953 lines
132 KiB
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953 lines
132 KiB
XML
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<mods:title id="D78F20D1148B057537879B22CBDD41F5">A neritid gastropod name “ Theodoxus dniestroviensis Put’, 1972 ” is a junior subjective synonym of Th. fluviatilis (L., 1758): decision based on the topotypic specimens study</mods:title>
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<mods:namePart id="60A78E09641187C3A33436583C9613EC">Anistratenko, Olga Yu.</mods:namePart>
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<mods:affiliation id="0AEA10758E90393A053A86C96022CEB1">I. I. Schmalhausen Institute of Zoology of NAS of Ukraine, B. Khmelnitsky Str. 15, Kiev 01054, UKRAINE. E-mail: anistrat @ izan. kiev. ua & Institute of Geological Sciences of NAS of Ukraine, O. Gontchar Str. 55 b, Kiev 01054, UKRAINE. E-mail: olga. anistrat @ gmail. com</mods:affiliation>
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<mods:affiliation id="434AC5879828CCC75A6789D3448BA7C2">I. I. Schmalhausen Institute of Zoology of NAS of Ukraine, B. Khmelnitsky Str. 15, Kiev 01054, UKRAINE. E-mail: anistrat @ izan. kiev. ua & National University of Life and Environmental Sciences of Ukraine, General Rodimcev Str. 19, Kiev 03041, UKRAINE. E-mail: oomit 99 @ ukr. net</mods:affiliation>
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<taxonomicName id="4C0B4D65FFB4C40BFA543B7925815D7C" ID-CoL="8YZBF" authority="Put’, 1957" box="[849,1187,1117,1146]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="dniestroviensis">
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<emphasis id="B97FEAF4FFB4C40BFA543B7925815D7C" box="[849,1187,1117,1146]" italics="true" pageId="1" pageNumber="62">Theodoxus dniestroviensis</emphasis>
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Put’, 1957
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<paragraph id="8BB436E6FFB4C40BFAF03B5B25AE5D9F" blockId="1.[849,1328,1117,1177]" box="[1013,1164,1151,1177]" pageId="1" pageNumber="62">
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(
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<figureCitation id="13302A63FFB4C40BFAFB3B5B25A65D9F" box="[1022,1156,1151,1177]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="1" pageNumber="62">Fig. 4 A–D</figureCitation>
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)
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</paragraph>
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<taxonomicName id="4C0B4D65FFB4C40BFA243B9F253C5DD6" box="[801,1054,1211,1232]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="dniestroviensis">
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<emphasis id="B97FEAF4FFB4C40BFA243B9F253C5DD6" box="[801,1054,1211,1232]" italics="true" pageId="1" pageNumber="62">Theodoxus dniestroviensis</emphasis>
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</taxonomicName>
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Put’, 1957: 98.
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<emphasis id="B97FEAF4FFB4C40BFA243BF3253C5DEA" box="[801,1054,1239,1260]" italics="true" pageId="1" pageNumber="62">Theodoxus dniestroviensis</emphasis>
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Put’, 1972: 80-82, textfig. 5.
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<emphasis id="B97FEAF4FFB4C40BFA243BD722185C0E" box="[801,826,1267,1288]" italics="true" pageId="1" pageNumber="62">Th</emphasis>
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[
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<emphasis id="B97FEAF4FFB4C40BFA463BD722B05C0E" box="[835,914,1267,1288]" italics="true" pageId="1" pageNumber="62">eodoxus</emphasis>
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]
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<emphasis id="B97FEAF4FFB4C40BFA9B3BD7250F5C0E" box="[926,1069,1267,1288]" italics="true" pageId="1" pageNumber="62">dniestroviensis</emphasis>
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</taxonomicName>
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Put’, 1972.
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<emphasis id="B97FEAF4FFB4C40BFDA43BD0258F5C0F" box="[1185,1197,1268,1289]" italics="true" pageId="1" pageNumber="62">–</emphasis>
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O.
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Anistratenko
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<emphasis id="B97FEAF4FFB4C40BFC493BD022785C22" italics="true" pageId="1" pageNumber="62">et al</emphasis>
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., 1999: 19
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</bibRefCitation>
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, fig. 4,
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<emphasis id="B97FEAF4FFB4C40BFD0F3A3425345C23" box="[1034,1046,1296,1317]" italics="true" pageId="1" pageNumber="62">8</emphasis>
|
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.
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</paragraph>
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<emphasis id="B97FEAF4FFB4C40BFA243A0822185C47" box="[801,826,1324,1345]" italics="true" pageId="1" pageNumber="62">Th</emphasis>
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[
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<emphasis id="B97FEAF4FFB4C40BFA473A0822B25C47" box="[834,912,1324,1345]" italics="true" pageId="1" pageNumber="62">eodoxus</emphasis>
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]
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<emphasis id="B97FEAF4FFB4C40BFA983A0825095C47" box="[925,1067,1324,1345]" italics="true" pageId="1" pageNumber="62">dniestroviensis</emphasis>
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</taxonomicName>
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Put’, 1972. –
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<collectionCode id="ED1AAE23FFB4C40BFDA93A08259F5C47" box="[1196,1213,1324,1345]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="62" type="Museum">V</collectionCode>
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.Anistratenko, O.
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<bibRefCitation id="EF9A4B17FFB4C40BFA403A6C25135C5B" author="Anistratenko V. V. & Anistratenko O. Yu." box="[837,1073,1352,1374]" pageId="1" pageNumber="62" pagination="1 - 240" refId="ref4501" refString="Anistratenko V. V., Anistratenko O. Yu. 2001. Class Polyplacophora or Chitons, Class Gastropoda - Cyclobranchia, Scutibranchia and Pectinibranchia (part). Fauna of Ukraine, Mollusca, 29 (1 / 1). Kiev: Veles: 1 - 240 [In Russian with extended English summary and captions]." type="book chapter" year="2001">Anistratenko, 2001: 108</bibRefCitation>
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.
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</paragraph>
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<taxonomicName id="4C0B4D65FFB4C40BFA243A4025015C7F" ID-CoL="8YZBF" authority="Put’, 1957" box="[801,1059,1380,1401]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="dniestroviensis">
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<emphasis id="B97FEAF4FFB4C40BFA243A4025015C7F" box="[801,1059,1380,1401]" italics="true" pageId="1" pageNumber="62">Theodoxus dniestroviensis</emphasis>
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</taxonomicName>
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Put’, 1972.
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<emphasis id="B97FEAF4FFB4C40BFDA73A41258C5C7C" box="[1186,1198,1381,1402]" italics="true" pageId="1" pageNumber="62">–</emphasis>
|
||
<treatmentCitation id="0AAA10F7FFB4C40BFDB23A40228A5C90" author="Vinarski M. V. & Kantor Yu. I." page="155" pageId="1" pageNumber="62" year="2016">
|
||
<bibRefCitation id="EF9A4B17FFB4C40BFDB23A40228A5C90" author="Vinarski M. V. & Kantor Yu. I." pageId="1" pageNumber="62" refId="ref5176" refString="Vinarski M. V., Kantor Yu. I. 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. Moscow, A. N. Severtsov Institute of Ecology and Evolution of RAS, 544 p." type="book" year="2016">Vinarski, Kantor, 2016: 155</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
(in the
|
||
<taxonomicName id="4C0B4D65FFB4C40BFAF73AA425525C93" ID-CoL="73WSQ" authorityName="Linnaeus" authorityYear="1758" box="[1010,1136,1408,1430]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB4C40BFAF73AA425525C93" box="[1010,1136,1408,1430]" italics="true" pageId="1" pageNumber="62">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
synonymy).
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB4C40BFA243AB8242C5CC8" blockId="1.[801,1376,1211,1486]" pageId="1" pageNumber="62">
|
||
<treatmentCitationGroup id="AB1B11C8FFB4C40BFA243AB822EE5CC8" pageId="1" pageNumber="62">
|
||
?
|
||
<taxonomicName id="4C0B4D65FFB4C40BFA293AB8250B5CB7" ID-CoL="8YZBF" authority="Put’, 1957" box="[812,1065,1436,1457]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB4C40BFA293AB8250B5CB7" box="[812,1065,1436,1457]" italics="true" pageId="1" pageNumber="62">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
Put’, 1972.
|
||
<emphasis id="B97FEAF4FFB4C40BFDA53AB9258E5CB4" box="[1184,1196,1437,1458]" italics="true" pageId="1" pageNumber="62">–</emphasis>
|
||
<collectionCode id="ED1AAE23FFB4C40BFDB63AB925E65CB4" box="[1203,1220,1437,1458]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="62" type="Museum">V</collectionCode>
|
||
.
|
||
<treatmentCitation id="0AAA10F7FFB4C40BFDC93AB822EE5CC8" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." page="120" pageId="1" pageNumber="62" year="2020">
|
||
<bibRefCitation id="EF9A4B17FFB4C40BFDC93AB822EE5CC8" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." pageId="1" pageNumber="62" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB4C40BFC483AB922785CCB" italics="true" pageId="1" pageNumber="62">et al</emphasis>
|
||
., 2020: 120
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
(in the
|
||
<taxonomicName id="4C0B4D65FFB4C40BFD133A9C25B75CCB" ID-CoL="73WSQ" authorityName="Linnaeus" authorityYear="1758" box="[1046,1173,1464,1485]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB4C40BFD133A9C25B75CCB" box="[1046,1173,1464,1485]" italics="true" pageId="1" pageNumber="62">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
synonymy).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311656DFFB4C408FA423AC723185C7F" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="materials_examined">
|
||
<paragraph id="8BB436E6FFB4C40BFA423AC722DC5E99" blockId="1.[799,1376,1506,2048]" pageId="1" pageNumber="62">
|
||
<emphasis id="B97FEAF4FFB4C40BFA423AC722D55CFA" bold="true" box="[839,1015,1506,1533]" pageId="1" pageNumber="62">Type material.</emphasis>
|
||
The types of
|
||
<taxonomicName id="4C0B4D65FFB4C40BFD9E3AC622AF5F1A" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB4C40BFD9E3AC622AF5F1A" italics="true" pageId="1" pageNumber="62">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
should be in the
|
||
<collectionCode id="ED1AAE23FFB4C40BFD58392725915F1B" box="[1117,1203,1539,1565]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="62" type="Museum">NMNH</collectionCode>
|
||
<collectionCode id="ED1AAE23FFB4C40BFDB83927242B5F1B" box="[1213,1289,1539,1565]" pageId="1" pageNumber="62">NASU</collectionCode>
|
||
collection; despite major effort, none of the labelled type specimens could be located. Since the “
|
||
<typeStatus id="54B08844FFB4C40BFDD73967241B5F5B" box="[1234,1337,1603,1629]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
” of
|
||
<taxonomicName id="4C0B4D65FFB4C40BFA24394725685F7B" box="[801,1098,1635,1661]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB4C40BFA24394725685F7B" box="[801,1098,1635,1661]" italics="true" pageId="1" pageNumber="62">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
was fixed by the author in 1972, but this nominal species taxon was introduced earlier in 1957, this specimen cannot possess
|
||
<typeStatus id="54B08844FFB4C40BFA2439E722A05FDB" box="[801,898,1731,1757]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
status [
|
||
<bibRefCitation id="EF9A4B17FFB4C40BFAD439E0257E5FD8" author="ICZN" box="[977,1116,1732,1758]" pageId="1" pageNumber="62" refId="ref4812" refString="ICZN 1999. International Code of Zoological Nomen- clature. Fourth Edition. London, The International Trust for Zoological Nomenclature, 306 p." type="book" year="1999">ICZN, 1999</bibRefCitation>
|
||
:Art. 72.4.7 and 73.1.3]. Thus, the entire type series dealt with by the author in both publications should be considered
|
||
<typeStatus id="54B08844FFB4C40BFDF23820247F5E18" box="[1271,1373,1796,1822]" pageId="1" pageNumber="62" type="syntype">syntypes</typeStatus>
|
||
. To fix the identity of
|
||
<taxonomicName id="4C0B4D65FFB4C40BFD15380025C15E38" box="[1040,1251,1828,1854]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="62" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB4C40BFD15380025C15E38" box="[1040,1251,1828,1854]" italics="true" pageId="1" pageNumber="62">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
we choose among the
|
||
<typeStatus id="54B08844FFB4C40BFAA23861252B5E59" box="[935,1033,1861,1887]" pageId="1" pageNumber="62" type="syntype">syntypes</typeStatus>
|
||
a single illustrated specimen [Put’, 1972, textfig. 5] and designate it here as the
|
||
<typeStatus id="54B08844FFB4C40BFA2438A022AA5E98" box="[801,904,1924,1950]" pageId="1" pageNumber="62" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<figureCitation id="13302A63FFB4C40BFA9238A022D05E99" box="[919,1010,1924,1951]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="1" pageNumber="62">Fig. 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB4C40BFA42388124425EF9" blockId="1.[799,1376,1506,2048]" pageId="1" pageNumber="62">
|
||
<materialsCitation id="3B633CBBFFB4C40BFA423881247E5EF9" country="Ukraine" location="Originally" pageId="1" pageNumber="62" specimenCount="1" stateProvince="Chernivtsi" typeStatus="holotype">
|
||
<emphasis id="B97FEAF4FFB4C40BFA42388122C05EB8" bold="true" box="[839,994,1956,1983]" pageId="1" pageNumber="62">
|
||
<typeStatus id="54B08844FFB4C40BFA42388122A35EB9" box="[839,897,1957,1983]" pageId="1" pageNumber="62">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8ED4603DFFB4C40BFAE23881257B5EB9" LSID="urn:lsid:plazi:treatment:03A287F0FFB4C40CFA543B7920AC5E60:8ED4603DFFB4C40BFAE23881257B5EB9" box="[999,1113,1957,1983]" country="Ukraine" name="Originally" pageId="1" pageNumber="62" stateProvince="Chernivtsi">Originally</location>
|
||
given as “the right bank of the Dniester River, Rukhotyn village, Khotyn District,
|
||
<collectingRegion id="49CFF804FFB4C40BFA5C38C125005EF9" box="[857,1058,2021,2047]" country="Ukraine" name="Chernivtsi" pageId="1" pageNumber="62">Chernivtsi Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31C7676FFB4C40BFD2F38C125A75EF9" box="[1066,1157,2021,2047]" name="Ukraine" pageId="1" pageNumber="62">Ukraine</collectingCountry>
|
||
” [Put’, 1957, 1972]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF74666EFFB7C408F9AB3F99232A5800" pageId="2" pageNumber="63" startId="2.[174,227,189,211]" targetType="table">
|
||
<paragraph id="8BB436E6FFB7C408F9AB3F99232A5800" blockId="2.[174,1383,189,262]" pageId="2" pageNumber="63">
|
||
Table 1. Examined material of
|
||
<taxonomicName id="4C0B4D65FFB7C408F8DC3F99236359D4" authorityName="Montfort" authorityYear="1810" box="[473,577,189,210]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97FEAF4FFB7C408F8DC3F99236359D4" box="[473,577,189,210]" italics="true" pageId="2" pageNumber="63">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
from the SW Ukraine. The numbers of localities refer to the text and figures. Coordinates of collecting sites sampled prior to 2009 are given approximately, based on data in museum catalogue cards, labels and corresponding publications.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF74666EFFB7C408F9AB3E3222925859" pageId="2" pageNumber="63" startId="2.[174,225,278,300]" targetType="table">
|
||
<paragraph id="8BB436E6FFB7C408F9AB3E3222925859" blockId="2.[174,1384,278,351]" pageId="2" pageNumber="63">
|
||
Табл. 1. ИЗученный материал по
|
||
<taxonomicName id="4C0B4D65FFB7C408F8FB3E332344582A" authorityName="Montfort" authorityYear="1810" box="[510,614,279,300]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97FEAF4FFB7C408F8FB3E332344582A" box="[510,614,279,300]" italics="true" pageId="2" pageNumber="63">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
иЗ ЮЗ Украины. Номера локалитетов соответствуют текстовым ссылкам и подписям к иллюстрациям. Координаты для местонахоЖдений до 2009 года приведены приблиЗительно с учетом данных муЗейных каталогов, Этикеток и соответствуюЩих публикаций.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB436E6FFB7C408F9B83EBE247D5D03" pageId="2" pageNumber="63">
|
||
<table id="F90BC446FFB73BF5F9B93EA9247D5D03" box="[188,1375,397,1029]" gridcols="8" gridrows="11" pageId="2" pageNumber="63">
|
||
<tr id="353B34A4FFB73BF5F9B93EA9247D58BF" box="[188,1375,397,441]" gridrow="0" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93EA921EA58BF" box="[188,200,397,441]" gridcol="0" gridrow="0" pageId="2" pageNumber="63">#</th>
|
||
<th id="76EA5DD8FFB73BF5F9DF3EA920A258BF" box="[218,384,397,441]" gridcol="1" gridrow="0" pageId="2" pageNumber="63">Taxon name</th>
|
||
<th id="76EA5DD8FFB73BF5F8913EA9238A58BF" box="[404,680,397,441]" gridcol="2" gridrow="0" pageId="2" pageNumber="63">Location</th>
|
||
<th id="76EA5DD8FFB73BF5FBB03EA9223758BF" box="[693,789,397,441]" gridcol="3" gridrow="0" pageId="2" pageNumber="63">Date</th>
|
||
<th id="76EA5DD8FFB73BF5FA2E3EA9228258BF" box="[811,928,397,441]" gridcol="4" gridrow="0" pageId="2" pageNumber="63">Latitude (N)</th>
|
||
<th id="76EA5DD8FFB73BF5FABC3EA9251358BF" box="[953,1073,397,441]" gridcol="5" gridrow="0" pageId="2" pageNumber="63">Longitude (E)</th>
|
||
<th id="76EA5DD8FFB73BF5FD4C3EA925C358BF" box="[1097,1249,397,441]" gridcol="6" gridrow="0" pageId="2" pageNumber="63">Collector</th>
|
||
<th id="76EA5DD8FFB73BF5FDF03EA9247D58BF" box="[1269,1375,397,441]" gridcol="7" gridrow="0" pageId="2" pageNumber="63">Collection and lot#</th>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93EE0247D5B0E" box="[188,1375,452,520]" gridrow="1" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93EE021EA5B0E" box="[188,200,452,520]" gridcol="0" gridrow="1" pageId="2" pageNumber="63">1</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3EE020A25B0E" box="[218,384,452,520]" gridcol="1" gridrow="1" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C0B4D65FFB7C408F9DF3EF8206958E9" authorityName="Linnaeus" authorityYear="1758" box="[218,331,476,495]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408F9DF3EF8206958E9" box="[218,331,476,495]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913EE0238A5B0E" box="[404,680,452,520]" gridcol="2" gridrow="1" pageId="2" pageNumber="63">Dniester River, Rukhotyn village, Khotyn District, Chernivtsi Region, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03EE022375B0E" box="[693,789,452,520]" gridcol="3" gridrow="1" pageId="2" pageNumber="63">26 May, 1950</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3EE022825B0E" box="[811,928,452,520]" gridcol="4" gridrow="1" pageId="2" pageNumber="63">48°31’33.83”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3EE025135B0E" box="[953,1073,452,520]" gridcol="5" gridrow="1" pageId="2" pageNumber="63">26°12’9.90”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3EE025C35B0E" box="[1097,1249,452,520]" gridcol="6" gridrow="1" pageId="2" pageNumber="63">A.L. Put’</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03EE0247D5B0E" box="[1269,1375,452,520]" gridcol="7" gridrow="1" pageId="2" pageNumber="63">NMNH NASU, PD, no number</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93D36247D5B3B" box="[188,1375,530,573]" gridrow="2" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93D3621EA5B3B" box="[188,200,530,573]" gridcol="0" gridrow="2" pageId="2" pageNumber="63">2</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3D3620A25B3B" box="[218,384,530,573]" gridcol="1" gridrow="2" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C0B4D65FFB7C408F9DF3D3A20695B37" authorityName="Linnaeus" authorityYear="1758" box="[218,331,542,561]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408F9DF3D3A20695B37" box="[218,331,542,561]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913D36238A5B3B" box="[404,680,530,573]" gridcol="2" gridrow="2" pageId="2" pageNumber="63">Seret River, Ternopil, Ternopil Region, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03D3622375B3B" box="[693,789,530,573]" gridcol="3" gridrow="2" pageId="2" pageNumber="63">10.08.2009</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3D3622825B3B" box="[811,928,530,573]" gridcol="4" gridrow="2" pageId="2" pageNumber="63">49°32’47.21”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3D3625135B3B" box="[953,1073,530,573]" gridcol="5" gridrow="2" pageId="2" pageNumber="63">25°34’47.45”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3D3625C35B3B" box="[1097,1249,530,573]" gridcol="6" gridrow="2" pageId="2" pageNumber="63">Yu.V. Tarasova</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03D36247D5B3B" box="[1269,1375,530,573]" gridcol="7" gridrow="2" pageId="2" pageNumber="63">IZAN, 614</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93D6C247D5B75" box="[188,1375,584,627]" gridrow="3" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93D6C21EA5B75" box="[188,200,584,627]" gridcol="0" gridrow="3" pageId="2" pageNumber="63">3</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3D6C20A25B75" box="[218,384,584,627]" gridcol="1" gridrow="3" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C0B4D65FFB7C408F9DF3D7020695B61" authorityName="Linnaeus" authorityYear="1758" box="[218,331,596,615]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408F9DF3D7020695B61" box="[218,331,596,615]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913D6C238A5B75" box="[404,680,584,627]" gridcol="2" gridrow="3" pageId="2" pageNumber="63">Strypa River, Buchach, Ternopil Region, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03D6C22375B75" box="[693,789,584,627]" gridcol="3" gridrow="3" pageId="2" pageNumber="63">10.08.2009</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3D6C22825B75" box="[811,928,584,627]" gridcol="4" gridrow="3" pageId="2" pageNumber="63">49° 3’38.85”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3D6C25135B75" box="[953,1073,584,627]" gridcol="5" gridrow="3" pageId="2" pageNumber="63">25°24’24.86”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3D6C25C35B75" box="[1097,1249,584,627]" gridcol="6" gridrow="3" pageId="2" pageNumber="63">Yu.V. Tarasova</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03D6C247D5B75" box="[1269,1375,584,627]" gridcol="7" gridrow="3" pageId="2" pageNumber="63">IZAN, 