159 lines
17 KiB
XML
159 lines
17 KiB
XML
<document id="BDF33AB070C3DC7406A8C62611C7241D" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="11755326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7828FFB1FEECFE9E9D86FEA1" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra monteverde Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="62" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="62" updateTime="1698190273211" updateUser="plazi">
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<mods:mods id="C01C55C5908AB2E7A087F11F3313CC9E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="775AC6321BCF6A9234FBE2F1E61BC69C">
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<mods:title id="D9E2EC778796FFD2F91B253295572486">The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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</mods:titleInfo>
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<mods:name id="57FC516DE001517A9DEE129F9B5BCCD4" type="personal">
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<mods:role id="DCEF8DD7D8314EC2B2E14CE8D15A2FDF">
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<mods:roleTerm id="A663749C7A3F40D74CDC98E25CF24D7B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6BD7A23F5B6711E12B319AD879F9E85E">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="2903946AD3DA0DB4C0D7B0F40C61C381">text</mods:typeOfResource>
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<mods:titleInfo id="4B3D5149232A11E7B0CBCA4FC6D020F7">
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<mods:title id="1DDFD9096E9616504202DF7095847DCC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BFB0CB98BBA977DF4E26A2651D45B09C">
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<mods:date id="8871BCC00692C24126BE92CA8D2463C6">2002</mods:date>
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<mods:detail id="9CDE758B528053CED32284631496B715" type="volume">
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<mods:number id="D652B837AB883248B646877F0FE9D7A0">119</mods:number>
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<mods:start id="80097C90F57510C412D33EE8F90DB699">1</mods:start>
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<mods:end id="0B2B8E0E6AA1CC39B777680099C0E921">68</mods:end>
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<mods:classification id="94A7354CE727F57CB688865856736451">journal article</mods:classification>
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<mods:identifier id="F33DCFF829D994399DAD8A8CC39F3920" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<mods:identifier id="16EA373FE5C7B58D049E96CA94F29091" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="D38E7C65EB85895BF6D392760C3358A9" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="3F4895C4D344CD41B8A0BB118ACD5146" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA7828FFB1FEECFE9E9D86FEA1" ID-DOI="http://doi.org/10.5281/zenodo.6278034" ID-GBIF-Taxon="121065206" ID-Zenodo-Dep="6278034" LSID="urn:lsid:plazi:treatment:03A287CA7828FFB1FEECFE9E9D86FEA1" httpUri="http://treatment.plazi.org/id/03A287CA7828FFB1FEECFE9E9D86FEA1" lastPageId="62" lastPageNumber="62" pageId="61" pageNumber="62">
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<subSubSection id="C31165577828FFB2FEECFE9E9E01FEA1" pageId="61" pageNumber="62" type="nomenclature">
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<paragraph id="8BB436DC7828FFB2FEECFE9E9EF8FECA" blockId="61.[264,716,283,350]" box="[264,716,283,310]" pageId="61" pageNumber="62">
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<heading id="D0FC81B07828FFB2FEECFE9E9EF8FECA" bold="true" box="[264,716,283,310]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
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<emphasis id="B97FEACE7828FFB2FEECFE9E9EF8FECA" bold="true" box="[264,716,283,310]" pageId="61" pageNumber="62">
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<taxonomicName id="4C0B4D5F7828FFB2FEECFE9E9E1BFECA" ID-CoL="B2RS" authority="Erwin" authorityName="Erwin" box="[264,559,283,310]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="monteverde" status="sp. nov.">
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<emphasis id="B97FEACE7828FFB2FEECFE9E9DECFEC9" bold="true" box="[264,472,284,310]" italics="true" pageId="61" pageNumber="62">Agra monteverde</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C57B57828FFB2FDD8FE999EF8FECA" box="[572,716,283,309]" pageId="61" pageNumber="62" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB436DC7828FFB2FEECFEC69E01FEA1" blockId="61.[264,716,283,350]" box="[264,565,324,350]" pageId="61" pageNumber="62">
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(
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<figureCitation id="13302A597828FFB2FEF5FEC69D9FFEA1" box="[273,427,324,350]" captionStart="FIGURE 29. A" captionStartId="61.[264,368,1565,1589]" captionTargetBox="[535,1055,926,1527]" captionTargetId="figure@61.[526,1060,913,1539]" captionTargetPageId="61" captionText="FIGURE 29. A. monteverde, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, ventral aspect; d) stylomere 2, ventral aspect. Measures (mm): HL = 2.8; WE = 1.9; SW = 3.6; S 2 L = 0.4." httpUri="https://zenodo.org/record/162843/files/figure.png" pageId="61" pageNumber="62">Figs. 29a, 29</figureCitation>
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b, 29c, 29d)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577828FFB2FEECFE119F26FE29" pageId="61" pageNumber="62" type="materials_examined">
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<paragraph id="8BB436DC7828FFB2FEECFE119F26FE29" blockId="61.[264,1324,403,870]" pageId="61" pageNumber="62">
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<bibRef id="52ABA22A7828FFB2FEECFE119F26FE29" pageId="61" pageNumber="62" type="journal article">
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<emphasis id="B97FEACE7828FFB2FEECFE119DB5FE52" bold="true" box="[264,385,403,429]" pageId="61" pageNumber="62">
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<title id="F90FDDEA7828FFB2FEECFE119D48FE52" box="[264,380,403,429]" jopScore="0.0" pageId="61" pageNumber="62" type="_" typeScore="3">
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<typeStatus id="54B0887E7828FFB2FEECFE119D48FE52" box="[264,380,403,429]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
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</title>
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.
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</emphasis>
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<author id="53B6DE397828FFB2FE6AFE169E99FE51" box="[398,685,404,430]" fns="I" npo="LF" pageId="61" pageNumber="62">
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Female,
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<collectingCountry id="F31C764C7828FFB2FE1CFE169E99FE51" box="[504,685,404,430]" name="Costa Rica" pageId="61" pageNumber="62">COSTA RICA</collectingCountry>
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</author>
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:
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<journalOrPublisher id="51CA3C107828FFB2FD59FE169FD6FE51" box="[701,994,404,430]" jopScore="0.0" pageId="61" pageNumber="62">Puntarenas, Monteverde</journalOrPublisher>
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,
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<quantity id="4CF39B397828FFB2FC16FE169872FE51" box="[1010,1094,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="61" pageNumber="62" unit="m" value="1380.0">
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<part id="FFAF9C417828FFB2FC16FE16981AFE51" box="[1010,1070,404,430]" pageId="61" pageNumber="62" type="volume">1380</part>
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m
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</quantity>
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,
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<geoCoordinate id="EE3F501B7828FFB2FBB3FE169882FE51" box="[1111,1206,404,430]" direction="north" orientation="latitude" pageId="61" pageNumber="62" precision="925" value="10.833333">
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<pagination id="B2C2C8287828FFB2FBB3FE169847FE51" box="[1111,1139,404,430]" pageId="61" pageNumber="62" type="pagination">10</pagination>
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º50'N
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</geoCoordinate>
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<geoCoordinate id="EE3F501B7828FFB2FB26FE169912FE51" box="[1218,1318,404,430]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="925" value="-85.61667">85º37'W</geoCoordinate>
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, January, (F.T. Hovore)(
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<collectionCode id="ED1AAE197828FFB2FDFDFE3E9E46FE29" LSID="urn:lsid:biocol.org:col:34871" box="[537,626,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="61" pageNumber="62">USNM</collectionCode>
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: ADP93375).
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</bibRef>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577828FFB2FEDCFE619DEEFDB1" pageId="61" pageNumber="62" type="diagnosis">
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<paragraph id="8BB436DC7828FFB2FEDCFE619DEEFDB1" blockId="61.[264,1324,403,870]" pageId="61" pageNumber="62">
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<emphasis id="B97FEACE7828FFB2FEDCFE619D83FE02" bold="true" box="[312,439,483,509]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
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This is the only species of Costa Rican
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<taxonomicName id="4C0B4D5F7828FFB2FC7BFE669FEEFE02" box="[927,986,484,509]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FEACE7828FFB2FC7BFE669FEEFE02" box="[927,986,484,509]" italics="true" pageId="61" pageNumber="62">Agra</emphasis>
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</taxonomicName>
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with aeneous elytral foveae and slightly oblique apex without a sutural tooth AND foveae separated by their own diameter or more.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577828FFB2FEDCFDD99DFEFC99" pageId="61" pageNumber="62" type="description">
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<paragraph id="8BB436DC7828FFB2FEDCFDD99DFEFC99" blockId="61.[264,1324,403,870]" pageId="61" pageNumber="62">
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<emphasis id="B97FEACE7828FFB2FEDCFDD99DE4FD8A" bold="true" box="[312,464,603,629]" pageId="61" pageNumber="62">Description.</emphasis>
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Color and luster: Castaneous. Legs paler than body and with infuscated knees; antennomeres infuscated apically. Form: Male unknown. Body and legs castaneous, legs paler than body and with infuscated knees. Head (
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<figureCitation id="13302A597828FFB2FC60FD2E9FD2FD39" box="[900,998,684,710]" captionStart="FIGURE 29. A" captionStartId="61.[264,368,1565,1589]" captionTargetBox="[535,1055,926,1527]" captionTargetId="figure@61.[526,1060,913,1539]" captionTargetPageId="61" captionText="FIGURE 29. A. monteverde, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, ventral aspect; d) stylomere 2, ventral aspect. Measures (mm): HL = 2.8; WE = 1.9; SW = 3.6; S 2 L = 0.4." httpUri="https://zenodo.org/record/162843/files/figure.png" pageId="61" pageNumber="62">Fig. 29a</figureCitation>
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) behind eyes elongate, posteriorly rounded. Female slender, with elytra slightly flared behind middle, apex (
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<figureCitation id="13302A597828FFB2FB5BFD569927FD11" box="[1215,1299,724,750]" captionStart="FIGURE 29. A" captionStartId="61.[264,368,1565,1589]" captionTargetBox="[535,1055,926,1527]" captionTargetId="figure@61.[526,1060,913,1539]" captionTargetPageId="61" captionText="FIGURE 29. A. monteverde, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, ventral aspect; d) stylomere 2, ventral aspect. Measures (mm): HL = 2.8; WE = 1.9; SW = 3.6; S 2 L = 0.4." httpUri="https://zenodo.org/record/162843/files/figure.png" pageId="61" pageNumber="62">Fig. 29</figureCitation>
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b) slightly oblique, lateral tooth very small, sutural apex rounded. Sternum VI (
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<figureCitation id="13302A597828FFB2FB62FD7E98E8FCE9" box="[1158,1244,764,790]" captionStart="FIGURE 29. A" captionStartId="61.[264,368,1565,1589]" captionTargetBox="[535,1055,926,1527]" captionTargetId="figure@61.[526,1060,913,1539]" captionTargetPageId="61" captionText="FIGURE 29. A. monteverde, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, ventral aspect; d) stylomere 2, ventral aspect. Measures (mm): HL = 2.8; WE = 1.9; SW = 3.6; S 2 L = 0.4." httpUri="https://zenodo.org/record/162843/files/figure.png" pageId="61" pageNumber="62">Fig. 29</figureCitation>
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c) with shallow emargination in female. Stylomere 2 (
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<figureCitation id="13302A597828FFB2FCC3FCA69F4AFCC1" box="[807,894,804,830]" captionStart="FIGURE 29. A" captionStartId="61.[264,368,1565,1589]" captionTargetBox="[535,1055,926,1527]" captionTargetId="figure@61.[526,1060,913,1539]" captionTargetPageId="61" captionText="FIGURE 29. A. monteverde, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, ventral aspect; d) stylomere 2, ventral aspect. Measures (mm): HL = 2.8; WE = 1.9; SW = 3.6; S 2 L = 0.4." httpUri="https://zenodo.org/record/162843/files/figure.png" pageId="61" pageNumber="62">Fig. 29</figureCitation>
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d). Size: medium,
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<quantity id="4CF39B397828FFB2FBB7FCA69881FCC1" box="[1107,1205,804,830]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="61" pageNumber="62" unit="mm" value="15.1">15.1mm</quantity>
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in length,
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<quantity id="4CF39B397828FFB2FEECFCCE9D6EFC99" box="[264,346,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="61" pageNumber="62" unit="mm" value="3.8">3.8mm</quantity>
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in width.
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</paragraph>
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</subSubSection>
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<caption id="DF7466547828FFB2FEECF99F9D04F985" httpUri="https://zenodo.org/record/162843/files/figure.png" pageId="61" pageNumber="62" targetBox="[535,1055,926,1527]" targetPageId="61">
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<paragraph id="8BB436DC7828FFB2FEECF99F9D04F985" blockId="61.[264,1324,1565,1658]" pageId="61" pageNumber="62">
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<emphasis id="B97FEACE7828FFB2FEECF99F9DA3F9CA" bold="true" box="[264,407,1565,1589]" pageId="61" pageNumber="62">FIGURE 29.</emphasis>
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<taxonomicName id="4C0B4D5F7828FFB2FE7AF99C9E03F9CA" box="[414,567,1566,1589]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="monteverde">
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<emphasis id="B97FEACE7828FFB2FE7AF99C9E03F9CA" box="[414,567,1566,1589]" italics="true" pageId="61" pageNumber="62">A. monteverde</emphasis>
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</taxonomicName>
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, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, ventral aspect; d) stylomere 2, ventral aspect. Measures (mm): HL = 2.8; WE = 1.9; SW = 3.6; S2L = 0.4.
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</paragraph>
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</caption>
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<subSubSection id="C31165577828FFB1FEDCF92F9D86FEA1" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="materials_examined">
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||
<paragraph id="8BB436DC7828FFB2FEDCF92F9F6AF8E7" blockId="61.[264,1323,1709,1896]" pageId="61" pageNumber="62">
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<emphasis id="B97FEACE7828FFB2FEDCF92F9EA4F938" bold="true" box="[312,656,1709,1735]" pageId="61" pageNumber="62">Other specimens examined.</emphasis>
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The female
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<typeStatus id="54B0887E7828FFB2FCC0F92C9FA5F937" box="[804,913,1710,1736]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
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is the only specimen I saw.
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<emphasis id="B97FEACE7828FFB2FEDCF9579DCAF910" bold="true" box="[312,510,1749,1775]" pageId="61" pageNumber="62">Specific epithet.</emphasis>
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The specific epithet,
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<taxonomicName id="4C0B4D5F7828FFB2FD1DF9549FB4F910" box="[761,896,1750,1775]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="monteverde">
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<emphasis id="B97FEACE7828FFB2FD1DF9549FB4F910" box="[761,896,1750,1775]" italics="true" pageId="61" pageNumber="62">monteverde</emphasis>
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</taxonomicName>
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, is used as a noun in apposition, and refers to the
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<typeStatus id="54B0887E7828FFB2FE7FF97C9DF9F8E7" box="[411,461,1790,1816]" pageId="61" pageNumber="62">type</typeStatus>
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locality,
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<taxonomicName id="4C0B4D5F7828FFB2FDDEF97C9EFFF8E7" box="[570,715,1790,1816]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="monteverde">Monteverde</taxonomicName>
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,
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<collectingCountry id="F31C764C7828FFB2FD32F97C9F6DF8E7" box="[726,857,1790,1816]" name="Costa Rica" pageId="61" pageNumber="62">Costa Rica</collectingCountry>
|
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.
|
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</paragraph>
|
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<paragraph id="8BB436DC7828FFB1FEDCF8A79D86FEA1" blockId="61.[264,1323,1709,1896]" lastBlockId="62.[264,1324,284,350]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
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||
<emphasis id="B97FEACE7828FFB2FEDCF8A79DB1F8C0" bold="true" box="[312,389,1829,1855]" pageId="61" pageNumber="62">Notes.</emphasis>
|
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The
|
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<typeStatus id="54B0887E7828FFB2FE2AF8A49E0FF8BF" box="[462,571,1830,1856]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
|
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is from the Arenal Tilarán Conservation Area. The closely related species.
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<taxonomicName id="4C0B4D5F7828FFB2FE20F8CC9E9DF897" authority="Bates" authorityName="Bates" box="[452,681,1870,1896]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="soccata">
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<emphasis id="B97FEACE7828FFB2FE20F8CC9E55F898" box="[452,609,1870,1895]" italics="true" pageId="61" pageNumber="62">Agra soccata</emphasis>
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Bates
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</taxonomicName>
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of
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<collectingCountry id="F31C764C7828FFB2FD35F8CC9F1AF897" box="[721,814,1870,1896]" name="Panama" pageId="61" pageNumber="62">Panamá</collectingCountry>
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with which this female agrees in attributes provided the information on male attributes, hence my assignment of this species to the
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<taxonomicName id="4C0B4D5F782BFFB1FEECFEC69D6AFEA1" box="[264,350,324,350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="soccata">soccata</taxonomicName>
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group.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |