180 lines
34 KiB
XML
180 lines
34 KiB
XML
<document id="7B85EB60C1F3D19E38F3EEE9CB4A6D7C" ID-CLB-Dataset="24572" ID-DOI="10.11646/zootaxa.4712.3.7" ID-GBIF-Dataset="4482b4cb-e1f0-48a8-a6e5-b5fda6e93bc9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3586422" ID-ZooBank="80BBA2AF-DDE8-456E-B983-3DA4F66A7181" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576823180524" checkinUser="plazi" docAuthor="Vandenberg, Natalia J." docDate="2019" docId="03A887B78238FE1AFF79A6029335FAB3" docLanguage="en" docName="zootaxa.4712.3.7.pdf" docOrigin="Zootaxa 4712 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Argosadalia Vandenberg 2019, gen. nov." docType="treatment" docVersion="7" lastPageNumber="418" masterDocId="FF91FFCF823CFE1FFFEEA15A9030FF80" masterDocTitle="A new monotypic genus and new species of lady beetle (Coleoptera: Coccinellidae Coccinellini) from western South America" masterLastPageNumber="422" masterPageNumber="413" pageNumber="417" updateTime="1698774946031" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5DDED7B5D2B0EA8982B947E3FA74AFF9">A new monotypic genus and new species of lady beetle (Coleoptera: Coccinellidae Coccinellini) from western South America</mods:title>
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<mods:namePart id="A97459928042F363F22B0822917734A5">Vandenberg, Natalia J.</mods:namePart>
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<mods:date id="40A851B5CF2B8E2A0C1A464FE9BDA0C1">2019</mods:date>
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<treatment id="03A887B78238FE1AFF79A6029335FAB3" ID-DOI="http://doi.org/10.5281/zenodo.5944550" ID-GBIF-Taxon="160797306" ID-Zenodo-Dep="5944550" LSID="urn:lsid:plazi:treatment:03A887B78238FE1AFF79A6029335FAB3" httpUri="http://treatment.plazi.org/id/03A887B78238FE1AFF79A6029335FAB3" lastPageId="5" lastPageNumber="418" pageId="4" pageNumber="417">
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<subSubSection id="C31B652A8238FE1BFF79A60291AFF8F3" box="[151,415,1880,1907]" pageId="4" pageNumber="417" type="nomenclature">
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<paragraph id="8BBE36A18238FE1BFF79A60291AFF8F3" blockId="4.[151,438,1880,1943]" box="[151,415,1880,1907]" pageId="4" pageNumber="417">
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<heading id="D0F681CD8238FE1BFF79A60291AFF8F3" bold="true" box="[151,415,1880,1907]" fontSize="11" level="1" pageId="4" pageNumber="417" reason="1">
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<emphasis id="B975EAB38238FE1BFF79A60291AFF8F3" bold="true" box="[151,415,1880,1907]" pageId="4" pageNumber="417">
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<taxonomicName id="4C014D228238FE1BFF79A602911AF8F3" authority="Vandenberg, 2019" authorityName="Vandenberg" authorityYear="2019" box="[151,298,1880,1907]" class="Insecta" family="Coccinellidae" genus="Argosadalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="417" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B975EAB38238FE1BFF79A602911AF8F3" bold="true" box="[151,298,1880,1907]" italics="true" pageId="4" pageNumber="417">Argosadalia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24657C88238FE1BFEDFA60391AFF8F3" box="[305,415,1881,1907]" pageId="4" pageNumber="417" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B652A8238FE1BFF79A62792AAF85F" pageId="4" pageNumber="417" type="description">
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<paragraph id="8BBE36A18238FE1BFF79A6279186F817" blockId="4.[151,438,1880,1943]" box="[151,438,1916,1943]" pageId="4" pageNumber="417">
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(
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<figureCitation id="133A2A248238FE1BFF71A627913BF817" box="[159,267,1917,1943]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1858,1883]" captionTargetBox="[155,1435,434,1797]" captionTargetId="figure@2.[151,1435,414,1830]" captionTargetPageId="2" captionText="FIGURES 1–10. External morphology of lady beetles in the tribe Coccinellini. 1–4, Argosadalia priscilla sp. nov. habitus illustrations: 1–3, holotype, male (scale bar lower left): 1, dorsal; 2, lateral; 3, frontal; 4, paratype, female (scale bar upper right), dorsal. 5–6, A. priscilla sp. nov., ventral view of mouthparts: 5, labial prementum and palpi; 6, maxillary palp. 7–10, abdominal postcoxal lines: 7, A. priscilla sp. nov.; 8, Adalia bipunctata L., 1758; 9, Neoharmonia venusta (Melsheimer, 1847); 10, Coccinella sp." figureDoi="http://doi.org/10.5281/zenodo.3586424" httpUri="https://zenodo.org/record/3586424/files/figure.png" pageId="4" pageNumber="417">Figs. 1–7</figureCitation>
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,
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<figureCitation id="133A2A248238FE1BFEF9A6279100F817" box="[279,304,1917,1943]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="4" pageNumber="417">11</figureCitation>
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a–e, 15–18)
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</paragraph>
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<paragraph id="8BBE36A18238FE1BFF79A69F92AAF85F" blockId="4.[151,666,1989,2015]" box="[151,666,1989,2015]" pageId="4" pageNumber="417">
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<emphasis id="B975EAB38238FE1BFF79A69F9104F85F" bold="true" box="[151,308,1989,2015]" pageId="4" pageNumber="417">
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<typeStatus id="54BA88038238FE1BFF79A69F90FDF85F" box="[151,205,1989,2015]" pageId="4" pageNumber="417">type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C014D228238FE1BFED2A69F9205F85F" authority="Vandenberg, 2019" authorityName="Vandenberg" authorityYear="2019" box="[316,565,1989,2015]" class="Insecta" family="Coccinellidae" genus="Argosadalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="417" phylum="Arthropoda" rank="species" species="priscilla" status="sp. nov.">
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<emphasis id="B975EAB38238FE1BFED2A69F9205F85F" box="[316,565,1989,2015]" italics="true" pageId="4" pageNumber="417">Argosadalia priscilla</emphasis>
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</taxonomicName>
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<emphasis id="B975EAB38238FE1BFDD2A69F92AAF85F" bold="true" box="[572,666,1989,2015]" pageId="4" pageNumber="417">
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<taxonomicNameLabel id="A24657C88238FE1BFDD2A69F92AAF85F" box="[572,666,1989,2015]" pageId="4" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B652A8239FE1AFF79A1C29221FE77" pageId="5" pageNumber="418" type="diagnosis">
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<paragraph id="8BBE36A18239FE1AFF79A1C29221FE77" blockId="5.[151,1437,152,1331]" pageId="5" pageNumber="418">
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<emphasis id="B975EAB38239FE1AFF79A1C29122FF33" bold="true" box="[151,274,152,179]" pageId="5" pageNumber="418">Diagnosis.</emphasis>
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Distinguished from other members of the
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<taxonomicName id="4C014D228239FE1AFD04A1C39315FF33" authorityName="Mulsant" authorityYear="1850" box="[746,805,153,179]" class="Insecta" family="Chrysopidae" genus="Neda" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
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<emphasis id="B975EAB38239FE1AFD04A1C39315FF33" box="[746,805,153,179]" italics="true" pageId="5" pageNumber="418">Neda</emphasis>
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</taxonomicName>
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group by a combination of: body oval, small-sized, with length
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<quantity id="4CF99B448239FE1AFF08A1E6910EFF57" box="[230,318,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="418" unit="mm" value="5.2">5.2 mm</quantity>
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or less, elytral margins narrowly but distinctly explanate, with marginal bead (
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<figureCitation id="133A2A248239FE1AFB25A1E79508FF57" box="[1227,1336,189,215]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1858,1883]" captionTargetBox="[155,1435,434,1797]" captionTargetId="figure@2.[151,1435,414,1830]" captionTargetPageId="2" captionText="FIGURES 1–10. External morphology of lady beetles in the tribe Coccinellini. 1–4, Argosadalia priscilla sp. nov. habitus illustrations: 1–3, holotype, male (scale bar lower left): 1, dorsal; 2, lateral; 3, frontal; 4, paratype, female (scale bar upper right), dorsal. 5–6, A. priscilla sp. nov., ventral view of mouthparts: 5, labial prementum and palpi; 6, maxillary palp. 7–10, abdominal postcoxal lines: 7, A. priscilla sp. nov.; 8, Adalia bipunctata L., 1758; 9, Neoharmonia venusta (Melsheimer, 1847); 10, Coccinella sp." figureDoi="http://doi.org/10.5281/zenodo.3586424" httpUri="https://zenodo.org/record/3586424/files/figure.png" pageId="5" pageNumber="418">Figs. 1–4</figureCitation>
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,
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<figureCitation id="133A2A248239FE1AFAA8A1E795BFFF58" box="[1350,1423,189,216]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">11a, d</figureCitation>
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). Prosternal intercoxal process lacking carinae. Elytral epipleuron (
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<figureCitation id="133A2A248239FE1AFC93A1BB93C4FF7C" box="[893,1012,225,252]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">Fig 11b, d</figureCitation>
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) narrow, with maximum width about 1/7 width of body, subhorizontal in anterior half, planar to weakly concave. Hind margin of abdominal ventrites 1–4 (inclusive of flexure bands) sublinear when abdomen is held flat (
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<figureCitation id="133A2A248239FE1AFC67A0739422FEC3" box="[905,1042,297,323]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">Figs. 11b, c</figureCitation>
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); postcoxal line of first abdominal ventrite joined to oblique line and obliterated beyond, forming modified V shape (
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<figureCitation id="133A2A248239FE1AFBD8A017944BFEE7" box="[1078,1147,333,359]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1858,1883]" captionTargetBox="[155,1435,434,1797]" captionTargetId="figure@2.[151,1435,414,1830]" captionTargetPageId="2" captionText="FIGURES 1–10. External morphology of lady beetles in the tribe Coccinellini. 1–4, Argosadalia priscilla sp. nov. habitus illustrations: 1–3, holotype, male (scale bar lower left): 1, dorsal; 2, lateral; 3, frontal; 4, paratype, female (scale bar upper right), dorsal. 5–6, A. priscilla sp. nov., ventral view of mouthparts: 5, labial prementum and palpi; 6, maxillary palp. 7–10, abdominal postcoxal lines: 7, A. priscilla sp. nov.; 8, Adalia bipunctata L., 1758; 9, Neoharmonia venusta (Melsheimer, 1847); 10, Coccinella sp." figureDoi="http://doi.org/10.5281/zenodo.3586424" httpUri="https://zenodo.org/record/3586424/files/figure.png" pageId="5" pageNumber="418">Fig. 7</figureCitation>
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,
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<figureCitation id="133A2A248239FE1AFB66A0179484FEE8" box="[1160,1204,333,360]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">11b</figureCitation>
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). Basal lobe of male genitalia widest before apex, apically bifurcate, terminating in pair of parallel tapered projections separated by Ushaped emargination (
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<figureCitation id="133A2A248239FE1AFE60A0CF91DBFE2F" box="[398,491,405,431]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">Fig. 11e</figureCitation>
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,
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<figureCitation id="133A2A248239FE1AFE19A0CF9226FE30" box="[503,534,405,432]" captionStart="FIGURES 15–19" captionStartId="4.[151,264,1666,1691]" captionTargetBox="[155,1435,190,1640]" captionTargetId="figure@4.[151,1435,181,1640]" captionTargetPageId="4" captionText="FIGURES 15–19. Genitalia of lady beetles in the Neda-group. 15–17, male genitalia of Argosadalia priscilla sp. nov.: 15, left lateral view of penis; 16, ventral view of phallobase; 17, right lateral view of phallobase. 18–19, Female genitalia in dorsal view, arrows to the right of each figure show the outer angle where the cornu joins the nodulus: 18, A. priscilla sp. nov.; 19, Neoharmonia venusta Melsheimer, 1847." figureDoi="http://doi.org/10.5281/zenodo.3586428" httpUri="https://zenodo.org/record/3586428/files/figure.png" pageId="5" pageNumber="418">16</figureCitation>
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). Spermatheca of female genitalia (
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<figureCitation id="133A2A248239FE1AFC4CA0CF93C3FE2F" box="[930,1011,405,431]" captionStart="FIGURES 15–19" captionStartId="4.[151,264,1666,1691]" captionTargetBox="[155,1435,190,1640]" captionTargetId="figure@4.[151,1435,181,1640]" captionTargetPageId="4" captionText="FIGURES 15–19. Genitalia of lady beetles in the Neda-group. 15–17, male genitalia of Argosadalia priscilla sp. nov.: 15, left lateral view of penis; 16, ventral view of phallobase; 17, right lateral view of phallobase. 18–19, Female genitalia in dorsal view, arrows to the right of each figure show the outer angle where the cornu joins the nodulus: 18, A. priscilla sp. nov.; 19, Neoharmonia venusta Melsheimer, 1847." figureDoi="http://doi.org/10.5281/zenodo.3586428" httpUri="https://zenodo.org/record/3586428/files/figure.png" pageId="5" pageNumber="418">Fig. 18</figureCitation>
|
||
) with nodulus joined to cornu to form distinct outer angle, shaped like the number “2,” but with apex of nodulus tapered distally, slightly down-curved at point of attachment to spermduct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B652A8239FE1AFF29A35B9467FBFF" pageId="5" pageNumber="418" type="discussion">
|
||
<paragraph id="8BBE36A18239FE1AFF29A35B93ADFD2B" blockId="5.[151,1437,152,1331]" pageId="5" pageNumber="418">
|
||
<emphasis id="B975EAB38239FE1AFF29A35B910BFD9B" bold="true" box="[199,315,513,539]" pageId="5" pageNumber="418">Remarks.</emphasis>
|
||
Within the
|
||
<taxonomicName id="4C014D228239FE1AFE53A35B91C8FD9B" authorityName="Mulsant" authorityYear="1850" box="[445,504,513,539]" class="Insecta" family="Chrysopidae" genus="Neda" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFE53A35B91C8FD9B" box="[445,504,513,539]" italics="true" pageId="5" pageNumber="418">Neda</emphasis>
|
||
</taxonomicName>
|
||
-group the new genus resembles some of the smaller members of
|
||
<taxonomicName id="4C014D228239FE1AFB3FA35B955BFD9B" authorityName="Crotch" authorityYear="1871" box="[1233,1387,513,539]" class="Insecta" family="Coccinellidae" genus="Neoharmonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFB3FA35B955BFD9B" box="[1233,1387,513,539]" italics="true" pageId="5" pageNumber="418">Neoharmonia</emphasis>
|
||
</taxonomicName>
|
||
, but the latter have the prosternal intercoxal process with a pair of convergent carinae separated by a triangular depression, abdominal postcoxal line continued laterally beyond the juncture with the oblique line (
|
||
<figureCitation id="133A2A248239FE1AFB5FA31394C6FDE3" box="[1201,1270,585,611]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1858,1883]" captionTargetBox="[155,1435,434,1797]" captionTargetId="figure@2.[151,1435,414,1830]" captionTargetPageId="2" captionText="FIGURES 1–10. External morphology of lady beetles in the tribe Coccinellini. 1–4, Argosadalia priscilla sp. nov. habitus illustrations: 1–3, holotype, male (scale bar lower left): 1, dorsal; 2, lateral; 3, frontal; 4, paratype, female (scale bar upper right), dorsal. 5–6, A. priscilla sp. nov., ventral view of mouthparts: 5, labial prementum and palpi; 6, maxillary palp. 7–10, abdominal postcoxal lines: 7, A. priscilla sp. nov.; 8, Adalia bipunctata L., 1758; 9, Neoharmonia venusta (Melsheimer, 1847); 10, Coccinella sp." figureDoi="http://doi.org/10.5281/zenodo.3586424" httpUri="https://zenodo.org/record/3586424/files/figure.png" pageId="5" pageNumber="418">Fig. 9</figureCitation>
|
||
), basal lobe of male genitalia with the apex undivided (
|
||
<figureCitation id="133A2A248239FE1AFDB8A3379285FD07" box="[598,693,621,647]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">Fig. 12e</figureCitation>
|
||
), and female spermatheca with a long straight tapered nodulus that flows directly from the cornu without a distinct outer angle (
|
||
<figureCitation id="133A2A248239FE1AFCD2A3CB93A0FD2B" box="[828,912,657,683]" captionStart="FIGURES 15–19" captionStartId="4.[151,264,1666,1691]" captionTargetBox="[155,1435,190,1640]" captionTargetId="figure@4.[151,1435,181,1640]" captionTargetPageId="4" captionText="FIGURES 15–19. Genitalia of lady beetles in the Neda-group. 15–17, male genitalia of Argosadalia priscilla sp. nov.: 15, left lateral view of penis; 16, ventral view of phallobase; 17, right lateral view of phallobase. 18–19, Female genitalia in dorsal view, arrows to the right of each figure show the outer angle where the cornu joins the nodulus: 18, A. priscilla sp. nov.; 19, Neoharmonia venusta Melsheimer, 1847." figureDoi="http://doi.org/10.5281/zenodo.3586428" httpUri="https://zenodo.org/record/3586428/files/figure.png" pageId="5" pageNumber="418">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBE36A18239FE1AFF29A3EF916EFC6F" blockId="5.[151,1437,152,1331]" pageId="5" pageNumber="418">
|
||
The genus
|
||
<taxonomicName id="4C014D228239FE1AFEAEA3EF914BFD4F" authorityName="Mulsant" authorityYear="1850" box="[320,379,693,719]" class="Insecta" family="Chrysopidae" genus="Neda" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFEAEA3EF914BFD4F" box="[320,379,693,719]" italics="true" pageId="5" pageNumber="418">Neda</emphasis>
|
||
</taxonomicName>
|
||
has the same form of the female spermathecal capsule as the new genus and is probably closely related. However, it can be easily distinguished (
|
||
<figureCitation id="133A2A248239FE1AFD23A383931AFD73" box="[717,810,729,755]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">Figs. 13</figureCitation>
|
||
a–e) by the larger body size (5.2–10.0 mm in length), broadly explanate elytral margin, epipleuron strongly concave and steeply descending externally, from about ¼–1/3 body width at widest part, hind margin of abdominal ventrite 4 distinctly arcuately emarginate in both sexes, and the basal lobe of the male genitalia widened apically, with projections diverging distally.
|
||
<taxonomicName id="4C014D228239FE1AFB66A21F9530FCDF" authorityName="Crotch" authorityYear="1874" box="[1160,1280,837,863]" class="Insecta" family="Coccinellidae" genus="Mononeda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFB66A21F9530FCDF" box="[1160,1280,837,863]" italics="true" pageId="5" pageNumber="418">Mononeda</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133A2A248239FE1AFAFEA21F955BFCDF" box="[1296,1387,837,863]" captionStart="FIGURES 11–14" captionStartId="3.[151,264,1834,1859]" captionTargetBox="[202,1386,186,1786]" captionTargetId="figure@3.[202,1386,181,1808]" captionTargetPageId="3" captionText="FIGURES 11–14. Genera of the Neda-group, key characters drawn from exemplars of the type species of each genus: a, body form in dorsal view, males; b, diagrammatic view of venter, males, elytral epipleuron shown in red, abdominal ventrites in white; c, apex of abdomen, female, arrow points to hind margin of 4th ventrite; d, shape of elytra in cross-sectional view through widest part; e, basal lobe of male genitalia in ventral view rescaled to the same length for comparative purposes. Scale bar applies to figures in rows a–d. 11, Argosadalia priscilla sp. nov.; 12, Neoharmonia venusta Melsheimer, 1847; 13, Neda norrisii (Guérin-Méneville, 1844); 14, Mononeda marginata (L., 1767)." figureDoi="http://doi.org/10.5281/zenodo.3586426" httpUri="https://zenodo.org/record/3586426/files/figure.png" pageId="5" pageNumber="418">Figs. 14</figureCitation>
|
||
a–e) also falls within the “
|
||
<taxonomicName id="4C014D228239FE1AFE62A23391F7FC03" authorityName="Mulsant" authorityYear="1850" box="[396,455,873,899]" class="Insecta" family="Chrysopidae" genus="Neda" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFE62A23391F7FC03" box="[396,455,873,899]" italics="true" pageId="5" pageNumber="418">Neda</emphasis>
|
||
</taxonomicName>
|
||
-group” and differs from the new genus by the much larger body size (length
|
||
<quantity id="4CF99B448239FE1AFAD4A23290F7FC27" metricMagnitude="-3" metricUnit="m" metricValue="9.95" metricValueMax="12.4" metricValueMin="7.5" pageId="5" pageNumber="418" unit="mm" value="9.95" valueMax="12.4" valueMin="7.5">7.5–12.4 mm</quantity>
|
||
), complete lack of an elytral marginal bead, elytral margin broadly explanate, and elytral epipleuron very wide, concave and descending externally, from about 1/3–1/2 body width at widest part, with ventral epipleural carina complete to apex.
|
||
</paragraph>
|
||
<paragraph id="8BBE36A18239FE1AFF29A2A39467FBFF" blockId="5.[151,1437,152,1331]" pageId="5" pageNumber="418">
|
||
The new genus superficially resembles the Holarctic genus
|
||
<taxonomicName id="4C014D228239FE1AFC97A2A393F4FB93" authorityName="Mulsant" authorityYear="1850" box="[889,964,1017,1043]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFC97A2A393F4FB93" box="[889,964,1017,1043]" italics="true" pageId="5" pageNumber="418">Adalia</emphasis>
|
||
</taxonomicName>
|
||
in its relatively small size, general body shape, and similar form of the abdominal postcoxal line (
|
||
<figureCitation id="133A2A248239FE1AFCFFA547934AFBB7" box="[785,890,1053,1079]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1858,1883]" captionTargetBox="[155,1435,434,1797]" captionTargetId="figure@2.[151,1435,414,1830]" captionTargetPageId="2" captionText="FIGURES 1–10. External morphology of lady beetles in the tribe Coccinellini. 1–4, Argosadalia priscilla sp. nov. habitus illustrations: 1–3, holotype, male (scale bar lower left): 1, dorsal; 2, lateral; 3, frontal; 4, paratype, female (scale bar upper right), dorsal. 5–6, A. priscilla sp. nov., ventral view of mouthparts: 5, labial prementum and palpi; 6, maxillary palp. 7–10, abdominal postcoxal lines: 7, A. priscilla sp. nov.; 8, Adalia bipunctata L., 1758; 9, Neoharmonia venusta (Melsheimer, 1847); 10, Coccinella sp." figureDoi="http://doi.org/10.5281/zenodo.3586424" httpUri="https://zenodo.org/record/3586424/files/figure.png" pageId="5" pageNumber="418">Figs. 7, 8</figureCitation>
|
||
), but
|
||
<taxonomicName id="4C014D228239FE1AFC54A5479435FBB7" authorityName="Mulsant" authorityYear="1850" box="[954,1029,1053,1079]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFC54A5479435FBB7" box="[954,1029,1053,1079]" italics="true" pageId="5" pageNumber="418">Adalia</emphasis>
|
||
</taxonomicName>
|
||
males have an undivided basal lobe, and a differently shaped penis with a strongly pigmented penis capsule.
|
||
<taxonomicName id="4C014D228239FE1AFC5AA51B93CFFBDB" authorityName="Mulsant" authorityYear="1850" box="[948,1023,1089,1115]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFC5AA51B93CFFBDB" box="[948,1023,1089,1115]" italics="true" pageId="5" pageNumber="418">Adalia</emphasis>
|
||
</taxonomicName>
|
||
females have a smaller spermathecal accessory gland and the spermduct with a more robust and heavily sclerotized sheath.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B652A8239FE1AFF29A5D39335FAB3" pageId="5" pageNumber="418" type="etymology">
|
||
<paragraph id="8BBE36A18239FE1AFF29A5D39335FAB3" blockId="5.[151,1437,152,1331]" pageId="5" pageNumber="418">
|
||
<emphasis id="B975EAB38239FE1AFF29A5D3917EFB23" bold="true" box="[199,334,1161,1187]" pageId="5" pageNumber="418">Etymology.</emphasis>
|
||
<taxonomicName id="4C014D228239FE1AFEB7A5D391D2FB23" authority="Vandenberg, 2019" authorityName="Vandenberg" authorityYear="2019" box="[345,482,1161,1187]" class="Insecta" family="Coccinellidae" genus="Argosadalia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="417" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B975EAB38239FE1AFEB7A5D391D2FB23" box="[345,482,1161,1187]" italics="true" pageId="5" pageNumber="418">Argosadalia</emphasis>
|
||
</taxonomicName>
|
||
(gender feminine) formed from a combination of Argos (= shining, bright, from Greek) +
|
||
<taxonomicName id="4C014D228239FE1AFEEFA5F7917CFB47" authorityName="Mulsant" authorityYear="1850" box="[257,332,1197,1223]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">Adalia</taxonomicName>
|
||
(= noble, from Hebrew). In Greek mythology, Argos was the name of the hundred-eyed servant of the goddess Hera, who had eyes all over his body. This name was selected because of the numerous white spots on the elytra which resemble eyes. The second part of the name was used because of the superficial similarity of this lady beetle to members of the lady beetle genus
|
||
<taxonomicName id="4C014D228239FE1AFD5AA44392CFFAB3" authorityName="Mulsant" authorityYear="1850" box="[692,767,1305,1331]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAB38239FE1AFD5AA44392CFFAB3" box="[692,767,1305,1331]" italics="true" pageId="5" pageNumber="418">Adalia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |