" box="[251,337,1160,1181]" italics="true" pageId="10" pageNumber="139">A. rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD4FFA7FE11FB34FE5AFB5F" box="[409,468,1160,1181]" italics="true" pageId="10" pageNumber="139">rigida</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="EF904B24FFD4FFA7FE77FB34FDF5FB5C" author="Strid, A." box="[511,635,1160,1182]" pageId="10" pageNumber="139" pagination="1 - 878" refId="ref8289" refString="Strid, A. (2016 b) Atlas of the Aegean flora, Part 2: Maps. Englera 33: 1 - 878. [Berlin: Botanic Garden and Botanical Museum Berlin, Freie Universitat Berlin]" type="journal article" year="2016">Strid (2016b</bibRefCitation>
|
||
: map 413).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31B655EFFD4FFA6FF35F9ACFB61FEC2" lastPageId="11" lastPageNumber="140" pageId="10" pageNumber="139" type="distribution">
|
||
<paragraph id="8BBE36D5FFD4FFA7FF35F9ACFB91F954" blockId="10.[136,1452,1228,2083]" pageId="10" pageNumber="139">
|
||
<emphasis id="B975EAC7FFD4FFA7FF35F9ACFE6FF9E8" bold="true" box="[189,481,1552,1579]" pageId="10" pageNumber="139">Distribution and habitat</emphasis>
|
||
:―This subspecies is endemic to Milos island group and has been located so far on the following islands and islets: Milos, Kimolos, Prasonisi, Poliegos, at elevations from sea level to ca. 87 meters. It has been found in a number of different habitats such as sandy shores, rocky and gravelly places next to the sea, phrygana formations, maquis and periodically flooded areas, all on acid volcanic substrates.
|
||
</paragraph>
|
||
<paragraph id="8BBE36D5FFD4FFA7FF35F91DFD69F7E0" blockId="10.[136,1452,1228,2083]" pageId="10" pageNumber="139">
|
||
<emphasis id="B975EAC7FFD4FFA7FF35F91DFE8EF979" bold="true" box="[189,256,1697,1723]" pageId="10" pageNumber="139">Notes</emphasis>
|
||
:―Although the pink-coloured ligules is a remarkable trait observed in this subspecies, it is not characteristic of all the known populations. Ligules in various shades of pink, from deep- to pale-pink, or white tinged pink, or totally white have been recorded, even among individuals from the same locality. The morphological features that separate
|
||
<taxonomicName id="4C014D56FFD4FFA7FF00F8B0FD85F8E4" authorityName="Biel & Kit Tan" authorityYear="2020" box="[136,523,1804,1830]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="runemarkii">
|
||
<emphasis id="B975EAC7FFD4FFA7FF00F8B0FEB5F8E4" box="[136,315,1804,1830]" italics="true" pageId="10" pageNumber="139">Anthemis rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD4FFA7FE07F8B0FD85F8E4" box="[399,523,1804,1830]" italics="true" pageId="10" pageNumber="139">runemarkii</emphasis>
|
||
</taxonomicName>
|
||
from the rest of the ligulate subspecies are mainly the narrow leaf lobes, the longer and wider involucre and the broad scarious margin of the outer involucral bracts (
|
||
<figureCitation id="133A2A50FFD4FFA7FBBAF88CFBF6F889" box="[1074,1144,1840,1867]" captionStart="FIGURE 5" captionStartId="9.[136,229,1943,1965]" captionTargetBox="[152,1435,191,1917]" captionTargetId="figure-17@9.[151,1436,190,1918]" captionTargetPageId="9" captionText="FIGURE 5. Photographs of Anthemis rigida in the field (a, c, e, g) and in cultivation (b, d, f, h): A. rigida subsp. rigida from Kriti (a & b); A. rigida subsp. liguliflora from Kithira (c & d); A. rigida subsp. runemarkii from Milos (e & f); A. rigida subsp. antri-neptuni from Karpathos (g & h) (all photos taken by the first author except c, by Stavroula Fatsea)." figureDoi="http://doi.org/10.5281/zenodo.5421348" httpUri="https://zenodo.org/record/5421348/files/figure.png" pageId="10" pageNumber="139">Fig. 5</figureCitation>
|
||
,
|
||
<tableCitation id="C683036EFFD4FFA7FB0DF88CFB52F889" box="[1157,1244,1840,1867]" captionStart="TABLE 5" captionStartId="7.[136,224,1534,1558]" captionText="TABLE 5. Comparison of morphological features among the five subspecies of Anthemis rigida in the Aegean. Uncommon character values are in parentheses. All measurements in mm." pageId="10" pageNumber="139">Table 5</tableCitation>
|
||
). The “adpressedpubescent” achenes that Biel & Kit Tan (2020) mention in their description raises serious questions about achene morphology for the entire genus: the only report of hairy achenes among the
|
||
<taxonomicName id="4C014D56FFD4FFA7FC67F8C4FB3EF850" box="[1007,1200,1912,1938]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="139" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B975EAC7FFD4FFA7FC67F8C4FBD9F850" box="[1007,1111,1912,1938]" italics="true" pageId="10" pageNumber="139">Anthemis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of Europe (
|
||
<bibRefCitation id="EF904B24FFD4FFA7FABEF8C4FF4AF875" author="Fernandes, R." pageId="10" pageNumber="139" pagination="145 - 159" refId="ref7207" refString="Fernandes, R. (1976) Anthemis L. In: Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A. (Eds.) Flora Europaea 4. Cambridge University Press, Cambridge, pp. 145 - 159." type="book chapter" year="1976">Fernandes 1976</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F3167645FFD4FFA7FF5EF820FEA8F874" box="[214,294,1948,1974]" name="Turkey" pageId="10" pageNumber="139">Turkey</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF904B24FFD4FFA7FEBBF820FDB4F875" author="Grierson, A. J. C. & Yavin, Z." box="[307,570,1948,1975]" pageId="10" pageNumber="139" pagination="174 - 221" refId="ref7540" refString="Grierson, A. J. C. & Yavin, Z. (1975) Anthemis L. In: Davis, P. H. (Ed.) Flora of Turkey and the East Aegean islands. Edinburgh University Press, Edinburgh, pp. 174 - 221." type="book chapter" year="1975">Grierson & Yavin 1975</bibRefCitation>
|
||
), N Africa (
|
||
<bibRefCitation id="EF904B24FFD4FFA7FD32F820FCF0F875" author="Oberprieler, C." box="[698,894,1948,1975]" pageId="10" pageNumber="139" pagination="5 - 328" refId="ref7787" refString="Oberprieler, C. (1998) The systematics of Anthemis L. (Compositae, Anthemideae) in W and C North Africa. Bocconea 9: 5 - 328." type="journal article" year="1998">Oberprieler 1998</bibRefCitation>
|
||
), Middle East (
|
||
<bibRefCitation id="EF904B24FFD4FFA7FBAEF820FB01F875" author="Eig, A." box="[1062,1167,1948,1975]" pageId="10" pageNumber="139" pagination="161 - 224" refId="ref7179" refString="Eig, A. (1938) Taxonomic studies on the oriental species of the genus Anthemis. Palestine Journal of Botany 1: 161 - 224." type="journal article" year="1938">Eig 1938</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF904B24FFD4FFA7FB12F820FACAF874" author="Iranshahr, M." box="[1178,1348,1948,1974]" pageId="10" pageNumber="139" pagination="5 - 44" refId="ref7615" refString="Iranshahr, M. (1986) Anthemis. In: Rechinger, K. H. (Ed.) Flora Iranica 158, Compositae: VI Anthemideae. Akademische Druck und Verlagsanstalt, Graz, pp. 5 - 44." type="book chapter" year="1986">Iranshahr 1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF904B24FFD4FFA7FAC6F820FE89F819" author="Ghafoor, A. & Ali, S. J." pageId="10" pageNumber="139" pagination="1 - 41" refId="ref7458" refString="Ghafoor, A. & Ali, S. J. (2002) The genus Anthemis L. (Compositae) in Iraq: a synopsis. Compositae Newsletter 38: 1 - 41." type="journal article" year="2002">Ghafoor & Ali 2002</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="F3167645FFD4FFA7FECBF87CFE5DF818" box="[323,467,1984,2010]" name="Saudi Arabia" pageId="10" pageNumber="139">Saudi Arabia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF904B24FFD4FFA7FE69F87CFD0FF819" author="Ghafoor, A." box="[481,641,1984,2011]" pageId="10" pageNumber="139" pagination="79 - 98" refId="ref7425" refString="Ghafoor, A. (2010) The genus Anthemis L. (Compositae-Anthemideae) in Arabian Peninsula: a taxonomic study. Pakistan Journal of Botany 42: 79 - 98." type="journal article" year="2010">Ghafoor 2010</bibRefCitation>
|
||
) concerns “a single poor specimen with somewhat hairy young achenes” that
|
||
<bibRefCitation id="EF904B24FFD4FFA7FF33F858FEB8F83D" author="Eig, A." box="[187,310,2020,2047]" pageId="10" pageNumber="139" pagination="161 - 224" refId="ref7179" refString="Eig, A. (1938) Taxonomic studies on the oriental species of the genus Anthemis. Palestine Journal of Botany 1: 161 - 224." type="journal article" year="1938">Eig (1938)</bibRefCitation>
|
||
mentions from Ramadi,
|
||
<collectingCountry id="F3167645FFD4FFA7FDDFF859FD06F83D" box="[599,648,2021,2047]" name="Iraq" pageId="10" pageNumber="139">Iraq</collectingCountry>
|
||
. A thorough examination of achenes in all the specimens collected on Milos do not indicate the presence of any pubescence.
|
||
</paragraph>
|
||
<paragraph id="8BBE36D5FFD5FFA6FF35FF22FB61FEC2" blockId="11.[136,1451,158,257]" pageId="11" pageNumber="140">
|
||
As previously mentioned,
|
||
<taxonomicName id="4C014D56FFD5FFA6FE60FF22FCE1FF7A" authorityName="Biel & Kit Tan" authorityYear="2020" box="[488,879,158,184]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="140" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="runemarkii">
|
||
<emphasis id="B975EAC7FFD5FFA6FE60FF22FD12FF7A" box="[488,668,158,184]" italics="true" pageId="11" pageNumber="140">Anthemis rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD5FFA6FD7BFF22FCE1FF7A" box="[755,879,158,184]" italics="true" pageId="11" pageNumber="140">runemarkii</emphasis>
|
||
</taxonomicName>
|
||
co-exist with
|
||
<taxonomicName id="4C014D56FFD5FFA6FB9AFF23FA9CFF7A" authority="subsp. rigida" box="[1042,1298,158,185]" class="Magnoliopsida" family="Asteraceae" genus="Anthemis" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="140" phylum="Tracheophyta" rank="subSpecies" species="rigida" subSpecies="rigida">
|
||
<emphasis id="B975EAC7FFD5FFA6FB9AFF23FBF8FF7A" box="[1042,1142,158,185]" italics="true" pageId="11" pageNumber="140">A. rigida</emphasis>
|
||
subsp.
|
||
<emphasis id="B975EAC7FFD5FFA6FB46FF22FA9CFF7A" box="[1230,1298,158,184]" italics="true" pageId="11" pageNumber="140">rigida</emphasis>
|
||
</taxonomicName>
|
||
on the island of Milos. In the areas where the two taxa meet, subpopulations with intermediate features have been recorded. Similar cases are reported by
|
||
<bibRefCitation id="EF904B24FFD5FFA6FEF3FF5AFD37FEC3" author="Greuter, W. & Rechinger, K. H." box="[379,697,230,257]" pageId="11" pageNumber="140" pagination="22 - 196" refId="ref7516" refString="Greuter, W. & Rechinger, K. H. (1967) Chloris Kythereia. Boissiera 13: 22 - 196." type="journal article" year="1967">Greuter & Rechinger (1967)</bibRefCitation>
|
||
for populations on Kithira and Antikithira islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |