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<document id="57B464C361B915FEA4B7BCDB29E720AA" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7809FF92FEECFB969DD2FDD9" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra sirena Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="30" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="29" updateTime="1698190273211" updateUser="plazi">
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<mods:title id="29C3848704522DAA21706259A18612D7">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="450588BAF78F256355E1E0A39DD709A2" type="personal">
<mods:role id="102E075ED94B61555C83FE4A81A59FEC">
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<mods:namePart id="B8122843E4AD97263FE57308D642FE36">Erwin, Terry L.</mods:namePart>
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<mods:title id="157B278CE7450F7AED068F8CD6F1BC3D">Zootaxa</mods:title>
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<mods:date id="BFF6E3726DD9481F3B1460B378D3CDBC">2002</mods:date>
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<mods:identifier id="7FC6503BFDF70CE88037850F6035B0E6" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="F1A0FC85300AE1403E1AEB1C4525C496" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
<mods:identifier id="C6FC34144C2EFEDE81641F258E1401B3" type="ISSN">1175­5326</mods:identifier>
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<treatment id="03A287CA7809FF92FEECFB969DD2FDD9" ID-DOI="http://doi.org/10.5281/zenodo.6277982" ID-GBIF-Taxon="121065211" ID-Zenodo-Dep="6277982" LSID="urn:lsid:plazi:treatment:03A287CA7809FF92FEECFB969DD2FDD9" httpUri="http://treatment.plazi.org/id/03A287CA7809FF92FEECFB969DD2FDD9" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C31165577809FF93FEECFB969DCFFBA9" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BB436DC7809FF93FEECFB969EBEFBD2" blockId="28.[264,650,1043,1110]" box="[264,650,1043,1070]" pageId="28" pageNumber="29">
<heading id="D0FC81B07809FF93FEECFB969EBEFBD2" bold="true" box="[264,650,1043,1070]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="B97FEACE7809FF93FEECFB969EBEFBD2" bold="true" box="[264,650,1043,1070]" pageId="28" pageNumber="29">
<taxonomicName id="4C0B4D5F7809FF93FEECFB969DDAFBD2" ID-CoL="B2XT" authority="Erwin" authorityName="Erwin" box="[264,494,1043,1070]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sirena" status="sp. nov.">
<emphasis id="B97FEACE7809FF93FEECFB969DA3FBD1" bold="true" box="[264,407,1044,1070]" italics="true" pageId="28" pageNumber="29">Agra sirena</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B57809FF93FE1FFB919EBEFBD2" box="[507,650,1043,1069]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC7809FF93FEECFBBE9DCFFBA9" blockId="28.[264,650,1043,1110]" box="[264,507,1084,1110]" pageId="28" pageNumber="29">
(
<figureCitation id="13302A597809FF93FEF5FBBE9DB8FBA9" box="[273,396,1084,1110]" captionStart="FIGURE 9. A" captionStartId="29.[264,367,1401,1425]" captionTargetBox="[466,1120,612,1361]" captionTargetId="figure@29.[463,1123,609,1375]" captionTargetPageId="29" captionText="FIGURE 9. A. sirena, male, dorsal aspect: a) head; b) dorsal aspect, left elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.9; WE = 2.4; male SW = 4.6, female SW = 4.6; AL = 5.0." httpUri="https://zenodo.org/record/162813/files/figure.png" pageId="28" pageNumber="29">Figs. 9a, 9</figureCitation>
b, 9c, 9d)
</paragraph>
</subSubSection>
<subSubSection id="C31165577809FF93FEECFB09984AFB31" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BB436DC7809FF93FEECFB09984AFB31" blockId="28.[264,1324,1163,1870]" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEECFB099DB5FB5A" bold="true" box="[264,385,1163,1189]" pageId="28" pageNumber="29">
<typeStatus id="54B0887E7809FF93FEECFB099D48FB5A" box="[264,380,1163,1189]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male.
<emphasis id="B97FEACE7809FF93FE33FB099EA3FB5A" bold="true" box="[471,663,1163,1189]" pageId="28" pageNumber="29">
<collectingCountry id="F31C764C7809FF93FE33FB099EA6FB5A" box="[471,658,1163,1189]" name="Costa Rica" pageId="28" pageNumber="29">COSTA RICA</collectingCountry>
:
</emphasis>
Heredia, Estacíon La Selva,
<geoCoordinate id="EE3F501B7809FF93FC12FB0E9860FB59" box="[1014,1108,1164,1190]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="10.45">10º27'N</geoCoordinate>
<geoCoordinate id="EE3F501B7809FF93FBB9FB0E98F6FB59" box="[1117,1218,1164,1190]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="-83.98333">83º59'W</geoCoordinate>
, August, (R. Vargas &amp; D. Brenes)(
<taxonomicName id="4C0B4D5F7809FF93FDD7FB369EB4FB31" box="[563,640,1204,1230]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197809FF93FDD7FB369EB4FB31" box="[563,640,1204,1230]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="28" pageNumber="29">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197809FF93FD6FFB369E89FB31" LSID="urn:lsid:biocol.org:col:14323" box="[651,701,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="28" pageNumber="29">CRI</collectionCode>
002­416609), at black light, L/09/442.
</paragraph>
</subSubSection>
<subSubSection id="C31165577809FF93FEDCFB5998FDFA91" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BB436DC7809FF93FEDCFB5998FDFA91" blockId="28.[264,1324,1163,1870]" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEDCFB599D84FB0A" bold="true" box="[312,432,1243,1269]" pageId="28" pageNumber="29">Diagnosis</emphasis>
. Femur
<taxonomicName id="4C0B4D5F7809FF93FDFDFB5E9E0BFB09" box="[537,575,1244,1270]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
infuscated in basal two­thirds; elytron with sutural apex prolonged, broad. Male middle and posterior tibiae excavate and densely setose in apical half; elytral apex shallowly arcuate between lateral and apical apices, sutural apex acute,
<taxonomicName id="4C0B4D5F7809FF93FAE1FAAE991EFAB9" box="[1285,1322,1324,1350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
prolonged; metasternum pubescent; abdominal sterna 2­5 with 2 bilateral patches.
</paragraph>
</subSubSection>
<subSubSection id="C31165577809FF93FEDCFAF99D67F929" pageId="28" pageNumber="29" type="description">
<paragraph id="8BB436DC7809FF93FEDCFAF99D67F929" blockId="28.[264,1324,1163,1870]" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEDCFAF99DFDFA6A" bold="true" box="[312,457,1403,1429]" pageId="28" pageNumber="29">Description</emphasis>
. Color and luster: Piceous. Appendages rufous. Surface shiny. Form: Head (
<figureCitation id="13302A597809FF93FEBDFA269D9BFA41" box="[345,431,1444,1470]" captionStart="FIGURE 9. A" captionStartId="29.[264,367,1401,1425]" captionTargetBox="[466,1120,612,1361]" captionTargetId="figure@29.[463,1123,609,1375]" captionTargetPageId="29" captionText="FIGURE 9. A. sirena, male, dorsal aspect: a) head; b) dorsal aspect, left elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.9; WE = 2.4; male SW = 4.6, female SW = 4.6; AL = 5.0." httpUri="https://zenodo.org/record/162813/files/figure.png" pageId="28" pageNumber="29">Fig. 9a</figureCitation>
) behind eyes moderately elongate, tapered and rounded behind in female, tapered in male. Male middle and posterior tibiae excavate and densely setose in apical half. Elytron (
<figureCitation id="13302A597809FF93FE51FA769E34F9F1" box="[437,512,1524,1550]" captionStart="FIGURE 9. A" captionStartId="29.[264,367,1401,1425]" captionTargetBox="[466,1120,612,1361]" captionTargetId="figure@29.[463,1123,609,1375]" captionTargetPageId="29" captionText="FIGURE 9. A. sirena, male, dorsal aspect: a) head; b) dorsal aspect, left elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.9; WE = 2.4; male SW = 4.6, female SW = 4.6; AL = 5.0." httpUri="https://zenodo.org/record/162813/files/figure.png" pageId="28" pageNumber="29">Fig. 9</figureCitation>
b) with apex shallowly arcuate between lateral and sutural apices, sutural angle
<taxonomicName id="4C0B4D5F7809FF93FE48F99E9DE5F9C9" box="[428,465,1564,1590]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
elongate. Metasternum and posterior trochanter pubescent. Abdominal sterna 2­5 each with 2 bilateral patches. Abdominal sternum VI (
<figureCitation id="13302A597809FF93FBF3F9C6986BF9A1" box="[1047,1119,1604,1630]" captionStart="FIGURE 9. A" captionStartId="29.[264,367,1401,1425]" captionTargetBox="[466,1120,612,1361]" captionTargetId="figure@29.[463,1123,609,1375]" captionTargetPageId="29" captionText="FIGURE 9. A. sirena, male, dorsal aspect: a) head; b) dorsal aspect, left elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.9; WE = 2.4; male SW = 4.6, female SW = 4.6; AL = 5.0." httpUri="https://zenodo.org/record/162813/files/figure.png" pageId="28" pageNumber="29">Fig. 9</figureCitation>
c) moderately Vnotched in male and in female. Aedeagus (
<figureCitation id="13302A597809FF93FD13F9EE9F09F979" box="[759,829,1644,1670]" captionStart="FIGURE 9. A" captionStartId="29.[264,367,1401,1425]" captionTargetBox="[466,1120,612,1361]" captionTargetId="figure@29.[463,1123,609,1375]" captionTargetPageId="29" captionText="FIGURE 9. A. sirena, male, dorsal aspect: a) head; b) dorsal aspect, left elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.9; WE = 2.4; male SW = 4.6, female SW = 4.6; AL = 5.0." httpUri="https://zenodo.org/record/162813/files/figure.png" pageId="28" pageNumber="29">Fig. 9</figureCitation>
d) with ostium moderately long, apex simple diamond shape. Size: moderately large,
<quantity id="4CF39B397809FF93FCF9F9169FEAF951" box="[797,990,1684,1710]" metricMagnitude="-2" metricUnit="m" metricValue="1.8649999999999998" metricValueMax="2.02" metricValueMin="1.71" pageId="28" pageNumber="29" unit="mm" value="18.65" valueMax="20.2" valueMin="17.1">17.1 to 20.2mm</quantity>
in length, 4.8 to 5.0mm in width.
</paragraph>
</subSubSection>
<subSubSection id="C31165577809FF92FEA6F9619EABFE29" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BB436DC7809FF92FEA6F961991FFEA1" blockId="28.[264,1324,1163,1870]" lastBlockId="29.[264,1324,284,550]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEA6F9619E94F902" bold="true" box="[322,672,1763,1789]" pageId="28" pageNumber="29">Other specimens examined.</emphasis>
<collectingCountry id="F31C764C7809FF93FD4EF9669F07F901" box="[682,819,1764,1790]" name="Costa Rica" pageId="28" pageNumber="29">Costa Rica</collectingCountry>
: 2
<typeStatus id="54B0887E7809FF93FCBDF9669FF8F901" box="[857,972,1764,1790]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
, females, from the following Conservation Area: Osa, Cordillera Central. I have seen
<specimenCount id="9D0DFD557809FF93FC57F88E9830F8D9" box="[947,1028,1804,1830]" pageId="28" pageNumber="29" type="male">1 male</specimenCount>
from Veracruz,
<collectingCountry id="F31C764C7809FF93FB2EF88E9913F8D9" box="[1226,1319,1804,1830]" name="Mexico" pageId="28" pageNumber="29">México</collectingCountry>
, and labeled it as a
<typeStatus id="54B0887E7809FF93FE07F8B69E78F8B1" box="[483,588,1844,1870]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
.
<collectingCountry id="F31C764C7809FF93FDB3F8B69EEFF8B1" box="[599,731,1844,1870]" name="Costa Rica" pageId="28" pageNumber="29">Costa Rica</collectingCountry>
, Cartago, 1f,
<taxonomicName id="4C0B4D5F7809FF93FC9FF8B69FDEF8B1" box="[891,1002,1844,1870]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
,
<geoCoordinate id="EE3F501B7809FF93FC12F8B69869F8B1" box="[1014,1117,1844,1870]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="-83.63333">83°38'W</geoCoordinate>
,
<geoCoordinate id="EE3F501B7809FF93FB8DF8B6988BF8B1" box="[1129,1215,1844,1870]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="9.883333">9°53'N</geoCoordinate>
, January, (C. Barfield)(HRBURKE: ADP54403); Heredia; Puntarenas, 1f, P.N. Corcovado, Estacíon
<taxonomicName id="4C0B4D5F7808FF92FEECFEC69D62FEA1" box="[264,342,324,350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sirena">Sirena</taxonomicName>
,
<quantity id="4CF39B397808FF92FE85FEC69DF1FEA1" box="[353,453,324,350]" metricMagnitude="1" metricUnit="m" metricValue="5.05" metricValueMax="10.0" metricValueMin="0.1" pageId="29" pageNumber="30" unit="m" value="50.5" valueMax="100.0" valueMin="1.0">1 ­100m</quantity>
, LS 270500,508300 #2853, April, (G. Fonseca)(
<taxonomicName id="4C0B4D5F7808FF92FC19FEC69865FEA1" box="[1021,1105,324,350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197808FF92FC19FEC69865FEA1" box="[1021,1105,324,350]" pageId="29" pageNumber="30">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197808FF92FBBFFEC698BAFEA1" LSID="urn:lsid:biocol.org:col:14323" box="[1115,1166,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="29" pageNumber="30">CRI</collectionCode>
001­798662).
</paragraph>
<paragraph id="8BB436DC7808FF92FEDCFEE99EABFE29" blockId="29.[264,1324,284,550]" pageId="29" pageNumber="30">
<emphasis id="B97FEACE7808FF92FEDCFEE99DCAFE7A" bold="true" box="[312,510,363,389]" pageId="29" pageNumber="30">Specific epithet.</emphasis>
The specific epithet,
<emphasis id="B97FEACE7808FF92FD19FEEE9F79FE7A" box="[765,845,364,389]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="4C0B4D5F7808FF92FD19FEEE9F7CFE7A" box="[765,840,364,389]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sirena">sirena</taxonomicName>
,
</emphasis>
is used as a noun in apposition based on the Estacíon
<taxonomicName id="4C0B4D5F7808FF92FE44FE169DDFFE51" box="[416,491,404,430]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sirena">Sirena</taxonomicName>
in Corcovado National Park in reference to one of the places in which members of this species are found.
</paragraph>
</subSubSection>
<subSubSection id="C31165577808FF92FEDCFE619DD2FDD9" pageId="29" pageNumber="30" type="description">
<paragraph id="8BB436DC7808FF92FEDCFE619DD2FDD9" blockId="29.[264,1324,284,550]" pageId="29" pageNumber="30">
<emphasis id="B97FEACE7808FF92FEDCFE619D4AFE02" bold="true" box="[312,382,483,509]" pageId="29" pageNumber="30">Notes</emphasis>
. The large size and non­infuscated femora make this species readily recognizable in
<collectingCountry id="F31C764C7808FF92FEBBFD8E9DD7FDD9" box="[351,483,524,550]" name="Costa Rica" pageId="29" pageNumber="30">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>