615</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93D5A247D5BAF" box="[188,1375,638,681]" gridrow="4" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93D5A21EA5BAF" box="[188,200,638,681]" gridcol="0" gridrow="4" pageId="2" pageNumber="63">4</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3D5A20A25BAF" box="[218,384,638,681]" gridcol="1" gridrow="4" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C0B4D65FFB7C408F9DF3DAE20695B9B" authorityName="Linnaeus" authorityYear="1758" box="[218,331,650,669]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408F9DF3DAE20695B9B" box="[218,331,650,669]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913D5A238A5BAF" box="[404,680,638,681]" gridcol="2" gridrow="4" pageId="2" pageNumber="63">Hnizna River, Terebovlia, Ternopil Region, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03D5A22375BAF" box="[693,789,638,681]" gridcol="3" gridrow="4" pageId="2" pageNumber="63">10.08.2009</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3D5A22825BAF" box="[811,928,638,681]" gridcol="4" gridrow="4" pageId="2" pageNumber="63">49°18’14.23”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3D5A25135BAF" box="[953,1073,638,681]" gridcol="5" gridrow="4" pageId="2" pageNumber="63">25°41’15.62”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3D5A25C35BAF" box="[1097,1249,638,681]" gridcol="6" gridrow="4" pageId="2" pageNumber="63">Yu.V. Tarasova</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03D5A247D5BAF" box="[1269,1375,638,681]" gridcol="7" gridrow="4" pageId="2" pageNumber="63">IZAN, 616</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93D90247D5A09" box="[188,1375,692,783]" gridrow="5" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93D9021EA5A09" box="[188,200,692,783]" gridcol="0" gridrow="5" pageId="2" pageNumber="63">5</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3D9020A25A09" box="[218,384,692,783]" gridcol="1" gridrow="5" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C0B4D65FFB7C408F9DF3DFC20695BED" authorityName="Linnaeus" authorityYear="1758" box="[218,331,728,747]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408F9DF3DFC20695BED" box="[218,331,728,747]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913D90238A5A09" box="[404,680,692,783]" gridcol="2" gridrow="5" pageId="2" pageNumber="63">Dniester River, Halych village, Ivano-Frankivsk District, Ivano-Frankivsk Region, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03D9022375A09" box="[693,789,692,783]" gridcol="3" gridrow="5" pageId="2" pageNumber="63">29.04.2012</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3D9022825A09" box="[811,928,692,783]" gridcol="4" gridrow="5" pageId="2" pageNumber="63">49º 7’4.59”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3D9025135A09" box="[953,1073,692,783]" gridcol="5" gridrow="5" pageId="2" pageNumber="63">24º45’ 7.56”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3D9025C35A09" box="[1097,1249,692,783]" gridcol="6" gridrow="5" pageId="2" pageNumber="63">E.V. Degtyarenko</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03D90247D5A09" box="[1269,1375,692,783]" gridcol="7" gridrow="5" pageId="2" pageNumber="63">IZAN, 335</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93C3E247D5A73" box="[188,1375,794,885]" gridrow="6" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93C3E21EA5A73" box="[188,200,794,885]" gridcol="0" gridrow="6" pageId="2" pageNumber="63">6</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3C3E20A25A73" box="[218,384,794,885]" gridcol="1" gridrow="6" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C0B4D65FFB7C408F9DF3C1A20695A57" authorityName="Linnaeus" authorityYear="1758" box="[218,331,830,849]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408F9DF3C1A20695A57" box="[218,331,830,849]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913C3E238A5A73" box="[404,680,794,885]" gridcol="2" gridrow="6" pageId="2" pageNumber="63">Dniester River, Monastyrok village, Ivano-Frankivsk District, Kolomyia Region, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03C3E22375A73" box="[693,789,794,885]" gridcol="3" gridrow="6" pageId="2" pageNumber="63">03.05.2012</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3C3E22825A73" box="[811,928,794,885]" gridcol="4" gridrow="6" pageId="2" pageNumber="63">48º49’24.39”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3C3E25135A73" box="[953,1073,794,885]" gridcol="5" gridrow="6" pageId="2" pageNumber="63">25º15’17.27”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3C3E25C35A73" box="[1097,1249,794,885]" gridcol="6" gridrow="6" pageId="2" pageNumber="63">E.V. Degtyarenko</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03C3E247D5A73" box="[1269,1375,794,885]" gridcol="7" gridrow="6" pageId="2" pageNumber="63">IZAN, 333</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93CA4247D5AAD" box="[188,1375,896,939]" gridrow="7" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93CA421EA5AAD" box="[188,200,896,939]" gridcol="0" gridrow="7" pageId="2" pageNumber="63">7</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3CA420A25AAD" box="[218,384,896,939]" gridcol="1" gridrow="7" pageId="2" pageNumber="63">
|
||
“
|
||
<taxonomicName id="4C0B4D65FFB7C408F9E13CA820435A99" box="[228,353,908,927]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="obliteratus">
|
||
<emphasis id="B97FEAF4FFB7C408F9E13CA820435A99" box="[228,353,908,927]" italics="true" pageId="2" pageNumber="63">Th. obliteratus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913CA4238A5AAD" box="[404,680,896,939]" gridcol="2" gridrow="7" pageId="2" pageNumber="63">Dniester River near Rybnitsa, Moldova</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03CA422375AAD" box="[693,789,896,939]" gridcol="3" gridrow="7" pageId="2" pageNumber="63">no date,?1902</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3CA422825AAD" box="[811,928,896,939]" gridcol="4" gridrow="7" pageId="2" pageNumber="63">47°47’26.59”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3CA425135AAD" box="[953,1073,896,939]" gridcol="5" gridrow="7" pageId="2" pageNumber="63">28°59’25.44”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3CA425C35AAD" box="[1097,1249,896,939]" gridcol="6" gridrow="7" pageId="2" pageNumber="63">A.A. Brauner</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03CA4247D5AAD" box="[1269,1375,896,939]" gridcol="7" gridrow="7" pageId="2" pageNumber="63">ZIN, 5912/1</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93C91247D5ACF" box="[188,1375,949,969]" gridrow="8" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93C9121EA5ACF" box="[188,200,949,969]" gridcol="0" gridrow="8" pageId="2" pageNumber="63">8</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3C9120A25ACF" box="[218,384,949,969]" gridcol="1" gridrow="8" pageId="2" pageNumber="63">
|
||
“
|
||
<taxonomicName id="4C0B4D65FFB7C408F9E13C9220525ACF" box="[228,368,950,969]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="alboguttatus">
|
||
<emphasis id="B97FEAF4FFB7C408F9E13C9220525ACF" box="[228,368,950,969]" italics="true" pageId="2" pageNumber="63">Th. alboguttatus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913C91238A5ACF" box="[404,680,949,969]" gridcol="2" gridrow="8" pageId="2" pageNumber="63">Odessa, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03C9122375ACF" box="[693,789,949,969]" gridcol="3" gridrow="8" pageId="2" pageNumber="63">28.12.1902</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3C9122825ACF" box="[811,928,949,969]" gridcol="4" gridrow="8" pageId="2" pageNumber="63">46°31’40.44”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3C9125135ACF" box="[953,1073,949,969]" gridcol="5" gridrow="8" pageId="2" pageNumber="63">30°43’18.74”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3C9125C35ACF" box="[1097,1249,949,969]" gridcol="6" gridrow="8" pageId="2" pageNumber="63">A.A. Brauner</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03C91247D5ACF" box="[1269,1375,949,969]" gridcol="7" gridrow="8" pageId="2" pageNumber="63">ZIN, 6051/1</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93CF7247D5AE1" box="[188,1375,979,999]" gridrow="9" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93CF721EA5AE1" box="[188,200,979,999]" gridcol="0" gridrow="9" pageId="2" pageNumber="63">8</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3CF720A25AE1" box="[218,384,979,999]" gridcol="1" gridrow="9" pageId="2" pageNumber="63">
|
||
“
|
||
<taxonomicName id="4C0B4D65FFB7C408F9E13CF0207C5AE1" box="[228,350,980,999]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="lacrymans">
|
||
<emphasis id="B97FEAF4FFB7C408F9E13CF0207C5AE1" box="[228,350,980,999]" italics="true" pageId="2" pageNumber="63">Th. lacrymans</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913CF7238A5AE1" box="[404,680,979,999]" gridcol="2" gridrow="9" pageId="2" pageNumber="63">Odessa, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03CF722375AE1" box="[693,789,979,999]" gridcol="3" gridrow="9" pageId="2" pageNumber="63">28.12.1902</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3CF722825AE1" box="[811,928,979,999]" gridcol="4" gridrow="9" pageId="2" pageNumber="63">46°31’40.44”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3CF725135AE1" box="[953,1073,979,999]" gridcol="5" gridrow="9" pageId="2" pageNumber="63">30°43’18.74”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3CF725C35AE1" box="[1097,1249,979,999]" gridcol="6" gridrow="9" pageId="2" pageNumber="63">A.A. Brauner</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03CF7247D5AE1" box="[1269,1375,979,999]" gridcol="7" gridrow="9" pageId="2" pageNumber="63">ZIN, 6052/1</td>
|
||
</tr>
|
||
<tr id="353B34A4FFB73BF5F9B93CD5247D5D03" box="[188,1375,1009,1029]" gridrow="10" pageId="2" pageNumber="63">
|
||
<th id="76EA5DD8FFB73BF5F9B93CD521EA5D03" box="[188,200,1009,1029]" gridcol="0" gridrow="10" pageId="2" pageNumber="63">8</th>
|
||
<td id="76EA5DD8FFB73BF5F9DF3CD520A25D03" box="[218,384,1009,1029]" gridcol="1" gridrow="10" pageId="2" pageNumber="63">
|
||
“
|
||
<taxonomicName id="4C0B4D65FFB7C408F9E13CD520555D02" box="[228,375,1009,1028]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="pulcherrimus">
|
||
<emphasis id="B97FEAF4FFB7C408F9E13CD520555D02" box="[228,375,1009,1028]" italics="true" pageId="2" pageNumber="63">Th. pulcherrimus</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
</td>
|
||
<td id="76EA5DD8FFB73BF5F8913CD5238A5D03" box="[404,680,1009,1029]" gridcol="2" gridrow="10" pageId="2" pageNumber="63">Odessa, Ukraine</td>
|
||
<td id="76EA5DD8FFB73BF5FBB03CD522375D03" box="[693,789,1009,1029]" gridcol="3" gridrow="10" pageId="2" pageNumber="63">28.12.1902</td>
|
||
<td id="76EA5DD8FFB73BF5FA2E3CD522825D03" box="[811,928,1009,1029]" gridcol="4" gridrow="10" pageId="2" pageNumber="63">46°31’40.44”</td>
|
||
<td id="76EA5DD8FFB73BF5FABC3CD525135D03" box="[953,1073,1009,1029]" gridcol="5" gridrow="10" pageId="2" pageNumber="63">30°43’18.74”</td>
|
||
<td id="76EA5DD8FFB73BF5FD4C3CD525C35D03" box="[1097,1249,1009,1029]" gridcol="6" gridrow="10" pageId="2" pageNumber="63">A.A. Brauner</td>
|
||
<td id="76EA5DD8FFB73BF5FDF03CD5247D5D03" box="[1269,1375,1009,1029]" gridcol="7" gridrow="10" pageId="2" pageNumber="63">ZIN, 6053/1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB7C408F9D13B7B23185C7F" blockId="2.[172,750,1119,2042]" pageId="2" pageNumber="63">
|
||
<emphasis id="B97FEAF4FFB7C408F9D13B7B20B25D7F" bold="true" box="[212,400,1119,1145]" pageId="2" pageNumber="63">Other material.</emphasis>
|
||
More than 100 shells labelled as “
|
||
<taxonomicName id="4C0B4D65FFB7C408F9D93B5B20EA5D9F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[220,456,1151,1177]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408F9D93B5B20EA5D9F" box="[220,456,1151,1177]" italics="true" pageId="2" pageNumber="63">Theodoxus fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
” collected from the type locality by Put’ on
|
||
<date id="FFB51026FFB7C408F8833B8423075DBC" box="[390,549,1183,1210]" pageId="2" pageNumber="63" value="1950-05-26">May 26, 1950</date>
|
||
were retrieved in the
|
||
<collectionCode id="ED1AAE23FFB7C408F9DE3BE420135DDC" box="[219,305,1216,1242]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="63" type="Museum">NMNH</collectionCode>
|
||
<collectionCode id="ED1AAE23FFB7C408F83E3B9B20A55DDF" box="[315,391,1215,1241]" pageId="2" pageNumber="63">NASU</collectionCode>
|
||
depository. The specimens are considered as
|
||
<typeStatus id="54B08844FFB7C408F8413BC4208C5DFC" box="[324,430,1248,1274]" pageId="2" pageNumber="63" type="topotype">topotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0B4D65FFB7C408F8C83BFB23C85DFF" box="[461,746,1247,1273]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB7C408F8C83BFB23C85DFF" box="[461,746,1247,1273]" italics="true" pageId="2" pageNumber="63">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
here. Additionally, over hundred individuals of
|
||
<taxonomicName id="4C0B4D65FFB7C408FBCC3BDB20375C3F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408FBCC3BDB20375C3F" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
from other points of the Dniester River and its tributaries, namely the Seret, Strypa, and Hnizna rivers (localities 2–6) were studied.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C311656DFFB7C40CF9D13A5B20AC5E60" lastPageId="6" lastPageNumber="67" pageId="2" pageNumber="63" type="discussion">
|
||
<paragraph id="8BB436E6FFB7C408F9D13A5B20EE5EBF" blockId="2.[172,750,1119,2042]" pageId="2" pageNumber="63">
|
||
<emphasis id="B97FEAF4FFB7C408F9D13A5B206A5C9F" bold="true" box="[212,328,1407,1433]" pageId="2" pageNumber="63">Remarks.</emphasis>
|
||
The species name “
|
||
<taxonomicName id="4C0B4D65FFB7C408FB3E3A5B20135CBF" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB7C408FB3E3A5B20135CBF" italics="true" pageId="2" pageNumber="63">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” has been considered as established by Put’ in 1972 [e.g. O.
|
||
<bibRefCitation id="EF9A4B17FFB7C408F8CD3A9B23CA5CDF" author="Anistratenko O. Yu. & Starobogatov Ya. I. & Anistratenko V. V." box="[456,744,1471,1498]" pageId="2" pageNumber="63" pagination="11 - 19" refId="ref4413" refString="Anistratenko O. Yu., Starobogatov Ya. I., Anistratenko V. V. 1999. Mollusks of the genus Theodoxus (Gastropoda, Pectinibranchia, Neritidae) of the Black and the Azov seas basin. Vestnik zoologii, 33: 11 - 19 [In Russian with English summary]." type="journal article" year="1999">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB7C408FB673AE423B75CDF" box="[610,661,1471,1497]" italics="true" pageId="2" pageNumber="63">et al</emphasis>
|
||
., 1999
|
||
</bibRefCitation>
|
||
;
|
||
<collectionCode id="ED1AAE23FFB7C408F9AB3AC421E05CFC" box="[174,194,1504,1530]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="63" type="Museum">V</collectionCode>
|
||
.
|
||
<bibRefCitation id="EF9A4B17FFB7C408F9CC3AFB20FC5CFF" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." box="[201,478,1502,1530]" pageId="2" pageNumber="63" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB7C408F85E3AC420A85CFF" box="[347,394,1503,1529]" italics="true" pageId="2" pageNumber="63">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
]. This dating uncritically relies on the article describing “
|
||
<taxonomicName id="4C0B4D65FFB7C408FB0F3ADB21D85F3F" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB7C408FB0F3ADB21D85F3F" italics="true" pageId="2" pageNumber="63">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
Put’ sp. nova.” in detail morphologically and providing a shell photo [Put’, 1972: 80-82, textfig. 5]. However, another publication by Put’ [1957] was recently found where the nominal name “
|
||
<taxonomicName id="4C0B4D65FFB7C408FB76395B20775FBF" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB7C408FB76395B20775FBF" italics="true" pageId="2" pageNumber="63">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” was established, and the species description satisfies the
|
||
<bibRefCitation id="EF9A4B17FFB7C408F8B2399B236B5FDF" author="ICZN" box="[439,585,1726,1753]" pageId="2" pageNumber="63" refId="ref4812" refString="ICZN 1999. International Code of Zoological Nomen- clature. Fourth Edition. London, The International Trust for Zoological Nomenclature, 306 p." type="book" year="1999">ICZN [1999]</bibRefCitation>
|
||
guidelines applied to species described between 1930 and 1999. In both these publications [Put’, 1957, 1972], the author clearly states that the new species name is introduced, lists the same diagnostic features and indicates the same
|
||
<typeStatus id="54B08844FFB7C408F884384420935E7C" box="[385,433,1888,1914]" pageId="2" pageNumber="63">type</typeStatus>
|
||
locality. Thus the name was made nomenclaturally available in 1957, not in 1972 as it was accepted before.
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB7C408F9D138E424065C1C" blockId="2.[172,750,1119,2042]" lastBlockId="2.[807,1386,1119,2042]" pageId="2" pageNumber="63">
|
||
To ascertain the validity of this name, one has to check whether the shells of “
|
||
<taxonomicName id="4C0B4D65FFB7C408F8E738FB238F5EFF" box="[482,685,2015,2041]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB7C408F8E738FB238F5EFF" box="[482,685,2015,2041]" italics="true" pageId="2" pageNumber="63">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” provide unique diagnostic features or their morphology falls within the range of variability of other species of the genus in this or neighboring region. In fact the
|
||
<typeStatus id="54B08844FFB7C408FA573BE422A05DDC" box="[850,898,1216,1242]" pageId="2" pageNumber="63">type</typeStatus>
|
||
locality of “
|
||
<taxonomicName id="4C0B4D65FFB7C408FD153B9B25C15DDF" box="[1040,1251,1215,1241]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB7C408FD153B9B25C15DDF" box="[1040,1251,1215,1241]" italics="true" pageId="2" pageNumber="63">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” is situated within the range of
|
||
<taxonomicName id="4C0B4D65FFB7C408FD053BFB25B05DFF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1024,1170,1247,1273]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408FD053BFB25B05DFF" box="[1024,1170,1247,1273]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
, the most common and widespread species of neritids in Europe.
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB7C408FA4A3A3B229A5F1C" blockId="2.[807,1386,1119,2042]" pageId="2" pageNumber="63">
|
||
The identity of the name
|
||
<taxonomicName id="4C0B4D65FFB7C408FD753A3B24275C3F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1136,1285,1311,1337]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408FD753A3B24275C3F" box="[1136,1285,1311,1337]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
is determined by its
|
||
<typeStatus id="54B08844FFB7C408FAB23A1B253C5C5F" box="[951,1054,1343,1369]" pageId="2" pageNumber="63" type="lectotype">lectotype</typeStatus>
|
||
[O.
|
||
<bibRefCitation id="EF9A4B17FFB7C408FD483A1B24415C5C" author="Anistratenko O. Yu. & Starobogatov Ya. I. & Anistratenko V. V." box="[1101,1379,1343,1370]" pageId="2" pageNumber="63" pagination="11 - 19" refId="ref4413" refString="Anistratenko O. Yu., Starobogatov Ya. I., Anistratenko V. V. 1999. Mollusks of the genus Theodoxus (Gastropoda, Pectinibranchia, Neritidae) of the Black and the Azov seas basin. Vestnik zoologii, 33: 11 - 19 [In Russian with English summary]." type="journal article" year="1999">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB7C408FDE13A6424365C5F" box="[1252,1300,1343,1369]" italics="true" pageId="2" pageNumber="63">et al</emphasis>
|
||
., 1999
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB7C408FA2D3A7B22DE5C7C" author="Anistratenko V. V." box="[808,1020,1375,1402]" pageId="2" pageNumber="63" pagination="3 - 10" refId="ref4469" refString="Anistratenko V. V. 2005. Lectotypes for Tricolia pul- lus, Gibbula divaricata and Theodoxus fluviatilis revisited. Vestnik zoologii, 39 (6): 3 - 10." type="journal article" year="2005">Anistratenko, 2005</bibRefCitation>
|
||
] though the species demonstrates a considerable variation in its shell colour pattern and shape [e.g.
|
||
<bibRefCitation id="EF9A4B17FFB7C408FAA73ABB25115CBC" author="Zettler M. L." box="[930,1075,1439,1466]" pageId="2" pageNumber="63" pagination="13 - 72" refId="ref5394" refString="Zettler M. L. 2008. Zur Taxonomie und Verbreitung der Gattung Theodoxus Montfort, 1810 in Deutschland. Darstellung historischer und rezenter Daten einschliesslich einer Bibliografie. Mollusca, 26: 13 - 72." type="journal article" year="2008">Zettler, 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB7C408FD393ABB242E5CBC" author="Gloer P & Pesic V" box="[1084,1292,1439,1466]" pageId="2" pageNumber="63" pagination="88 - 92" refId="ref4761" refString="Gloer P, Pesic V (2015) The morphological plasticity of Theodoxus fluviatilis (Linnaeus, 1758) (Mollusca: Gastropoda: Neritidae). Ecologica Montenegrina, 2 (2): 88 - 92. https: // doi. org / 10.37828 / em. 2015.2.10" type="journal article" year="2015">Glöer, Pešić, 2015</bibRefCitation>
|
||
]. That is why a large number of nominal taxa are considered as synonyms of
|
||
<taxonomicName id="4C0B4D65FFB7C408FAE33AFB25595CFF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[998,1147,1503,1529]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB7C408FAE33AFB25595CFF" box="[998,1147,1503,1529]" italics="true" pageId="2" pageNumber="63">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
[see
|
||
<collectionCode id="ED1AAE23FFB7C408FDBD3AC425EE5CFC" box="[1208,1228,1504,1530]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="63" type="Museum">V</collectionCode>
|
||
.
|
||
<bibRefCitation id="EF9A4B17FFB7C408FDD23AFB228F5F1C" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." pageId="2" pageNumber="63" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB7C408FA2C392422785F1F" box="[809,858,1535,1561]" italics="true" pageId="2" pageNumber="63">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB7C408FA4A393B256C5EDC" blockId="2.[807,1386,1119,2042]" pageId="2" pageNumber="63">
|
||
The original description of
|
||
<taxonomicName id="4C0B4D65FFB7C408FD90393B24485F3F" box="[1173,1386,1567,1593]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="2" pageNumber="63" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB7C408FD90393B24485F3F" box="[1173,1386,1567,1593]" italics="true" pageId="2" pageNumber="63">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
was based on a relatively few morphological characters. No peculiarities in the size and/or proportions of the shell are mentioned [Put’, 1957, 1972] or depicted (
|
||
<figureCitation id="13302A63FFB7C408FA3439BB22AE5FBC" box="[817,908,1695,1722]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="2" pageNumber="63">Fig. 4D</figureCitation>
|
||
). The diagnosis is focused only on periostracum colouration and number of light spiral stripes on the last whorl of the shell: these can be accounted from 3–10 up to 15 depending on individuals’ age [Put’, 1957, 1972]. As it is known, these characters are prone to considerable intraspecific variation [
|
||
<bibRefCitation id="EF9A4B17FFB7C408FA2A387B22E75E7C" author="Bandel K." box="[815,965,1887,1914]" pageId="2" pageNumber="63" pagination="65 - 164" refId="ref4721" refString="Bandel K. 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mit- teilungen aus dem Geologisch-Palaontologischen Institut Universitat der Hamburg, 85: 65 - 164." type="journal article" year="2001">Bandel, 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB7C408FAC8387B25F85E7C" author="Anistratenko V. V. & Zettler M. L. & Anistratenko O. Yu." box="[973,1242,1887,1914]" pageId="2" pageNumber="63" pagination="213 - 226" refId="ref4650" refString="Anistratenko V. V., Zettler M. L., Anistratenko O. Yu. 2017. On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region. Archiv fur Molluskenkunde, 146 (2): 213 - 226. https: // doi. org / 10.1127 / arch. moll / 146 / 213 - 226" type="journal article" year="2017">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB7C408FD5A384425AF5E7F" box="[1119,1165,1887,1913]" italics="true" pageId="2" pageNumber="63">et al</emphasis>
|
||
., 2017
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9A4B17FFB7C408FDE73844243E5E7C" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." box="[1250,1308,1888,1914]" pageId="2" pageNumber="63" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">2020</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB7C408FC23387B228E5E9C" author="Sands A. F. & Gloer P. & Gurlek M. E. & Albrecht C. & T. A" pageId="2" pageNumber="63" pagination="25 - 66" refId="ref5061" refString="Sands A. F., Gloer P., Gurlek M. E., Albrecht C., Neu- bauer T. A. 2020. A revision of the extant species of Theodoxus (Gastropoda, Neritidae) inAsia, with the description of three new species. Zoosystematics and Evolution, 96 (1): 25 - 66. https: // zse. pensoft. net / article / 48312 /" type="journal article" year="2020">
|
||
Sands
|
||
<emphasis id="B97FEAF4FFB7C408FA2238A4227B5E9F" box="[807,857,1919,1945]" italics="true" pageId="2" pageNumber="63">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
] and certainly related to environmental conditions [e.g.
|
||
<bibRefCitation id="EF9A4B17FFB7C408FADD388425A95EBC" author="Neumann D." box="[984,1163,1951,1978]" pageId="2" pageNumber="63" pagination="349 - 411" refId="ref4870" refString="Neumann D. 1959. Morphologische und experimentelle Untersuchungen uber die Variabilitat der Farb- muster auf der Schale von Theodoxus fluviatilis L. Zeitschrift fur Morphologie und Okologie der Tiere, 48 (4): 349 - 411. https: // doi. org / 10.1007 / BF 00408578" type="journal article" year="1959">Neumann, 1959</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB7C408FD913884242C5EBC" author="Rust J." box="[1172,1294,1952,1978]" pageId="2" pageNumber="63" pagination="37 - 180" refId="ref5027" refString="Rust J. 1997. Evolution, Systematik, Palaookologie und stratigraphischer Nutzenneogener Suss- und Brackwassergastropoden im Nord-Agais-Raum. Palaeontographica Abteilung, A 243 (1 - 6): 37 - 180." type="journal article" year="1997">Rust, 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB7C408FC1D38BB22465EDC" author="Zettler M. L." pageId="2" pageNumber="63" pagination="245 - 251" refId="ref5356" refString="Zettler M. L. 2007. A redescription of Theodoxus schul- tzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea. Journal of Conchology, 39: 245 - 251." type="journal article" year="2007">Zettler, 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB7C408FA75389B25615EDC" author="Gloer P & Pesic V" box="[880,1091,1983,2010]" pageId="2" pageNumber="63" pagination="88 - 92" refId="ref4761" refString="Gloer P, Pesic V (2015) The morphological plasticity of Theodoxus fluviatilis (Linnaeus, 1758) (Mollusca: Gastropoda: Neritidae). Ecologica Montenegrina, 2 (2): 88 - 92. https: // doi. org / 10.37828 / em. 2015.2.10" type="journal article" year="2015">Glöer, Pešić, 2015</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB7C409FA4A38C422915E9C" blockId="2.[807,1386,1119,2042]" lastBlockId="3.[800,1377,1792,2043]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
|
||
In general, the great conchological variability makes the reliable identification of some
|
||
<taxonomicName id="4C0B4D65FFB6C409FB6E382421D55E3C" pageId="3" pageNumber="64">
|
||
<emphasis id="B97FEAF4FFB6C409FB6E382423C65E1C" box="[619,740,1792,1818]" italics="true" pageId="3" pageNumber="64">Theodoxus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
difficult and raises debates on their validity. For example, there is no complete consensus on the taxonomy and nomenclature of
|
||
<taxonomicName id="4C0B4D65FFB6C409FB16384423C75E7C" box="[531,741,1888,1914]" pageId="3" pageNumber="64">
|
||
<emphasis id="B97FEAF4FFB6C409FB16384423AE5E7C" box="[531,652,1888,1914]" italics="true" pageId="3" pageNumber="64">Theodoxus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
inhabiting the river basins of the Black Sea region [
|
||
<bibRefCitation id="EF9A4B17FFB6C409F9A9388420E85EBC" author="Anistratenko V. V. & Zettler M. L. & Anistratenko O. Yu." box="[172,458,1952,1978]" pageId="3" pageNumber="64" pagination="213 - 226" refId="ref4650" refString="Anistratenko V. V., Zettler M. L., Anistratenko O. Yu. 2017. On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region. Archiv fur Molluskenkunde, 146 (2): 213 - 226. https: // doi. org / 10.1127 / arch. moll / 146 / 213 - 226" type="journal article" year="2017">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB6C409F84D388520585EBC" box="[328,378,1952,1978]" italics="true" pageId="3" pageNumber="64">et al</emphasis>
|
||
., 2017
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9A4B17FFB6C409F8D3388423305EBC" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." box="[470,530,1952,1978]" pageId="3" pageNumber="64" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">2020</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB6C409FB18388421C35EDC" author="Wesselingh F. P. & Neubauer T. A. & Anistratenko V. V. & Vinarski M. V. & Yanina T. & ter Poorten J. J. & Kijashko P. & Albrecht C. & Anistratenko O. & Yu & D'Hont A. & Frolov P. & Gandara A. M. & Gittenberger A. & Gogaladze A. & Karpinsky M. & Lattuada M. & Popa L. & Sands A. F. & van de Velde S. & Vandendorpe J. & Wilke T." pageId="3" pageNumber="64" pagination="31 - 124" refId="ref5221" refString="Wesselingh F. P., Neubauer T. A., Anistratenko V. V., Vinarski M. V., Yanina T., ter Poorten J. J., Kijashko P., Albrecht C., Anistratenko O. Yu, D'Hont A., Frolov P., Gandara A. M., Gittenberger A., Gogaladze A., Karpinsky M., Lattuada M., Popa L., Sands A. F., van de Velde S., Vandendorpe J., Wilke T. 2019. Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys, 827: 31 - 124. https: // doi. org / 10.3897 / zookeys. 827.31365" type="journal article" year="2019">
|
||
Wesselingh
|
||
<emphasis id="B97FEAF4FFB6C409FBA3388523F55EBC" box="[678,727,1952,1978]" italics="true" pageId="3" pageNumber="64">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB6C409F9EE38E420915EDC" author="Sands A. F. & Gloer P. & Gurlek M. E. & Albrecht C. & T. A" box="[235,435,1983,2010]" pageId="3" pageNumber="64" pagination="25 - 66" refId="ref5061" refString="Sands A. F., Gloer P., Gurlek M. E., Albrecht C., Neu- bauer T. A. 2020. A revision of the extant species of Theodoxus (Gastropoda, Neritidae) inAsia, with the description of three new species. Zoosystematics and Evolution, 96 (1): 25 - 66. https: // zse. pensoft. net / article / 48312 /" type="journal article" year="2020">
|
||
Sands
|
||
<emphasis id="B97FEAF4FFB6C409F83438E520435EDC" box="[305,353,1984,2010]" italics="true" pageId="3" pageNumber="64">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
].At the same time it is demonstrated that the most widely-distributed in Europe
|
||
<taxonomicName id="4C0B4D65FFB6C409FA25382422905E1C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[800,946,1792,1818]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="64" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB6C409FA25382422905E1C" box="[800,946,1792,1818]" italics="true" pageId="3" pageNumber="64">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
comprises many different variations in shell shape and colour pattern [O.
|
||
<bibRefCitation id="EF9A4B17FFB6C409FDB4380422A85E5C" author="Anistratenko O. Yu. & Starobogatov Ya. I. & Anistratenko V. V." pageId="3" pageNumber="64" pagination="11 - 19" refId="ref4413" refString="Anistratenko O. Yu., Starobogatov Ya. I., Anistratenko V. V. 1999. Mollusks of the genus Theodoxus (Gastropoda, Pectinibranchia, Neritidae) of the Black and the Azov seas basin. Vestnik zoologii, 33: 11 - 19 [In Russian with English summary]." type="journal article" year="1999">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB6C409FC49380522155E5C" italics="true" pageId="3" pageNumber="64">et al</emphasis>
|
||
., 1999
|
||
</bibRefCitation>
|
||
;
|
||
<collectionCode id="ED1AAE23FFB6C409FA9C3864228F5E5C" box="[921,941,1856,1882]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="64" type="Museum">V</collectionCode>
|
||
. Anistratenko, O.
|
||
<bibRefCitation id="EF9A4B17FFB6C409FD7B386424795E5C" author="Anistratenko V. V. & Anistratenko O. Yu." box="[1150,1371,1856,1882]" pageId="3" pageNumber="64" pagination="1 - 240" refId="ref4501" refString="Anistratenko V. V., Anistratenko O. Yu. 2001. Class Polyplacophora or Chitons, Class Gastropoda - Cyclobranchia, Scutibranchia and Pectinibranchia (part). Fauna of Ukraine, Mollusca, 29 (1 / 1). Kiev: Veles: 1 - 240 [In Russian with extended English summary and captions]." type="book chapter" year="2001">Anistratenko, 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB6C409FA24384422995E7C" author="Bandel K." box="[801,955,1888,1914]" pageId="3" pageNumber="64" pagination="65 - 164" refId="ref4721" refString="Bandel K. 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mit- teilungen aus dem Geologisch-Palaontologischen Institut Universitat der Hamburg, 85: 65 - 164." type="journal article" year="2001">Bandel, 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB6C409FACC384425495E7C" author="Shubrat Yu." box="[969,1131,1888,1914]" pageId="3" pageNumber="64" pagination="154 - 159" refId="ref5138" refString="Shubrat Yu. 2007. Complex analysis of the conchological indications of Theodoxus species of Ukraine. Visnyk of Lviv University, Biology series, 45: 154 - 159 [In Ukrainian with English summary]." type="journal article" year="2007">Shubrat, 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB6C409FD7D3844242F5E7C" author="Zettler M. L." box="[1144,1293,1888,1914]" pageId="3" pageNumber="64" pagination="13 - 72" refId="ref5394" refString="Zettler M. L. 2008. Zur Taxonomie und Verbreitung der Gattung Theodoxus Montfort, 1810 in Deutschland. Darstellung historischer und rezenter Daten einschliesslich einer Bibliografie. Mollusca, 26: 13 - 72." type="journal article" year="2008">Zettler, 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB6C409FC1F3844228A5E9C" author="Gloer P & Pesic V" pageId="3" pageNumber="64" pagination="88 - 92" refId="ref4761" refString="Gloer P, Pesic V (2015) The morphological plasticity of Theodoxus fluviatilis (Linnaeus, 1758) (Mollusca: Gastropoda: Neritidae). Ecologica Montenegrina, 2 (2): 88 - 92. https: // doi. org / 10.37828 / em. 2015.2.10" type="journal article" year="2015">Glöer, Pešić, 2015</bibRefCitation>
|
||
].
|
||
</paragraph>
|
||
<caption id="DF74666EFFB6C409F9AD391C203E5FA0" ID-DOI="http://doi.org/10.5281/zenodo.11040988" ID-Zenodo-Dep="11040988" httpUri="https://zenodo.org/record/11040988/files/figure.png" pageId="3" pageNumber="64" startId="3.[168,210,1592,1613]" targetBox="[168,1376,189,1562]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BB436E6FFB6C409F9AD391C20745F60" blockId="3.[168,1376,1591,1702]" pageId="3" pageNumber="64">
|
||
FIG. 1. Map of the study area with indication of sampling points of
|
||
<taxonomicName id="4C0B4D65FFB6C409FA30391322BF5F4A" authorityName="Montfort" authorityYear="1810" box="[821,925,1591,1612]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="64" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97FEAF4FFB6C409FA30391322BF5F4A" box="[821,925,1591,1612]" italics="true" pageId="3" pageNumber="64">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
. The numbers of localities correspond to those in the Table 1.
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB6C409F9AD395C203E5FA0" blockId="3.[168,1376,1591,1702]" pageId="3" pageNumber="64">
|
||
РИС. 1. Карта иЗученного региона с укаЗанием точек сбора
|
||
<taxonomicName id="4C0B4D65FFB6C409FA0A395322555F8A" authorityName="Montfort" authorityYear="1810" box="[783,887,1655,1676]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="64" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97FEAF4FFB6C409FA0A395322555F8A" box="[783,887,1655,1676]" italics="true" pageId="3" pageNumber="64">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
. Номера локалитетов соответствуют таковым в Табл. 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB436E6FFB6C40CFA423884200E5C06" blockId="3.[800,1377,1792,2043]" lastBlockId="6.[172,750,188,1894]" lastPageId="6" lastPageNumber="67" pageId="3" pageNumber="64">
|
||
The distribution range of the nominal species “
|
||
<taxonomicName id="4C0B4D65FFB6C409FA2B38E425755EDC" box="[814,1111,1984,2010]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="64" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB6C409FA2B38E425755EDC" box="[814,1111,1984,2010]" italics="true" pageId="3" pageNumber="64">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” is limited by the type locality only [Put’, 1957, 1972]. The
|
||
<typeStatus id="54B08844FFB6C409FDD138C424635EFC" box="[1236,1345,2016,2042]" pageId="3" pageNumber="64" type="topotype">topotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9AB3F98205E59D0" box="[174,380,188,214]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB3C40CF9AB3F98205E59D0" box="[174,380,188,214]" italics="true" pageId="6" pageNumber="67">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
show a diversity of pattern (
|
||
<figureCitation id="13302A63FFB3C40CFBA83F9821C059F1" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4 A–C</figureCitation>
|
||
) that is registered in
|
||
<taxonomicName id="4C0B4D65FFB3C40CF8B83FF9236E59F1" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[445,588,221,247]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB3C40CF8B83FF9236E59F1" box="[445,588,221,247]" italics="true" pageId="6" pageNumber="67">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
populations in many parts of its range [
|
||
<collectionCode id="ED1AAE23FFB3C40CF8B63FDA20EB581E" box="[435,457,254,280]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="67" type="Museum">V</collectionCode>
|
||
.
|
||
<bibRefCitation id="EF9A4B17FFB3C40CF8D63FD923CB5811" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." box="[467,745,253,280]" pageId="6" pageNumber="67" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB3C40CFB6C3FDA23BB5811" box="[617,665,253,279]" italics="true" pageId="6" pageNumber="67">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9A4B17FFB3C40CF9A93E392055583E" author="Sands A. F. & Gloer P. & Gurlek M. E. & Albrecht C. & T. A" box="[172,375,285,312]" pageId="6" pageNumber="67" pagination="25 - 66" refId="ref5061" refString="Sands A. F., Gloer P., Gurlek M. E., Albrecht C., Neu- bauer T. A. 2020. A revision of the extant species of Theodoxus (Gastropoda, Neritidae) inAsia, with the description of three new species. Zoosystematics and Evolution, 96 (1): 25 - 66. https: // zse. pensoft. net / article / 48312 /" type="journal article" year="2020">
|
||
Sands
|
||
<emphasis id="B97FEAF4FFB3C40CF9F33E3A20045831" box="[246,294,285,311]" italics="true" pageId="6" pageNumber="67">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
]. It should be noted that among all available
|
||
<typeStatus id="54B08844FFB3C40CF81E3E1A20AA585E" box="[283,392,318,344]" pageId="6" pageNumber="67" type="topotype">topotypes</typeStatus>
|
||
no shell exactly corresponding in colouration and pattern to the one depicted in the 1972 paper [Put’, 1972: 80-82, textfig. 5] has been found (
|
||
<figureCitation id="13302A63FFB3C40CF8013EBA204658BF" box="[260,356,414,441]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4D</figureCitation>
|
||
). However, we found shells with longitudinal stripes in
|
||
<taxonomicName id="4C0B4D65FFB3C40CF8BA3E9B237958DF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[447,603,447,473]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB3C40CF8BA3E9B237958DF" box="[447,603,447,473]" italics="true" pageId="6" pageNumber="67">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
populations from other localities of the Dniester River basin (
|
||
<figureCitation id="13302A63FFB3C40CF9B33EDB20185B1C" box="[182,314,511,538]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4 E–K</figureCitation>
|
||
). “
|
||
<taxonomicName id="4C0B4D65FFB3C40CF85F3EDB23A05B1F" box="[346,642,511,537]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB3C40CF85F3EDB23A05B1F" box="[346,642,511,537]" italics="true" pageId="6" pageNumber="67">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” in Put’s interpretation also shows a clear similarity in pattern to some
|
||
<taxonomicName id="4C0B4D65FFB3C40CF8023D6420B55B5C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[263,407,576,602]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB3C40CF8023D6420B55B5C" box="[263,407,576,602]" italics="true" pageId="6" pageNumber="67">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
morphotypes earlier described by
|
||
<bibRefCitation id="EF9A4B17FFB3C40CF9CA3D4420B15B7C" author="Lindholm W. A." box="[207,403,607,634]" pageId="6" pageNumber="67" pagination="199 - 232" refId="ref4837" refString="Lindholm W. A. 1908. Materialien zur Molluskenfauena [sic] von Sudwestrussland, Polen und der Krim. Zapiski Novorossijskago Obshchestva Estestvois- pytatelej, 31: 199 - 232." type="journal article" year="1908">Lindholm [1908]</bibRefCitation>
|
||
: about 10–12 stripes are found in
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9D53DA420585B9C" box="[208,378,640,666]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="obliteratus">
|
||
<emphasis id="B97FEAF4FFB3C40CF9D53DA420585B9C" box="[208,378,640,666]" italics="true" pageId="6" pageNumber="67">Th. obliteratus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype (
|
||
<figureCitation id="13302A63FFB3C40CFB1A3DA4235B5B9D" box="[543,633,640,667]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4L</figureCitation>
|
||
) and 5–7 stripes in
|
||
<taxonomicName id="4C0B4D65FFB3C40CF8243D8520C35BBD" box="[289,481,673,699]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="pulcherrimus">
|
||
<emphasis id="B97FEAF4FFB3C40CF8243D8520C35BBD" box="[289,481,673,699]" italics="true" pageId="6" pageNumber="67">Th. pulcherrimus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype (
|
||
<figureCitation id="13302A63FFB3C40CFB863D8523C25BBD" box="[643,736,673,699]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4O</figureCitation>
|
||
). A pattern of broken bands forming the spiral rows of large light spots is also commonly registered e.g. in
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9D53C2520555A1D" box="[208,375,769,795]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="lacrymans">
|
||
<emphasis id="B97FEAF4FFB3C40CF9D53C2520555A1D" box="[208,375,769,795]" italics="true" pageId="6" pageNumber="67">Th. lacrymans</emphasis>
|
||
</taxonomicName>
|
||
-morphotype (
|
||
<figureCitation id="13302A63FFB3C40CFB193C25235C5A1A" box="[540,638,769,796]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4M</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C0B4D65FFB3C40CFBCD3C2520185A3A" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="alboguttatus">
|
||
<emphasis id="B97FEAF4FFB3C40CFBCD3C2520185A3A" italics="true" pageId="6" pageNumber="67">Th. alboguttatus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype (
|
||
<figureCitation id="13302A63FFB3C40CF8DE3C0623155A3A" box="[475,567,802,828]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4N</figureCitation>
|
||
). It is important to note that the descriptions of
|
||
<taxonomicName id="4C0B4D65FFB3C40CFB2D3C6623C75A5A" box="[552,741,834,860]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="alboguttatus">
|
||
<emphasis id="B97FEAF4FFB3C40CFB2D3C6623C75A5A" box="[552,741,834,860]" italics="true" pageId="6" pageNumber="67">Th. alboguttatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9AB3C4620725A7A" box="[174,336,866,892]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="lacrymans">
|
||
<emphasis id="B97FEAF4FFB3C40CF9AB3C4620725A7A" box="[174,336,866,892]" italics="true" pageId="6" pageNumber="67">Th. lacrymans</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0B4D65FFB3C40CF8953C4623735A7A" box="[400,593,866,892]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="pulcherrimus">
|
||
<emphasis id="B97FEAF4FFB3C40CF8953C4623735A7A" box="[400,593,866,892]" italics="true" pageId="6" pageNumber="67">Th. pulcherrimus</emphasis>
|
||
</taxonomicName>
|
||
morphotypes [
|
||
<bibRefCitation id="EF9A4B17FFB3C40CF9B03CA720485A9B" author="Lindholm W. A." box="[181,362,898,925]" pageId="6" pageNumber="67" pagination="199 - 232" refId="ref4837" refString="Lindholm W. A. 1908. Materialien zur Molluskenfauena [sic] von Sudwestrussland, Polen und der Krim. Zapiski Novorossijskago Obshchestva Estestvois- pytatelej, 31: 199 - 232." type="journal article" year="1908">Lindholm, 1908</bibRefCitation>
|
||
] were based on specimens collected near Odessa (
|
||
<tableCitation id="C689035DFFB3C40CF84C3C8720825ABB" box="[329,416,931,957]" captionStart="Table 1" captionStartId="2.[174,227,189,211]" captionText="Table 1. Examined material of Theodoxus from the SW Ukraine. The numbers of localities refer to the text and figures. Coor- dinates of collecting sites sampled prior to 2009 are given approximately, based on data in museum catalogue cards, labels and corresponding publications." pageId="6" pageNumber="67">Table 1</tableCitation>
|
||
): in the
|
||
<collectionCode id="ED1AAE23FFB3C40CFB033C8723175ABB" box="[518,565,931,957]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="67" type="Museum">ZIN</collectionCode>
|
||
catalogue these samples are erroneously labelled as “
|
||
<emphasis id="B97FEAF4FFB3C40CFB6A3CE720225AFB" italics="true" pageId="6" pageNumber="67">Black Sea, Odessa</emphasis>
|
||
” but these molluscs do not inhabit the Black Sea and are evidently originated from the neighboring limans. Meanwhile, the only shell of Lindholm’s
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9A93B6020725D58" box="[172,336,1092,1118]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="obliteratus">
|
||
<emphasis id="B97FEAF4FFB3C40CF9A93B6020725D58" box="[172,336,1092,1118]" italics="true" pageId="6" pageNumber="67">Th. obliteratus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype, that makes up the type series of this taxon, was collected by A. Brauner in Dniester River near Rybnitsa,
|
||
<collectingCountry id="F31C7676FFB3C40CFB083BA123505D99" box="[525,626,1157,1183]" name="Moldova" pageId="6" pageNumber="67">Moldova</collectingCountry>
|
||
(
|
||
<tableCitation id="C689035DFFB3C40CFB813BA123F95D99" box="[644,731,1157,1183]" captionStart="Table 1" captionStartId="2.[174,227,189,211]" captionText="Table 1. Examined material of Theodoxus from the SW Ukraine. The numbers of localities refer to the text and figures. Coor- dinates of collecting sites sampled prior to 2009 are given approximately, based on data in museum catalogue cards, labels and corresponding publications." pageId="6" pageNumber="67">Table 1</tableCitation>
|
||
), i.e. not far from the type locality of “
|
||
<taxonomicName id="4C0B4D65FFB3C40CFB763B8120775DD9" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB3C40CFB763B8120775DD9" italics="true" pageId="6" pageNumber="67">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” at least from the same part of the river basin.
|
||
</paragraph>
|
||
<caption id="DF74666EFFB1C40EF9AB3DDD225C5A6F" ID-DOI="http://doi.org/10.5281/zenodo.11040992" ID-Zenodo-Dep="11040992" httpUri="https://zenodo.org/record/11040992/files/figure.png" pageId="4" pageNumber="65" startId="4.[174,216,761,782]" targetBox="[177,1382,192,720]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BB436E6FFB1C40EF9AB3DDD22155A2F" blockId="4.[174,1384,761,873]" pageId="4" pageNumber="65">
|
||
FIG. 2. Some lots of river nerites collected by A.L. Put’ on May 26, 1950 from the Dniester River near Rukhotyn village, the type locality of “
|
||
<taxonomicName id="4C0B4D65FFB1C40EF8733C3723515A2E" box="[374,627,787,808]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="65" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB1C40EF8733C3723515A2E" box="[374,627,787,808]" italics="true" pageId="4" pageNumber="65">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” (Fig. 1, locality 1).
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB1C40EF9AB3C1D225C5A6F" blockId="4.[174,1384,761,873]" pageId="4" pageNumber="65">
|
||
РИС. 2. Некоторые пробы лунок, собранные А.Л. Путем 26 мая 1950 года иЗ р. Днестр воЗле с. Рухотин, типового местонахоЖдения “
|
||
<taxonomicName id="4C0B4D65FFB1C40EF89F3C7723B55A6E" box="[410,663,851,872]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="65" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB1C40EF89F3C7723B55A6E" box="[410,663,851,872]" italics="true" pageId="4" pageNumber="65">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” (Рис. 1, локалитет 1).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF74666EFFB1C40EF9AB38A1206D5EF2" ID-DOI="http://doi.org/10.5281/zenodo.11040996" ID-Zenodo-Dep="11040996" httpUri="https://zenodo.org/record/11040996/files/figure.png" pageId="4" pageNumber="65" startId="4.[174,216,1925,1946]" targetBox="[174,1379,990,1851]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BB436E6FFB1C40EF9AB38A120F75EB3" blockId="4.[174,1382,1925,2036]" pageId="4" pageNumber="65">
|
||
FIG. 3. Selected ecotopes of the Dniester River near Monastyrok village, locality 6 (
|
||
<emphasis id="B97FEAF4FFB1C40EFAE138A122D75E9C" bold="true" box="[996,1013,1925,1946]" pageId="4" pageNumber="65">A</emphasis>
|
||
,
|
||
<emphasis id="B97FEAF4FFB1C40EFD0738A125305E9C" bold="true" box="[1026,1042,1925,1946]" pageId="4" pageNumber="65">B</emphasis>
|
||
) and Halych village, locality 5 (
|
||
<emphasis id="B97FEAF4FFB1C40EFC4B38A1247D5E9D" bold="true" box="[1358,1375,1925,1947]" pageId="4" pageNumber="65">C</emphasis>
|
||
,
|
||
<emphasis id="B97FEAF4FFB1C40EF9D138BB21C75EB2" bold="true" box="[212,229,1951,1972]" pageId="4" pageNumber="65">D</emphasis>
|
||
). Photo E. Degtyarenko.
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB1C40EF9AB38E1206D5EF2" blockId="4.[174,1382,1925,2036]" pageId="4" pageNumber="65">
|
||
РИС. 3. Некоторые Экотопы р. Днестр воЗле с. Монастырок, локалитет 6 (
|
||
<emphasis id="B97FEAF4FFB1C40EFA9A38E122925EDC" bold="true" box="[927,944,1989,2010]" pageId="4" pageNumber="65">A</emphasis>
|
||
,
|
||
<emphasis id="B97FEAF4FFB1C40EFAB938E122EE5EDC" bold="true" box="[956,972,1989,2010]" pageId="4" pageNumber="65">B</emphasis>
|
||
) и с. Галич, локалитет 5 (
|
||
<emphasis id="B97FEAF4FFB1C40EFDD138E125C75EDD" bold="true" box="[1236,1253,1989,2011]" pageId="4" pageNumber="65">C</emphasis>
|
||
,
|
||
<emphasis id="B97FEAF4FFB1C40EFDF438E124205EDC" bold="true" box="[1265,1282,1989,2010]" pageId="4" pageNumber="65">D</emphasis>
|
||
). Фото Е. Дегтяренко.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF74666EFFB0C40FF9A3390524395E8B" ID-DOI="http://doi.org/10.5281/zenodo.11041000" ID-Zenodo-Dep="11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="5" pageNumber="66" startId="5.[166,208,1569,1590]" targetBox="[166,1354,212,1500]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BB436E6FFB0C40FF9A33905247E5FC8" blockId="5.[166,1376,1569,1933]" pageId="5" pageNumber="66">
|
||
FIG. 4. Shells of
|
||
<taxonomicName id="4C0B4D65FFB0C40FF840390523945F31" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[325,694,1569,1591]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB0C40FF840390523285F30" box="[325,522,1569,1590]" italics="true" pageId="5" pageNumber="66">Theodoxus fluviatilis</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B97FEAF4FFB0C40FFBB8390523CE5F31" bold="true" box="[701,748,1569,1591]" pageId="5" pageNumber="66">A–C</emphasis>
|
||
. Three topotypes of “
|
||
<taxonomicName id="4C0B4D65FFB0C40FFABC390521CE5F56" authority="", NMNH NASU, PD" authorityName="NMNH NASU, PD" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB0C40FFABC390525965F30" box="[953,1204,1569,1590]" italics="true" pageId="5" pageNumber="66">Theodoxus dniestroviensis</emphasis>
|
||
”, NMNH NASU, PD
|
||
</taxonomicName>
|
||
, no number, locality 1.
|
||
<emphasis id="B97FEAF4FFB0C40FF8C0391E20F45F49" bold="true" box="[453,470,1594,1615]" pageId="5" pageNumber="66">D</emphasis>
|
||
. Lectotype of
|
||
<taxonomicName id="4C0B4D65FFB0C40FFB58391E222D5F49" box="[605,783,1594,1615]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB0C40FFB58391E222D5F49" box="[605,783,1594,1615]" italics="true" pageId="5" pageNumber="66">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
, designated here, locality 1, photo reproduced after Put’, 1972.
|
||
<emphasis id="B97FEAF4FFB0C40FF9CF397021F85F6F" bold="true" box="[202,218,1620,1641]" pageId="5" pageNumber="66">E</emphasis>
|
||
. Specimen from Seret River, IZAN #614, locality 2.
|
||
<emphasis id="B97FEAF4FFB0C40FFBDA397023CF5F6F" bold="true" box="[735,749,1620,1641]" pageId="5" pageNumber="66">F</emphasis>
|
||
. Specimen from Hnizna River, IZAN #616, locality 4.
|
||
<emphasis id="B97FEAF4FFB0C40FFC06397724375F6F" bold="true" box="[1283,1301,1619,1641]" pageId="5" pageNumber="66">G</emphasis>
|
||
. Specimen from Strypa River, IZAN #615, locality 3.
|
||
<emphasis id="B97FEAF4FFB0C40FFB91394923855F84" bold="true" box="[660,679,1645,1666]" pageId="5" pageNumber="66">H</emphasis>
|
||
,
|
||
<emphasis id="B97FEAF4FFB0C40FFBB63949239E5F84" bold="true" box="[691,700,1645,1666]" pageId="5" pageNumber="66">I</emphasis>
|
||
. Specimens from Dniester River, IZAN #333, locality 6.
|
||
<emphasis id="B97FEAF4FFB0C40FFDE7394925D75F84" bold="true" box="[1250,1269,1645,1666]" pageId="5" pageNumber="66">K</emphasis>
|
||
. Specimen from Dniester River, IZAN #335, locality 5.
|
||
<emphasis id="B97FEAF4FFB0C40FFB7E39A223A95F9D" bold="true" box="[635,651,1670,1691]" pageId="5" pageNumber="66">L</emphasis>
|
||
.
|
||
<taxonomicName id="4C0B4D65FFB0C40FFB9239A222075F9D" box="[663,805,1670,1691]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="obliteratus">
|
||
<emphasis id="B97FEAF4FFB0C40FFB9239A222075F9D" box="[663,805,1670,1691]" italics="true" pageId="5" pageNumber="66">Th. obliteratus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype, ZIN #5912/1, locality 7.
|
||
<emphasis id="B97FEAF4FFB0C40FFD9839A225915F9D" bold="true" box="[1181,1203,1670,1691]" pageId="5" pageNumber="66">M</emphasis>
|
||
. Lectotype of
|
||
<taxonomicName id="4C0B4D65FFB0C40FFC4539A220125FB2" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="lacrymans">
|
||
<emphasis id="B97FEAF4FFB0C40FFC4539A220125FB2" italics="true" pageId="5" pageNumber="66">Th. lacrymans</emphasis>
|
||
</taxonomicName>
|
||
-morphotype, ZIN #6052/1, locality 8.
|
||
<emphasis id="B97FEAF4FFB0C40FFB9A398423925FB3" bold="true" box="[671,688,1696,1717]" pageId="5" pageNumber="66">N</emphasis>
|
||
. Lectotype of
|
||
<taxonomicName id="4C0B4D65FFB0C40FFA3C39BB22F75FB2" box="[825,981,1695,1716]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="alboguttatus">
|
||
<emphasis id="B97FEAF4FFB0C40FFA3C39BB22F75FB2" box="[825,981,1695,1716]" italics="true" pageId="5" pageNumber="66">Th. alboguttatus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype, ZIN #6051/1, locality 8.
|
||
<emphasis id="B97FEAF4FFB0C40FFC4039BB247A5FB3" bold="true" box="[1349,1368,1695,1717]" pageId="5" pageNumber="66">O</emphasis>
|
||
. Lectotype of
|
||
<taxonomicName id="4C0B4D65FFB0C40FF849399D20D35FC8" box="[332,497,1721,1742]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="pulcherrimus">
|
||
<emphasis id="B97FEAF4FFB0C40FF849399D20D35FC8" box="[332,497,1721,1742]" italics="true" pageId="5" pageNumber="66">Th. pulcherrimus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype, ZIN #6053/1, locality 8.
|
||
<emphasis id="B97FEAF4FFB0C40FFA6D399D22B55FC9" bold="true" box="[872,919,1721,1743]" pageId="5" pageNumber="66">L–O</emphasis>
|
||
. Reproduced after V.
|
||
<bibRefCitation id="EF9A4B17FFB0C40FFD62399D247A5FC8" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." box="[1127,1368,1721,1743]" pageId="5" pageNumber="66" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB0C40FFDEC399D24315FC8" box="[1257,1299,1721,1742]" italics="true" pageId="5" pageNumber="66">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB436E6FFB0C40FF9A339C424395E8B" blockId="5.[166,1376,1569,1933]" pageId="5" pageNumber="66">
|
||
РИС. 4.
|
||
<taxonomicName id="4C0B4D65FFB0C40FF9F339FB23465FF3" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[246,612,1759,1781]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB0C40FF9F339FB20995FF2" box="[246,443,1759,1780]" italics="true" pageId="5" pageNumber="66">Theodoxus fluviatilis</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B97FEAF4FFB0C40FFB6F39FB23B85FF3" bold="true" box="[618,666,1759,1781]" pageId="5" pageNumber="66">A–C</emphasis>
|
||
. Топотипы “
|
||
<taxonomicName id="4C0B4D65FFB0C40FFA1C39FB25C65FF3" authority="", NMNH NASU, PD" authorityName="NMNH NASU, PD" box="[793,1252,1759,1781]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB0C40FFA1C39FB25375FF2" box="[793,1045,1759,1780]" italics="true" pageId="5" pageNumber="66">Theodoxus dniestroviensis</emphasis>
|
||
”, NMNH NASU, PD
|
||
</taxonomicName>
|
||
, беЗ номера, локалитет 1.
|
||
<emphasis id="B97FEAF4FFB0C40FF85639DD20465E08" bold="true" box="[339,356,1785,1806]" pageId="5" pageNumber="66">D</emphasis>
|
||
. Лектотип
|
||
<taxonomicName id="4C0B4D65FFB0C40FF8DC39DD23AC5E08" box="[473,654,1785,1806]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB0C40FF8DC39DD23AC5E08" box="[473,654,1785,1806]" italics="true" pageId="5" pageNumber="66">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972.
|
||
<emphasis id="B97FEAF4FFB0C40FFC4C39DD247B5E08" bold="true" box="[1353,1369,1785,1806]" pageId="5" pageNumber="66">E</emphasis>
|
||
. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2.
|
||
<emphasis id="B97FEAF4FFB0C40FFBBE383623E85E21" bold="true" box="[699,714,1810,1831]" pageId="5" pageNumber="66">F</emphasis>
|
||
. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4.
|
||
<emphasis id="B97FEAF4FFB0C40FFDD7383625C75E2E" bold="true" box="[1234,1253,1810,1832]" pageId="5" pageNumber="66">G</emphasis>
|
||
. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3.
|
||
<emphasis id="B97FEAF4FFB0C40FFB5E3808234C5E47" bold="true" box="[603,622,1836,1857]" pageId="5" pageNumber="66">H</emphasis>
|
||
,
|
||
<emphasis id="B97FEAF4FFB0C40FFB7C380823A05E47" bold="true" box="[633,642,1836,1857]" pageId="5" pageNumber="66">I</emphasis>
|
||
. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6.
|
||
<emphasis id="B97FEAF4FFB0C40FFD9B380825935E47" bold="true" box="[1182,1201,1836,1857]" pageId="5" pageNumber="66">K</emphasis>
|
||
. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5.
|
||
<emphasis id="B97FEAF4FFB0C40FFB1E386123095E5C" bold="true" box="[539,555,1861,1882]" pageId="5" pageNumber="66">L</emphasis>
|
||
. Морфотип
|
||
<taxonomicName id="4C0B4D65FFB0C40FFBA3386122475E5C" authority=", ZIN" authorityName="ZIN" box="[678,869,1861,1882]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="obliteratus">
|
||
<emphasis id="B97FEAF4FFB0C40FFBA3386122105E5C" box="[678,818,1861,1882]" italics="true" pageId="5" pageNumber="66">Th. obliteratus</emphasis>
|
||
, ZIN
|
||
</taxonomicName>
|
||
#5912/1, локалитет 7.
|
||
<emphasis id="B97FEAF4FFB0C40FFD41386125795E5C" bold="true" box="[1092,1115,1861,1882]" pageId="5" pageNumber="66">M</emphasis>
|
||
. Лектотип морфотипа
|
||
<taxonomicName id="4C0B4D65FFB0C40FFC45386120405E72" authority=", ZIN" authorityName="ZIN" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="lacrymans">
|
||
<emphasis id="B97FEAF4FFB0C40FFC45386120125E75" italics="true" pageId="5" pageNumber="66">Th. lacrymans</emphasis>
|
||
, ZIN
|
||
</taxonomicName>
|
||
#6052/1, локалитет 8.
|
||
<emphasis id="B97FEAF4FFB0C40FFB44387A23705E75" bold="true" box="[577,594,1886,1907]" pageId="5" pageNumber="66">N</emphasis>
|
||
. Лектотип морфотипа
|
||
<taxonomicName id="4C0B4D65FFB0C40FFA33387A25265E72" authority=", ZIN" authorityName="ZIN" box="[822,1028,1886,1908]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="alboguttatus">
|
||
<emphasis id="B97FEAF4FFB0C40FFA33387A22F05E75" box="[822,978,1886,1907]" italics="true" pageId="5" pageNumber="66">Th. alboguttatus</emphasis>
|
||
, ZIN
|
||
</taxonomicName>
|
||
#6051/1, локалитет 8.
|
||
<emphasis id="B97FEAF4FFB0C40FFDE7387A25D75E72" bold="true" box="[1250,1269,1886,1908]" pageId="5" pageNumber="66">O</emphasis>
|
||
. Лектотип морфотипа
|
||
<taxonomicName id="4C0B4D65FFB0C40FF8463853233F5E8B" authority=", ZIN" authorityName="ZIN" box="[323,541,1911,1933]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="66" phylum="Mollusca" rank="species" species="pulcherrimus">
|
||
<emphasis id="B97FEAF4FFB0C40FF846385320CB5E8A" box="[323,489,1911,1932]" italics="true" pageId="5" pageNumber="66">Th. pulcherrimus</emphasis>
|
||
, ZIN
|
||
</taxonomicName>
|
||
#6053/1, локалитет 8.
|
||
<emphasis id="B97FEAF4FFB0C40FFA07385C22135E8B" bold="true" box="[770,817,1911,1933]" pageId="5" pageNumber="66">L–O</emphasis>
|
||
. ВоспроиЗведены по V.
|
||
<bibRefCitation id="EF9A4B17FFB0C40FFD23385324355E8B" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." box="[1062,1303,1911,1933]" pageId="5" pageNumber="66" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB0C40FFDAD385C25F05E8A" box="[1192,1234,1911,1933]" italics="true" pageId="5" pageNumber="66">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB436E6FFB3C40CF9D13A2220AC5E60" blockId="6.[172,750,188,1894]" pageId="6" pageNumber="67">
|
||
Based on conchological features as well as periostracum colouration and patterning it is hardly possible to distinguish
|
||
<taxonomicName id="4C0B4D65FFB3C40CF8CC3A6323855C67" box="[457,679,1351,1377]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB3C40CF8CC3A6323855C67" box="[457,679,1351,1377]" italics="true" pageId="6" pageNumber="67">Th. dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
from morphotypes of
|
||
<taxonomicName id="4C0B4D65FFB3C40CF86E3A4320DC5C87" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[363,510,1383,1409]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB3C40CF86E3A4320DC5C87" box="[363,510,1383,1409]" italics="true" pageId="6" pageNumber="67">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
given the variability in colour and patterns among the topotypic material (
|
||
<figureCitation id="13302A63FFB3C40CF9B33A8320115CC4" box="[182,307,1447,1474]" captionStart="FIG" captionStartId="5.[166,208,1569,1590]" captionTargetBox="[166,1354,212,1500]" captionTargetId="figure-13@5.[166,1354,212,1500]" captionTargetPageId="5" captionText="FIG. 4. Shells ofTheodoxus fluviatilis (Linnaeus, 1758).A–C. Three topotypes of “Theodoxus dniestroviensis”, NMNH NASU, PD, no number, locality 1. D. Lectotype ofTh. dniestroviensis, designated here, locality 1, photo reproduced after Put’, 1972. E. Specimen from Seret River, IZAN #614, locality 2. F. Specimen from Hnizna River, IZAN #616, locality 4. G. Specimen from Strypa River, IZAN #615, locality 3. H, I. Specimens from Dniester River, IZAN #333, locality 6. K. Specimen from Dniester River, IZAN #335, locality 5. L. Th. obliteratus-morphotype, ZIN #5912/1, locality 7. M. Lectotype of Th. lacrymans-morphotype, ZIN #6052/1, locality 8.N. Lectotype of Th. alboguttatus-morphotype, ZIN #6051/1, locality 8.O. Lectotype of Th. pulcherrimus-morphotype, ZIN #6053/1, locality 8. L–O. Reproduced after V. Anistratenko et al., 2020. РИС.4. Theodoxus fluviatilis (Linnaeus, 1758).A–C. Топотипы “Theodoxus dniestroviensis”, NMNH NASU, PD, беЗ номера, локалитет 1. D. Лектотип Th. dniestroviensis, обоЗначен Здесь, локалитет 1, фото воспроиЗведено по Put’, 1972. E. ЭкЗемпляр иЗ р. Серет, IZAN #614, локалитет 2. F. ЭкЗемпляр иЗ р. ГниЗна, IZAN #616, локалитет 4. G. ЭкЗемпляр иЗ р. Стрыпа, IZAN #615, локалитет 3. H, I. ЭкЗемпляры иЗ р. Днестр, IZAN #333, локалитет 6. K. ЭкЗемпляр иЗ р. Днестр, IZAN #335, локалитет 5. L. Морфотип Th. obliteratus, ZIN #5912/1, локалитет 7. M. Лектотип морфотипа Th. lacrymans, ZIN #6052/1, локалитет 8. N. Лектотип морфотипа Th. alboguttatus, ZIN #6051/1, локалитет 8. O. Лектотип морфотипа Th. pulcherrimus, ZIN #6053/1, локалитет 8. L–O. ВоспроиЗведены по V. Anistratenko et al., 2020." figureDoi="http://doi.org/10.5281/zenodo.11041000" httpUri="https://zenodo.org/record/11041000/files/figure.png" pageId="6" pageNumber="67">Fig. 4 A–C</figureCitation>
|
||
). Since the original description contains no other differential characteristics apart the pattern, “
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9BF3ACC20C65F04" box="[186,484,1512,1538]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="dniestroviensis">
|
||
<emphasis id="B97FEAF4FFB3C40CF9BF3ACC20C65F04" box="[186,484,1512,1538]" italics="true" pageId="6" pageNumber="67">Theodoxus dniestroviensis</emphasis>
|
||
</taxonomicName>
|
||
” should be interpreted as a local morphotype of the polymorphic species
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9AB390D20605F45" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[174,322,1577,1603]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="B97FEAF4FFB3C40CF9AB390D20605F45" box="[174,322,1577,1603]" italics="true" pageId="6" pageNumber="67">Th. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
(L., 1758) as it was suggested earlier [O.
|
||
<bibRefCitation id="EF9A4B17FFB3C40CF9D8396D23245F65" author="Anistratenko O. Yu. & Starobogatov Ya. I. & Anistratenko V. V." box="[221,518,1608,1635]" pageId="6" pageNumber="67" pagination="11 - 19" refId="ref4413" refString="Anistratenko O. Yu., Starobogatov Ya. I., Anistratenko V. V. 1999. Mollusks of the genus Theodoxus (Gastropoda, Pectinibranchia, Neritidae) of the Black and the Azov seas basin. Vestnik zoologii, 33: 11 - 19 [In Russian with English summary]." type="journal article" year="1999">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB3C40CF87C396E208D5F65" box="[377,431,1609,1635]" italics="true" pageId="6" pageNumber="67">et al</emphasis>
|
||
., 1999
|
||
</bibRefCitation>
|
||
]. The present study contributes additional data to the recently published overview of the
|
||
<typeStatus id="54B08844FFB3C40CF86F39AE20B85FA2" box="[362,410,1674,1700]" pageId="6" pageNumber="67">type</typeStatus>
|
||
series of nominal taxa of the genus
|
||
<taxonomicName id="4C0B4D65FFB3C40CF9F0398E204C5FC2" authorityName="Montfort" authorityYear="1810" box="[245,366,1706,1732]" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="67" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97FEAF4FFB3C40CF9F0398E204C5FC2" box="[245,366,1706,1732]" italics="true" pageId="6" pageNumber="67">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
inhabiting the Ponto-Caspian area [
|
||
<collectionCode id="ED1AAE23FFB3C40CF9B639EF21EB5FE3" box="[179,201,1739,1765]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="67" type="Museum">V</collectionCode>
|
||
.
|
||
<bibRefCitation id="EF9A4B17FFB3C40CF9D139EE20CF5FE2" author="Anistratenko V. V. & Sitnikova T. Ya. & Kijashko P. V. & Vinarski M. V. & Anistratenko O. Yu." box="[212,493,1737,1765]" pageId="6" pageNumber="67" pagination="115 - 134" refId="ref4562" refString="Anistratenko V. V., Sitnikova T. Ya., Kijashko P. V., Vinarski M. V., Anistratenko O. Yu. 2020. A review of species of the genus Theodoxus (Gastropoda: Neritidae) of the Ponto-Caspian region, with considerations on available type materials. Ruthenica, Russian Malacological Journal, 30 (2): 115 - 134. https: // doi. org / 10.35885 / ruthenica. 2021.30 (2). 5" type="journal article" year="2020">
|
||
Anistratenko
|
||
<emphasis id="B97FEAF4FFB3C40CF86E39EF20B95FE2" box="[363,411,1738,1764]" italics="true" pageId="6" pageNumber="67">et al</emphasis>
|
||
., 2020
|
||
</bibRefCitation>
|
||
]. It adds knowledge on the river nerites’ variability range as well as sheds more light on understanding how many valid species of this genus exist in a region and what the proper names for them are.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |