753 lines
82 KiB
XML
753 lines
82 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.938.49349" ID-GBIF-Dataset="1070bebf-afc9-40dd-a16e-24bd6821469d" ID-PMC="PMC7286948" ID-Pensoft-Pub="1313-2970-938-1" ID-Pensoft-UUID="A7E05382096E5182BD65560002045C47" ID-PubMed="32549744" ID-ZooBank="76B7FF03FEB14884AD2955BE864F0EBF" ModsDocID="1313-2970-938-1" checkinTime="1591212709532" checkinUser="pensoft" docAuthor="Neal, Lenka, Paterson, Gordon L. J., Blockley, David, Scott, Ben, Sherlock, Emma, Huque, Cate & Glover, Adrian G." docDate="2020" docId="D4382279AE655258BC05E2AAA5D4AD57" docLanguage="en" docName="ZooKeys 938: 1-86" docOrigin="ZooKeys 938" docSource="http://dx.doi.org/10.3897/zookeys.938.49349" docTitle="Prosphaerosyllis modinouae Neal & Paterson 2020, sp. nov." docType="treatment" docUuid="4FE5FE0D-7909-40C1-A8AE-F641192A6048" docUuidSource="ZooBank" docVersion="5" id="A7E05382096E5182BD65560002045C47" lastPageNumber="1" masterDocId="A7E05382096E5182BD65560002045C47" masterDocTitle="Biodiversity data and new species descriptions of polychaetes from offshore waters of the Falkland Islands, an area undergoing hydrocarbon exploration" masterLastPageNumber="86" masterPageNumber="1" pageNumber="1" updateTime="1668168912013" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Biodiversity data and new species descriptions of polychaetes from offshore waters of the Falkland Islands, an area undergoing hydrocarbon exploration</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neal, Lenka</mods:namePart>
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<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
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<mods:nameIdentifier type="email">l.nealova@nhm.ac.uk</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Paterson, Gordon L. J.</mods:namePart>
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<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Blockley, David</mods:namePart>
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<mods:affiliation>Greenland Institute of Natural Resources, Nuuk, Greenland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6368-1101</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Scott, Ben</mods:namePart>
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<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5590-7174</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sherlock, Emma</mods:namePart>
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<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huque, Cate</mods:namePart>
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<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Glover, Adrian G.</mods:namePart>
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<mods:affiliation>Natural History Museum, London, UK</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>938</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>86</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.938.49349</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.938.49349</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-938-1</mods:identifier>
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<mods:identifier type="ZooBank">76B7FF03FEB14884AD2955BE864F0EBF</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A7E05382096E5182BD65560002045C47</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164479465" LSID="urn:lsid:zoobank.org:act:4FE5FE0D-7909-40C1-A8AE-F641192A6048" httpUri="http://treatment.plazi.org/id/D4382279AE655258BC05E2AAA5D4AD57" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="D4382279-AE65-5258-BC05-E2AAA5D4AD57" authority="Neal & Paterson" authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Polynoidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae" status="sp. nov.">Prosphaerosyllis modinouae Neal & Paterson</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">Figures 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100; specimen Shirla-stained) a complete specimen in dorsal view b pattern of dorsal body papillae (= darkly stained dots) in anterior end c pattern of dorsal body papillae (= darkly stained dots) in mid-body segments d pattern of ventral papillae (= darkly stained dots). Scale bars: 500 µm (a); 200 µm (b, d)." figureDoi="10.3897/zookeys.938.49349.figure6" httpUri="https://binary.pensoft.net/fig/417684" pageId="0" pageNumber="1">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">, 8</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100) a specimen in partial ventral view b, c examples of ventral cirri (Vc) and elongated lateral body papillae (Lp) d example of parapodium e detail of aciculae. Scale bars: 250 µm (a); 50 µm (c, d); 20 µm (e)." figureDoi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1">, 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">, 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Comparative figure of Sphaerosyllis palpopapillata (holotype, ZMH P- 20751) (a) and Prosphaerosyllis modinouae sp. nov. (b, c). Scale bars: 1 mm." figureDoi="10.3897/zookeys.938.49349.figure11" httpUri="https://binary.pensoft.net/fig/417689" pageId="0" pageNumber="1">, 11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Materials.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Sample 63MFA, 448 m,
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<geoCoordinate degrees="49.2457310" direction="south" orientation="latitude" precision="1" value="-49.24573">-49.2457310</geoCoordinate>
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,
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<geoCoordinate degrees="59.1254934" direction="west" orientation="longitude" precision="1" value="-59.125492">-59.1254934</geoCoordinate>
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, coll. 17/03/2012, ind. 1,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">holotype</emphasis>
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(NHM.2018.25100). Sample 21MFC, 445 m
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<geoCoordinate degrees="49.2866453" direction="south" orientation="latitude" precision="1" value="-49.286644">-49.2866453</geoCoordinate>
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,
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<geoCoordinate degrees="59.1130539" direction="west" orientation="longitude" precision="1" value="-59.113052">-59.1130539</geoCoordinate>
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, coll. 16/04/2012, ind. 1,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paratype</emphasis>
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(NHM.2018.24236). Sample 28MFB, 451 m,
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<geoCoordinate degrees="49.3044189" direction="south" orientation="latitude" precision="1" value="-49.30442">-49.3044189</geoCoordinate>
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,
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<geoCoordinate degrees="59.0852225" direction="west" orientation="longitude" precision="1" value="-59.085224">-59.0852225</geoCoordinate>
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, coll. 23/04/2012, ind. 1,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paratype</emphasis>
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(NHM.2018.24386).
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<emphasis bold="true" pageId="0" pageNumber="1">Other materials</emphasis>
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: Sample 15MFB, 454 m,
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<geoCoordinate degrees="49.2686572" direction="south" orientation="latitude" precision="1" value="-49.268658">-49.2686572</geoCoordinate>
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,
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<geoCoordinate degrees="59.1133764" direction="west" orientation="longitude" precision="1" value="-59.113377">-59.1133764</geoCoordinate>
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, coll. 16/04/2012, ind. 1, NHM.2018.24080. Sample 25MFA, 447 m,
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<geoCoordinate degrees="49.3050428" direction="south" orientation="latitude" precision="1" value="-49.305042">-49.3050428</geoCoordinate>
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,
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<geoCoordinate degrees="59.1677492" direction="west" orientation="longitude" precision="1" value="-59.167747">-59.1677492</geoCoordinate>
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, coll. 16/04/2012, ind. 1, NHM.2018.24302. Sample 25MFC, 438 m, -49.3050428, Longitude: -59.1677492, coll. 16/04/2012, ind. 1, NHM.2018.24324. Sample 35MFC, 450 m,
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<geoCoordinate degrees="49.3221858" direction="south" orientation="latitude" precision="1" value="-49.322186">-49.3221858</geoCoordinate>
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,
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<geoCoordinate degrees="59.0573711" direction="west" orientation="longitude" precision="1" value="-59.05737">-59.0573711</geoCoordinate>
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, coll. 15/04/2012, ind. 1, NHM.2018.24498. Sample 40MFA, 450 m,
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<geoCoordinate degrees="49.3403947" direction="south" orientation="latitude" precision="1" value="-49.340393">-49.3403947</geoCoordinate>
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,
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<geoCoordinate degrees="59.0845558" direction="west" orientation="longitude" precision="1" value="-59.084557">-59.0845558</geoCoordinate>
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, coll. 23/04/2012, ind. 1, NHM.2018.24605.Sample 41MFA, 439 m,
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<geoCoordinate degrees="49.3401736" direction="south" orientation="latitude" precision="1" value="-49.34017">-49.3401736</geoCoordinate>
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,
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<geoCoordinate degrees="59.0570275" direction="west" orientation="longitude" precision="1" value="-59.057026">-59.0570275</geoCoordinate>
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, coll. 14/04/2012, ind. 1, NHM.2018.24637. Sample 44MFB, 429 m,
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<geoCoordinate degrees="49.3585975" direction="south" orientation="latitude" precision="1" value="-49.358597">-49.3585975</geoCoordinate>
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,
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<geoCoordinate degrees="59.1117606" direction="west" orientation="longitude" precision="1" value="-59.11176">-59.1117606</geoCoordinate>
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, coll. 14/04/2012, ind. 1, NHM.2018.24719. Sample 60MFC, 450 m,
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<geoCoordinate degrees="49.2545270" direction="south" orientation="latitude" precision="1" value="-49.254528">-49.2545270</geoCoordinate>
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,
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<geoCoordinate degrees="59.0494527" direction="west" orientation="longitude" precision="1" value="-59.049454">-59.0494527</geoCoordinate>
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, coll. 19/03/2012, ind. 1, NHM.2018.25062. Sample 64MFC, 447 m,
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<geoCoordinate degrees="49.2455380" direction="south" orientation="latitude" precision="1" value="-49.245537">-49.2455380</geoCoordinate>
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,
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<geoCoordinate degrees="59.1060962" direction="west" orientation="longitude" precision="1" value="-59.106094">-59.1060962</geoCoordinate>
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, coll. 18/03/2012, ind. 1, NHM.2018.25127. Sample 69MFC, 442 m,
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<geoCoordinate degrees="49.2887900" direction="south" orientation="latitude" precision="1" value="-49.28879">-49.2887900</geoCoordinate>
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,
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<geoCoordinate degrees="59.1005700" direction="west" orientation="longitude" precision="1" value="-59.10057">-59.1005700</geoCoordinate>
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, coll. 19/03/2012, ind. 1, NHM.2018.2521.
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<emphasis bold="true" pageId="0" pageNumber="1">Comparative materials</emphasis>
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:
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<taxonomicName baseAuthorityName="McIntosh" baseAuthorityYear="1885" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis kerguelensis" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="kerguelensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis kerguelensis</emphasis>
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</taxonomicName>
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: Kerguelen Islands (off Cumberland Bay), 232 m,
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<geoCoordinate degrees="48.750000" direction="south" orientation="latitude" precision="1" value="-48.75">-48.750000</geoCoordinate>
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,
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<geoCoordinate degrees="69.233333" direction="west" orientation="longitude" precision="1" value="-69.23333">-69.233333</geoCoordinate>
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, holotype, BMNH.85.12.1.155.
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<taxonomicName class="Polychaeta" family="Syllidae" genus="Sphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerosyllis palpopapillata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="palpopapillata">
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<emphasis italics="true" pageId="0" pageNumber="1">Sphaerosyllis palpopapillata</emphasis>
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</taxonomicName>
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: Antarctic Peninsula, 300 m,
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<geoCoordinate degrees="63.27777778" direction="south" orientation="latitude" precision="1" value="-63.27778">-63.27777778</geoCoordinate>
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,
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<geoCoordinate degrees="63.72166667" direction="west" orientation="longitude" precision="1" value="-63.72167">-63.72166667</geoCoordinate>
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, holotype, ZMH P-20751.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype (NHM.2018.25100) a complete, very small, slender specimen, 4.5 mm long and 0.5 mm wide (at mid-body) for 31 chaetigers. Paratype (NHM.2018.24236), complete specimen 2.8 mm long and 0.4 mm wide (at mid-body) for 31 chaetigers. Paratype (SEM specimen, NHM.2018.24386) complete specimen 3.5 mm long and 0.4 mm wide (at mid-body) for 30 chaetigers. Paratype (SEM specimen, NHM.2018.24236) incomplete specimen 1.2 mm long and 0.25 mm wide for 12 chaetigers. Integument of body appearing smooth under light microscopy (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5a</figureCitation>
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), upon staining (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100; specimen Shirla-stained) a complete specimen in dorsal view b pattern of dorsal body papillae (= darkly stained dots) in anterior end c pattern of dorsal body papillae (= darkly stained dots) in mid-body segments d pattern of ventral papillae (= darkly stained dots). Scale bars: 500 µm (a); 200 µm (b, d)." figureDoi="10.3897/zookeys.938.49349.figure6" httpUri="https://binary.pensoft.net/fig/417684" pageId="0" pageNumber="1">6a-d</figureCitation>
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) and under SEM (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">7a, c, f</figureCitation>
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) sparse papillation detected dorsally with two alternating longitudinal rows of digitiform papillae on each side of the body (= 4 rows in total); palps with some diffused small papillae (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8b</figureCitation>
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); papillation on body venter diffused (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100; specimen Shirla-stained) a complete specimen in dorsal view b pattern of dorsal body papillae (= darkly stained dots) in anterior end c pattern of dorsal body papillae (= darkly stained dots) in mid-body segments d pattern of ventral papillae (= darkly stained dots). Scale bars: 500 µm (a); 200 µm (b, d)." figureDoi="10.3897/zookeys.938.49349.figure6" httpUri="https://binary.pensoft.net/fig/417684" pageId="0" pageNumber="1">6d</figureCitation>
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);parapodial with few tiny papillae (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100) a specimen in partial ventral view b, c examples of ventral cirri (Vc) and elongated lateral body papillae (Lp) d example of parapodium e detail of aciculae. Scale bars: 250 µm (a); 50 µm (c, d); 20 µm (e)." figureDoi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1">9d</figureCitation>
|
|
); slender and elongated lateral body papillae observed in-between parapodia (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100) a specimen in partial ventral view b, c examples of ventral cirri (Vc) and elongated lateral body papillae (Lp) d example of parapodium e detail of aciculae. Scale bars: 250 µm (a); 50 µm (c, d); 20 µm (e)." figureDoi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1">9b, c</figureCitation>
|
|
). Colour in alcohol pale yellow, no pigmentation observed (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5a</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
|
|
<taxonomicName authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (paratype, NHM.2018.24386)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
complete specimen in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
|
|
detail of anterior end in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
|
|
detail of prostomium and palps in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
detail of eyes
|
|
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
|
|
anterior dorsal cirrus
|
|
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
|
|
posterior dorsal cirrus
|
|
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
|
|
posterior parapodium (arrow marking acicula)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
|
|
detail of acicula. Scale bars: 1000
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
); 100
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b, c</emphasis>
|
|
); 25
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">e, f</emphasis>
|
|
); 50
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Prostomium short, wider than long (Figs
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5b, c</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8a</figureCitation>
|
|
). Three antennae present (Figs
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">7d</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8a</figureCitation>
|
|
), all pyriform and very small (difficult to observe under stereo microscope); median antenna positioned near posterior margin of prostomium, ca. 30
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
long; lateral antennae positioned lateromedially on prostomium, smaller than median antenna, ca. 25
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
long (Figs
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">7d</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8a</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure6" httpUri="https://binary.pensoft.net/fig/417684" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
|
|
<taxonomicName authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (holotype, NHM.2018.25100; specimen Shirla-stained)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
complete specimen in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
|
|
pattern of dorsal body papillae (= darkly stained dots) in anterior end
|
|
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
|
|
pattern of dorsal body papillae (= darkly stained dots) in mid-body segments
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
pattern of ventral papillae (= darkly stained dots). Scale bars: 500
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
); 200
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b, d</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Palps very short, almost entirely obscured by prostomium in dorsal view (Figs
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5b, c</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">7a-d</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8a</figureCitation>
|
|
) with only their lateral margins visible in dorsal view; fully fused along their length; provided with some small papillae (Fig.
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8b</figureCitation>
|
|
). Two pairsof large red eyes present (Figs
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5b-d</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8a</figureCitation>
|
|
), in trapezoid arrangement, with anterior and posterior pair close together (almost appearing as a single eye), anterior pair cup-shaped, posterior pair circular; presence of additional eyespots not confirmed.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
|
|
<taxonomicName authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (SEM micrograph)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
specimen (paratype, NHM.2018.24386) in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
|
|
anterior end in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
|
|
paratype (NHM.2018.24236) in dorso-lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
detail of prostomium of paratype (NHM.2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
|
|
detail of dorsal cirri with those on chaetigers 2-4 (paratype, NHM.2018.24386)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
|
|
pattern of dorsal body papillae (marked by white circles) (paratype, NHM.2018.24386). Scale bars: 500
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
); 100
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b, e</emphasis>
|
|
); 200
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
|
|
); 50
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Proventricle starts between chaetigers 3-4 and ends between chaetigers 6-7, with ca. 20 muscle rows (Figs
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5b</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100; specimen Shirla-stained) a complete specimen in dorsal view b pattern of dorsal body papillae (= darkly stained dots) in anterior end c pattern of dorsal body papillae (= darkly stained dots) in mid-body segments d pattern of ventral papillae (= darkly stained dots). Scale bars: 500 µm (a); 200 µm (b, d)." figureDoi="10.3897/zookeys.938.49349.figure6" httpUri="https://binary.pensoft.net/fig/417684" pageId="0" pageNumber="1">6b</figureCitation>
|
|
). First segment achaetous; with small pair of tentacular cirri, ca. 20
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
long, pyriform (similar to antennae) (Figs
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">7d</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8a</figureCitation>
|
|
). Pharynx everted in paratype (NHM.2018.24236) (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">7c, d</figureCitation>
|
|
), without terminal papillae, pharyngeal tooth far from anterior margin.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
|
|
Drawing of anterior end of
|
|
<taxonomicName authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in dorsal view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
) and detail of palps (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Parapodia uniramous, short but distinct, conical (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100) a specimen in partial ventral view b, c examples of ventral cirri (Vc) and elongated lateral body papillae (Lp) d example of parapodium e detail of aciculae. Scale bars: 250 µm (a); 50 µm (c, d); 20 µm (e)." figureDoi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1">9a-c</figureCitation>
|
|
). Dorsal cirri in all chaetigers, including chaetiger 2 (Figs
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Prosphaerosyllis modinouae sp. nov. (SEM micrograph) a specimen (paratype, NHM. 2018.24386) in dorsal view b anterior end in dorsal view c paratype (NHM. 2018.24236) in dorso-lateral view d detail of prostomium of paratype (NHM. 2018.24236) with palps (P), median antenna (Ma), lateral antennae (La), tentacular cirrus (Tc) and dorsal cirrus (Dc) e detail of dorsal cirri with those on chaetigers 2 - 4 (paratype, NHM. 2018.24386) f pattern of dorsal body papillae (marked by white circles) (paratype, NHM. 2018.24386). Scale bars: 500 µm (a); 100 µm (b, e); 200 µm (c); 50 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure7" httpUri="https://binary.pensoft.net/fig/417685" pageId="0" pageNumber="1">7e</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawing of anterior end of Prosphaerosyllis modinouae sp. nov. in dorsal view (a) and detail of palps (b)." figureDoi="10.3897/zookeys.938.49349.figure8" httpUri="https://binary.pensoft.net/fig/417686" pageId="0" pageNumber="1">8a</figureCitation>
|
|
) (missing in some parapodia due to damage); dorsal cirri in anterior chaetigers small (but easy to observe, ca. 40
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
in length, Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5e</figureCitation>
|
|
), similar in form to tentacular cirri and antennae, becoming more elongated in posterior chaetigers (ca. 55
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
long, Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5f</figureCitation>
|
|
). Ventral cirri as extremely slender cirriform structures, inserted near the base of neuropodia (Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100) a specimen in partial ventral view b, c examples of ventral cirri (Vc) and elongated lateral body papillae (Lp) d example of parapodium e detail of aciculae. Scale bars: 250 µm (a); 50 µm (c, d); 20 µm (e)." figureDoi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1">9a-c</figureCitation>
|
|
), becoming progressively longer posteriorly.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
|
|
<taxonomicName authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (holotype, NHM.2018.25100)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
specimen in partial ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b, c</emphasis>
|
|
examples of ventral cirri (Vc) and elongated lateral body papillae (Lp)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
example of parapodium
|
|
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
|
|
detail of aciculae. Scale bars: 250
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
); 50
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">c, d</emphasis>
|
|
); 20
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Chaetae often missing (broken off). One simple dorsal chaeta commonly observed (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10a</figureCitation>
|
|
), present from chaetiger 1; straight and smooth; increasing in size throughout body measuring 30
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
in anterior parapodia, 85
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
in mid-body parapodia and 110
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
in some posterior parapodia (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10b</figureCitation>
|
|
). Simple ventral chaeta observed only in posteriormost parapodia, slightly sigmoid and smooth (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10c</figureCitation>
|
|
). Other chaetae compound falcigers (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10d-g</figureCitation>
|
|
); ca. six per fascicle; all blades unidentate and serrated to greater (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10d, g</figureCitation>
|
|
) or lesser degree (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10e, f</figureCitation>
|
|
). Blades of falcigers of varying lengths with greatest length difference between dorsalmost (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10d</figureCitation>
|
|
) and ventral most (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10e</figureCitation>
|
|
) chaetae in anterior parapodia, this difference becomes particularly pronounced in mid-body parapodia, where long dorsalmost chaeta (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10f</figureCitation>
|
|
) bears particularly short blade (as ratio to length of its shaft); length of blades 5-13
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(total chaetal length 35-60
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
) in anterior chaetigers; 5-13.5
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(total chaetal length 75-100
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
) in mid-body chaetigers and 12-20
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(total chaetal length ca.100
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
) in posterior chaetigers (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10g</figureCitation>
|
|
). Acicula in anterior and posterior chaetigers mostly solitary, acuminate (Figs
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5g, h</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100) a specimen in partial ventral view b, c examples of ventral cirri (Vc) and elongated lateral body papillae (Lp) d example of parapodium e detail of aciculae. Scale bars: 250 µm (a); 50 µm (c, d); 20 µm (e)." figureDoi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1">9e</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
|
|
<taxonomicName authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (paratype NHM.2018.24386)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
simple dorsal chaeta from anterior chaetiger
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
|
|
simple dorsal chaeta from posterior chaetiger
|
|
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
|
|
simple ventral chaeta from posterior chaetgier
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
dorsalmost falciger from anterior chaetiger
|
|
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
|
|
ventralmost falciger from anterior chaetiger
|
|
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
|
|
dorsalmost chaeta from mid body chaetiger
|
|
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
|
|
falcigers from posterior chaetiger. Scale bar: 50
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Pygidium broad, rounded, pygidial cirri not observed in any specimens examined (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosphaerosyllis modinouae sp. nov. (paratype, NHM. 2018.24386) a complete specimen in dorsal view b detail of anterior end in dorsal view c detail of prostomium and palps in dorsal view d detail of eyes e anterior dorsal cirrus f posterior dorsal cirrus g posterior parapodium (arrow marking acicula) h detail of acicula. Scale bars: 1000 µm (a); 100 µm (b, c); 25 µm (e, f); 50 µm (g)." figureDoi="10.3897/zookeys.938.49349.figure5" httpUri="https://binary.pensoft.net/fig/417683" pageId="0" pageNumber="1">5a</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Falkland Island specimens were assigned to genus
|
|
<taxonomicName authorityName="San Martin" authorityYear="1984" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis</emphasis>
|
|
</taxonomicName>
|
|
San
|
|
<normalizedToken originalValue="Martín">Martin</normalizedToken>
|
|
, 1984 based on morphological characters only as no reproductive specimens were observed.
|
|
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.57.2005.1438" author="San Martin, G" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="39 - 152" refId="B53" refString="San Martin, G, 2005. Exogoninae (Polychaeta: Syllidae) from Australia with the description of a new genus and twenty-two new species. Records of the Australian Museum 57 (1): 39 - 152, DOI: https://doi.org/10.3853/j.0067-1975.57.2005.1438" title="Exogoninae (Polychaeta: Syllidae) from Australia with the description of a new genus and twenty-two new species." url="https://doi.org/10.3853/j.0067-1975.57.2005.1438" volume="57" year="2005">
|
|
San
|
|
<normalizedToken originalValue="Martín">Martin</normalizedToken>
|
|
(2005)
|
|
</bibRefCitation>
|
|
emended the generic diagnosis and further distinguished
|
|
<taxonomicName authorityName="San Martin" authorityYear="1984" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis</emphasis>
|
|
</taxonomicName>
|
|
from the similar genus
|
|
<taxonomicName class="Polychaeta" family="Syllidae" genus="Sphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerosyllis" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Sphaerosyllis</emphasis>
|
|
</taxonomicName>
|
|
, however here we provide a comparison for species in both genera with the type locality inthe southern waters because not all species have been revised.
|
|
<taxonomicName authorityName="Gravier" authorityYear="1906" class="Polychaeta" family="Syllidae" genus="Sphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerosyllis antarctica" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="antarctica">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Sphaerosyllis antarctica</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="S. hirsuta" pageId="0" pageNumber="1" rank="species" species="hirsuta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. hirsuta</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="S. sublaevis" pageId="0" pageNumber="1" rank="species" species="sublaevis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. sublaevis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="S. capensis" pageId="0" pageNumber="1" rank="species" species="capensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. capensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="S. dubiosa" pageId="0" pageNumber="1" rank="species" species="dubiosa">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. dubiosa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="S. lateropapillata subsp. uteae" pageId="0" pageNumber="1" rank="subSpecies" species="lateropapillata" subSpecies="uteae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. lateropapillata uteae</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName baseAuthorityName="McIntosh" baseAuthorityYear="1885" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis kerguelensis" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="kerguelensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis kerguelensis</emphasis>
|
|
</taxonomicName>
|
|
(holotype BMNH.85.12.1.155 examined as part of this study) can be easily distinguished from the Falkland Island species due to absence of dorsal cirrus on chaetiger 2. Of species with a dorsal cirrus on chaetiger 2 present (
|
|
<taxonomicName lsidName="S. semiveruccosa" pageId="0" pageNumber="1" rank="species" species="semiveruccosa">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. semiveruccosa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="P. joinvillensis" pageId="0" pageNumber="1" rank="species" species="joinvillensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. joinvillensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="P. capensis subsp. chilensis" pageId="0" pageNumber="1" rank="subSpecies" species="capensis" subSpecies="chilensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. capensis chilensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="S. brandhorsti" pageId="0" pageNumber="1" rank="species" species="brandhorsti">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. brandhorsti</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName lsidName="P. brachycephala" pageId="0" pageNumber="1" rank="species" species="brachycephala">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. brachycephala</emphasis>
|
|
</taxonomicName>
|
|
) the new species can be easily distinguished by having small antennae and short palps (often with only their lateral margins observable in dorsal view, otherwise mostly obscured by prostomium).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Falkland Islands species is most similar to
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
De Nogueira, San
|
|
<normalizedToken originalValue="Martín">Martin</normalizedToken>
|
|
& Amaral, 2001 described from intertidal depths in Brazil in having short antennae and a sparse distribution of body papillae. However, other than length of the palps,
|
|
<taxonomicName lsidName="P. modinouae" pageId="0" pageNumber="1" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be further differentiated by having all falcigerousblades serrated (these are smooth from mid-body chaetigers in
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
), in lacking iridescent inclusions in the dorsal cirri and in the dorsal cirri becoming elongated throughout the body in
|
|
<taxonomicName lsidName="P. modinouae" pageId="0" pageNumber="1" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. There also appear to be greater differences in length of falcigerous blades in
|
|
<taxonomicName lsidName="P. modinouae" pageId="0" pageNumber="1" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10d-g</figureCitation>
|
|
) compared to
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
(see
|
|
<bibRefCitation DOI="https://doi.org/10.1080/00222930152667096" author="De Nogueira, JM" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="1" pagination="1773 - 1794" refId="B19" refString="De Nogueira, JM, San Martin, G, Amaral, AC, 2001. Description of five new species of Exogoninae Rioja, 1925 (Polychaeta: Syllidae) associated with the stony coral Mussismilia hispida (Verrill, 1868) in Sao Paulo State, Brazil. Journal of Natural History 35 (12): 1773 - 1794, DOI: https://doi.org/10.1080/00222930152667096" title="Description of five new species of Exogoninae Rioja, 1925 (Polychaeta: Syllidae) associated with the stony coral Mussismilia hispida (Verrill, 1868) in Sao Paulo State, Brazil." url="https://doi.org/10.1080/00222930152667096" volume="35" year="2001">DeNogueira et al. 2001</bibRefCitation>
|
|
;
|
|
<bibRefCitation DOI="https://doi.org/10.1017/S0025315409000095" author="Fukuda, MV" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="0" pageNumber="1" pagination="1443 - 1454" refId="B24" refString="Fukuda, MV, Yunda-Guarin, G, Nogueira, JM, 2009. The genus Prosphaerosyllis (Polychaeta: Syllidae: Exogoninae) in Brazil, with description of a new species. Journal of the Marine Biological Association of the United Kingdom 89 (7): 1443 - 1454, DOI: https://doi.org/10.1017/S0025315409000095" title="The genus Prosphaerosyllis (Polychaeta: Syllidae: Exogoninae) in Brazil, with description of a new species." url="https://doi.org/10.1017/S0025315409000095" volume="89" year="2009">Fukuda et al. 2009</bibRefCitation>
|
|
). We also suggest that Brazilian specimens from bathyal depths assigned to
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
by
|
|
<bibRefCitation DOI="https://doi.org/10.1017/S0025315409000095" author="Fukuda, MV" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="0" pageNumber="1" pagination="1443 - 1454" refId="B24" refString="Fukuda, MV, Yunda-Guarin, G, Nogueira, JM, 2009. The genus Prosphaerosyllis (Polychaeta: Syllidae: Exogoninae) in Brazil, with description of a new species. Journal of the Marine Biological Association of the United Kingdom 89 (7): 1443 - 1454, DOI: https://doi.org/10.1017/S0025315409000095" title="The genus Prosphaerosyllis (Polychaeta: Syllidae: Exogoninae) in Brazil, with description of a new species." url="https://doi.org/10.1017/S0025315409000095" volume="89" year="2009">Fukuda et al. (2009)</bibRefCitation>
|
|
may in fact represent a different species, even more closely aligned to
|
|
<taxonomicName lsidName="P. modinouae" pageId="0" pageNumber="1" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. This suggestion is based mainly on much deeper distribution (down to 650 m) reported by
|
|
<bibRefCitation DOI="https://doi.org/10.1017/S0025315409000095" author="Fukuda, MV" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="0" pageNumber="1" pagination="1443 - 1454" refId="B24" refString="Fukuda, MV, Yunda-Guarin, G, Nogueira, JM, 2009. The genus Prosphaerosyllis (Polychaeta: Syllidae: Exogoninae) in Brazil, with description of a new species. Journal of the Marine Biological Association of the United Kingdom 89 (7): 1443 - 1454, DOI: https://doi.org/10.1017/S0025315409000095" title="The genus Prosphaerosyllis (Polychaeta: Syllidae: Exogoninae) in Brazil, with description of a new species." url="https://doi.org/10.1017/S0025315409000095" volume="89" year="2009">Fukuda et al. (2009)</bibRefCitation>
|
|
compared to 4-7 m depth at the type locality of
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
, which was also reported to be associated with coral colonies (
|
|
<bibRefCitation DOI="https://doi.org/10.1080/00222930152667096" author="De Nogueira, JM" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="1" pagination="1773 - 1794" refId="B19" refString="De Nogueira, JM, San Martin, G, Amaral, AC, 2001. Description of five new species of Exogoninae Rioja, 1925 (Polychaeta: Syllidae) associated with the stony coral Mussismilia hispida (Verrill, 1868) in Sao Paulo State, Brazil. Journal of Natural History 35 (12): 1773 - 1794, DOI: https://doi.org/10.1080/00222930152667096" title="Description of five new species of Exogoninae Rioja, 1925 (Polychaeta: Syllidae) associated with the stony coral Mussismilia hispida (Verrill, 1868) in Sao Paulo State, Brazil." url="https://doi.org/10.1080/00222930152667096" volume="35" year="2001">De Nogueira et al. 2001</bibRefCitation>
|
|
). However, specimens of
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
were not available for examination as part of this study.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Another similar species is
|
|
<taxonomicName class="Polychaeta" family="Syllidae" genus="Sphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerosyllis palpopapillata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="palpopapillata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Sphaerosyllis palpopapillata</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Hartmann-Schröder">Hartmann-Schroeder</normalizedToken>
|
|
& Rosenfeldt, 1992 described from the Antarctic Peninsula, 300 m depth. Unfortunately, the description and drawings provided by
|
|
<bibRefCitation author="Hartmann-Schroeder, G" journalOrPublisher="Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="1" pagination="85 - 124" refId="B31" refString="Hartmann-Schroeder, G, Rosenfeldt, P, 1992. Die Polychaeten der "Polarstern" -Reise ANT V/1 in die Antarktis 1986. Teil 1: Euphrosinidae bis Iphitimidae. Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut 89: 85 - 124" title="Die Polychaeten der " Polarstern " - Reise ANT V / 1 in die Antarktis 1986. Teil 1: Euphrosinidae bis Iphitimidae." volume="89" year="1992">
|
|
<normalizedToken originalValue="Hartmann-Schröder">Hartmann-Schroeder</normalizedToken>
|
|
and Rosenfeldt (1992)
|
|
</bibRefCitation>
|
|
are of limited value. Therefore, the holotype (ZMH P-20751) was loaned from Zoologisches Museum Hamburg and photographed here for the first time (Figs
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Comparative figure of Sphaerosyllis palpopapillata (holotype, ZMH P- 20751) (a) and Prosphaerosyllis modinouae sp. nov. (b, c). Scale bars: 1 mm." figureDoi="10.3897/zookeys.938.49349.figure11" httpUri="https://binary.pensoft.net/fig/417689" pageId="0" pageNumber="1">11a</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Sphaerosyllis palpopapillata (holotype ZMH P- 20751) a anterior end in dorsal view b, c examples of ventral cirri (marked by arrows) d example of lateral body papillae (marked by arrow) e acicula and simple ventral chaeta f falcigers. Scale bars: 1000 µm (a); 20 µm (c, f), 25 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure12" httpUri="https://binary.pensoft.net/fig/417690" pageId="0" pageNumber="1">12a-f</figureCitation>
|
|
). The holotype, which is the only known specimen of this species, was found to be a small anterior fragment, with structures such as the antennae now missing andsome chaetae broken off. As a result,
|
|
<taxonomicName lsidName="S. palpopapillata" pageId="0" pageNumber="1" rank="species" species="palpopapillata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. palpopapillata</emphasis>
|
|
</taxonomicName>
|
|
remains a poorly known species, until new material from the type locality becomes available. The holotype (ZMH P-20751) (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Comparative figure of Sphaerosyllis palpopapillata (holotype, ZMH P- 20751) (a) and Prosphaerosyllis modinouae sp. nov. (b, c). Scale bars: 1 mm." figureDoi="10.3897/zookeys.938.49349.figure11" httpUri="https://binary.pensoft.net/fig/417689" pageId="0" pageNumber="1">11a</figureCitation>
|
|
) is similar to
|
|
<taxonomicName lsidName="P. modinouae" pageId="0" pageNumber="1" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Comparative figure of Sphaerosyllis palpopapillata (holotype, ZMH P- 20751) (a) and Prosphaerosyllis modinouae sp. nov. (b, c). Scale bars: 1 mm." figureDoi="10.3897/zookeys.938.49349.figure11" httpUri="https://binary.pensoft.net/fig/417689" pageId="0" pageNumber="1">11b, c</figureCitation>
|
|
) in the form of the palps and also possesses the elongated lateral body papillae (Fig.
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Sphaerosyllis palpopapillata (holotype ZMH P- 20751) a anterior end in dorsal view b, c examples of ventral cirri (marked by arrows) d example of lateral body papillae (marked by arrow) e acicula and simple ventral chaeta f falcigers. Scale bars: 1000 µm (a); 20 µm (c, f), 25 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure12" httpUri="https://binary.pensoft.net/fig/417690" pageId="0" pageNumber="1">12d</figureCitation>
|
|
), which were not reported by
|
|
<bibRefCitation author="Hartmann-Schroeder, G" journalOrPublisher="Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="1" pagination="85 - 124" refId="B31" refString="Hartmann-Schroeder, G, Rosenfeldt, P, 1992. Die Polychaeten der "Polarstern" -Reise ANT V/1 in die Antarktis 1986. Teil 1: Euphrosinidae bis Iphitimidae. Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut 89: 85 - 124" title="Die Polychaeten der " Polarstern " - Reise ANT V / 1 in die Antarktis 1986. Teil 1: Euphrosinidae bis Iphitimidae." volume="89" year="1992">
|
|
<normalizedToken originalValue="Hartmann-Schröder">Hartmann-Schroeder</normalizedToken>
|
|
and Rosenfeldt (1992)
|
|
</bibRefCitation>
|
|
, although the rows of dorsal papillae were not confirmed by us. The main differences observed were the form and length of ventral cirri, which are distinctly longer and slender in
|
|
<taxonomicName lsidName="S. modinouae" pageId="0" pageNumber="1" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (Figs
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Prosphaerosyllis modinouae sp. nov. (holotype, NHM. 2018.25100) a specimen in partial ventral view b, c examples of ventral cirri (Vc) and elongated lateral body papillae (Lp) d example of parapodium e detail of aciculae. Scale bars: 250 µm (a); 50 µm (c, d); 20 µm (e)." figureDoi="10.3897/zookeys.938.49349.figure9" httpUri="https://binary.pensoft.net/fig/417687" pageId="0" pageNumber="1">9b, c</figureCitation>
|
|
;
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Sphaerosyllis palpopapillata (holotype ZMH P- 20751) a anterior end in dorsal view b, c examples of ventral cirri (marked by arrows) d example of lateral body papillae (marked by arrow) e acicula and simple ventral chaeta f falcigers. Scale bars: 1000 µm (a); 20 µm (c, f), 25 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure12" httpUri="https://binary.pensoft.net/fig/417690" pageId="0" pageNumber="1">12b, c</figureCitation>
|
|
). While both species have unidentate falcigers, their blades in
|
|
<taxonomicName lsidName="S. palpopapillata" pageId="0" pageNumber="1" rank="species" species="palpopapillata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. palpopapillata</emphasis>
|
|
</taxonomicName>
|
|
are shorter (6-7
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
) and similar in size where observed (Fig.
|
|
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Sphaerosyllis palpopapillata (holotype ZMH P- 20751) a anterior end in dorsal view b, c examples of ventral cirri (marked by arrows) d example of lateral body papillae (marked by arrow) e acicula and simple ventral chaeta f falcigers. Scale bars: 1000 µm (a); 20 µm (c, f), 25 µm (d)." figureDoi="10.3897/zookeys.938.49349.figure12" httpUri="https://binary.pensoft.net/fig/417690" pageId="0" pageNumber="1">12e, f</figureCitation>
|
|
), but in the new species their lengths are more variable (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Prosphaerosyllis modinouae sp. nov. (paratype NHM. 2018.24386) a simple dorsal chaeta from anterior chaetiger b simple dorsal chaeta from posterior chaetiger c simple ventral chaeta from posterior chaetgier d dorsalmost falciger from anterior chaetiger e ventralmost falciger from anterior chaetiger f dorsalmost chaeta from mid body chaetiger g falcigers from posterior chaetiger. Scale bar: 50 µm." figureDoi="10.3897/zookeys.938.49349.figure10" httpUri="https://binary.pensoft.net/fig/417688" pageId="0" pageNumber="1">10d-g</figureCitation>
|
|
) as already discussed in comparison with
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
. Furthermore, the image provided in the original description (
|
|
<bibRefCitation author="Hartmann-Schroeder, G" journalOrPublisher="Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="1" pagination="85 - 124" refId="B31" refString="Hartmann-Schroeder, G, Rosenfeldt, P, 1992. Die Polychaeten der "Polarstern" -Reise ANT V/1 in die Antarktis 1986. Teil 1: Euphrosinidae bis Iphitimidae. Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut 89: 85 - 124" title="Die Polychaeten der " Polarstern " - Reise ANT V / 1 in die Antarktis 1986. Teil 1: Euphrosinidae bis Iphitimidae." volume="89" year="1992">
|
|
<normalizedToken originalValue="Hartmann-Schröder">Hartmann-Schroeder</normalizedToken>
|
|
and Rosenfeldt 1992
|
|
</bibRefCitation>
|
|
: fig. 34) suggests that antennae in
|
|
<taxonomicName lsidName="S. palpopapillata" pageId="0" pageNumber="1" rank="species" species="palpopapillata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. palpopapillata</emphasis>
|
|
</taxonomicName>
|
|
are large, not small as in the new species or in
|
|
<taxonomicName lsidName="P. isabellae" pageId="0" pageNumber="1" rank="species" species="isabellae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. isabellae</emphasis>
|
|
</taxonomicName>
|
|
. Unfortunately, as already mentioned, the antennae have since been lost in the holotype of
|
|
<taxonomicName lsidName="S. palpopapillata" pageId="0" pageNumber="1" rank="species" species="palpopapillata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">S. palpopapillata</emphasis>
|
|
</taxonomicName>
|
|
and this character cannot be verified.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure11" httpUri="https://binary.pensoft.net/fig/417689" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
|
|
Comparative figure of
|
|
<taxonomicName class="Polychaeta" family="Syllidae" genus="Sphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerosyllis palpopapillata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="palpopapillata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Sphaerosyllis palpopapillata</emphasis>
|
|
</taxonomicName>
|
|
(holotype, ZMH P-20751) (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
) and
|
|
<taxonomicName authorityName="Neal & Paterson" authorityYear="2020" class="Polychaeta" family="Syllidae" genus="Prosphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosphaerosyllis modinouae" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="modinouae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Prosphaerosyllis modinouae</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b, c</emphasis>
|
|
). Scale bars: 1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">The lack of detailed descriptions and reliable drawings/images of the known species from the southern waters, complicate the efforts in describing new species. While we believe that observations provided in this study justify the establishment of a new species from the Falkland Islands material, the known species are clearly in need of revision. However, such an undertaking is beyond the scope of this study.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">This species is dedicated to Yvett Modinou, a passionate science communicator, who inspired the fourth author (BS) to join the NHM London.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">This species is only known from its type locality, North Falklands Basin, ca. 450 m depth.</paragraph>
|
|
<caption doi="10.3897/zookeys.938.49349.figure12" httpUri="https://binary.pensoft.net/fig/417690" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
|
|
<taxonomicName class="Polychaeta" family="Syllidae" genus="Sphaerosyllis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerosyllis palpopapillata" order="Phyllodocida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="palpopapillata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Sphaerosyllis palpopapillata</emphasis>
|
|
</taxonomicName>
|
|
(holotype ZMH P-20751)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
anterior end in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="1">b, c</emphasis>
|
|
examples of ventral cirri (marked by arrows)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
example of lateral body papillae (marked by arrow)
|
|
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
|
|
acicula and simple ventral chaeta
|
|
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
|
|
falcigers. Scale bars: 1000
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
|
|
); 20
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">c, f</emphasis>
|
|
), 25
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |