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<document id="37782232FA2A7B763C70C8A1268BE0DA" ID-DOI="10.11646/zootaxa.1161.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5058330" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625190197324" checkinUser="felipe" docAuthor="Hippa, Heikki" docDate="2006" docId="5D4C87852848FFA15C1A0B68FE99FC3C" docLanguage="en" docName="zt01161p049.pdf" docOrigin="Zootaxa 1161 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Manota pollex Hippa 2006, sp. n." docType="treatment" docUuid="B44478C0-2B26-44C5-9752-A559A8C462CA" docUuidSource="ZooBank" docVersion="4" lastPageNumber="15" masterDocId="A175FFFD2844FFAF5D120E76FFC5FFE7" masterDocTitle="Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species" masterLastPageNumber="49" masterPageNumber="1" pageNumber="13" updateTime="1698983105622" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="435DE6C388F704C41666D6C57FDF1B5C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FC5B043AB8816FB07843D4A560C3B572">Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species</mods:title>
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<mods:namePart id="CDBC598583B8512166CBC2DFE7F20E12">Hippa, Heikki</mods:namePart>
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<mods:title id="AC238AF2B10BECA15DD89523E1BB7BBA">Zootaxa</mods:title>
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<mods:part id="01F8434887C1C26944958BCDD12B68BB">
<mods:date id="2DCCEF4818CD074168B3C507E54799FF">2006</mods:date>
<mods:detail id="B66AC507D99E399B4DCC4B91B47EB098" type="pubDate">
<mods:number id="64BDF3C090709374C9D7753C72F3B3AE">2006-03-30</mods:number>
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<mods:detail id="F46EE430F46757AB1FDA6360D40697FD" type="volume">
<mods:number id="35B745A4A2CCA60C7C0E67F72C04DD32">1161</mods:number>
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<mods:url id="9D41BECB1D0C2864B9F7D641DAD18034">https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1</mods:url>
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<mods:classification id="099A722E35D4D378F1FB01A8E6A1F13D">journal article</mods:classification>
<mods:identifier id="7DAF93327677327AF89EB654AEFF86F5" type="DOI">10.11646/zootaxa.1161.1.1</mods:identifier>
<mods:identifier id="7789E0F0156EB9FE0AB977E3056AD7FF" type="ISSN">1175­5334</mods:identifier>
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<treatment id="5D4C87852848FFA15C1A0B68FE99FC3C" ID-DOI="http://doi.org/10.5281/zenodo.5059284" ID-Zenodo-Dep="5059284" LSID="urn:lsid:zoobank.org:act:B44478C0-2B26-44C5-9752-A559A8C462CA" httpUri="http://treatment.plazi.org/id/5D4C87852848FFA15C1A0B68FE99FC3C" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="9DFF65182848FFA35C1A0B68FDC5FADF" box="[264,512,1310,1336]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="D55A36932848FFA35C1A0B68FDC5FADF" blockId="12.[264,512,1310,1371]" box="[264,512,1310,1336]" pageId="12" pageNumber="13">
<heading id="8E1281FF2848FFA35C1A0B68FDC5FADF" bold="true" box="[264,512,1310,1336]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="E791EA812848FFA35C1A0B68FDC5FADF" bold="true" box="[264,512,1310,1336]" pageId="12" pageNumber="13">
<taxonomicName id="12E54D102848FFA35C1A0B68FE73FADF" ID-CoL="3XVY5" authority="Hippa, 2006" authorityName="Hippa" authorityYear="2006" box="[264,438,1310,1336]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pollex" status="sp. nov.">
<emphasis id="E791EA812848FFA35C1A0B68FE73FADF" bold="true" box="[264,438,1310,1336]" italics="true" pageId="12" pageNumber="13">Manota pollex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCA257FA2848FFA35CAC0B68FDC5FADF" box="[446,512,1310,1336]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9DFF65182848FFA15C1A0B37FE99FC3C" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="D55A36932848FFA35C1A0B37FE66FABC" blockId="12.[264,512,1310,1371]" box="[264,419,1345,1371]" pageId="12" pageNumber="13">
(
<figureCitation id="4DDE2A162848FFA35C030B37FE5FFABC" box="[273,410,1345,1371]" captionStart="FIGURE 4" captionStartId="13.[264,368,1799,1823]" captionTargetBox="[407,1166,319,1782]" captionTargetId="figure-59@13.[381,1206,307,1806]" captionTargetPageId="13" captionText="FIGURE 4. Antennal flagellomere 4, lateral view (A, C) and hypopygium, ventral view (C, D). A, B. Manota yongi sp. n. (holotype). C, D. Manota pollex sp. n. (holotype). Scale for A and C 0.05 mm, for B and D 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058340" httpUri="https://zenodo.org/record/5058340/files/figure.png" pageId="12" pageNumber="13">Figs. 4C, D</figureCitation>
)
</paragraph>
<paragraph id="D55A36932848FFA35C1A0BFAFDD6FA41" blockId="12.[264,1323,1420,1846]" box="[264,531,1420,1446]" pageId="12" pageNumber="13">
A small­sized
<taxonomicName id="12E54D102848FFA35CA30BFAFDC9FA42" authorityName="Williston" authorityYear="1896" box="[433,524,1420,1445]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="E791EA812848FFA35CA30BFAFDC9FA42" box="[433,524,1420,1445]" italics="true" pageId="12" pageNumber="13">Manota</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D55A36932848FFA15C2A0BC2FDA0FE49" blockId="12.[264,1323,1420,1846]" lastBlockId="14.[264,1323,284,470]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
Male.
<emphasis id="E791EA812848FFA35C910BC2FE03FA29" bold="true" box="[387,454,1460,1486]" pageId="12" pageNumber="13">Head</emphasis>
. Flagellomere 4,
<figureCitation id="4DDE2A162848FFA35F860BC2FD2AFA29" box="[660,751,1460,1486]" captionStart="FIGURE 4" captionStartId="13.[264,368,1799,1823]" captionTargetBox="[407,1166,319,1782]" captionTargetId="figure-59@13.[381,1206,307,1806]" captionTargetPageId="13" captionText="FIGURE 4. Antennal flagellomere 4, lateral view (A, C) and hypopygium, ventral view (C, D). A, B. Manota yongi sp. n. (holotype). C, D. Manota pollex sp. n. (holotype). Scale for A and C 0.05 mm, for B and D 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058340" httpUri="https://zenodo.org/record/5058340/files/figure.png" pageId="12" pageNumber="13">Fig. 4C</figureCitation>
. Maxillary palpus with palpomere 3 bearing 4–5 apically expanded curved sensilla; palpomere 4 with parasegment.
<emphasis id="E791EA812848FFA3590B0BAAFBB8FA11" bold="true" box="[1049,1149,1500,1526]" pageId="12" pageNumber="13">Thorax.</emphasis>
Anepisternum setose. Anterior basalare non­setose. Preepisternum 2 setose. Laterotergite setose. Episternum 3 setose.
<emphasis id="E791EA812848FFA35F18085AFD8BF9A1" bold="true" box="[522,590,1580,1606]" pageId="12" pageNumber="13">Wing</emphasis>
. Length
<quantity id="121D9B762848FFA35FA9085AFCDFF9A1" box="[699,794,1580,1606]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="13" unit="mm" value="1.7">1.7 mm</quantity>
. Wing membrane dorsally with few setae at posterior margin. Sc distally of h non­setose. A1 indicated by few setae.
<emphasis id="E791EA812848FFA359950822FAEFF989" bold="true" box="[1159,1322,1620,1646]" pageId="12" pageNumber="13">Hypopygium</emphasis>
(
<figureCitation id="4DDE2A162848FFA35C03080AFEB1F971" box="[273,372,1660,1686]" captionStart="FIGURE 4" captionStartId="13.[264,368,1799,1823]" captionTargetBox="[407,1166,319,1782]" captionTargetId="figure-59@13.[381,1206,307,1806]" captionTargetPageId="13" captionText="FIGURE 4. Antennal flagellomere 4, lateral view (A, C) and hypopygium, ventral view (C, D). A, B. Manota yongi sp. n. (holotype). C, D. Manota pollex sp. n. (holotype). Scale for A and C 0.05 mm, for B and D 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058340" httpUri="https://zenodo.org/record/5058340/files/figure.png" pageId="12" pageNumber="13">Fig. 4D</figureCitation>
). Sternite 9 short, one­fourth of ventral length of gonocoxa, laterally sharply delimited, posterior margin not clearly contiguous so that sternite gives impression of being medially divided, setae stronger than ventral setae of gonocoxa. Gonocoxa ventrally with rounded lobe apicomesially, lobe with stronger setae deviating from adjacent setosity; dorsally in position I with rounded lobe bearing 1 megaseta, 1 flattened megaseta in position III which basally may have a branch, no apicolateral triangular lobe. Gonostylus elongate, laterally angularly convex, mesially nearly straight, apically almost pointed. Parastylar lobe lacking. Tegmen long and unusually narrow, with slight lateral shoulders. Setae on sternite 10 scattered.
</paragraph>
<caption id="819A661B2849FFA25C1A0971FDEFF880" ID-DOI="http://doi.org/10.5281/zenodo.5058340" ID-Zenodo-Dep="5058340" httpUri="https://zenodo.org/record/5058340/files/figure.png" pageId="13" pageNumber="14" startId="13.[264,368,1799,1823]" targetBox="[407,1166,319,1782]" targetPageId="13">
<paragraph id="D55A36932849FFA25C1A0971FDEFF880" blockId="13.[264,1323,1799,1895]" pageId="13" pageNumber="14">
<emphasis id="E791EA812849FFA25C1A0971FE4FF8F8" bold="true" box="[264,394,1799,1823]" pageId="13" pageNumber="14">FIGURE 4.</emphasis>
Antennal flagellomere 4, lateral view (A, C) and hypopygium, ventral view (C, D).
<emphasis id="E791EA812849FFA258030971FEE5F8A3" bold="true" pageId="13" pageNumber="14">A, B.</emphasis>
<taxonomicName id="12E54D102849FFA25C3A095AFE78F8A4" authority="Hippa, 2006" authorityName="Hippa" authorityYear="2006" box="[296,445,1836,1859]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="yongi" status="sp. nov.">
<emphasis id="E791EA812849FFA25C3A095AFE78F8A4" box="[296,445,1836,1859]" italics="true" pageId="13" pageNumber="14">Manota yongi</emphasis>
</taxonomicName>
<emphasis id="E791EA812849FFA25CD6095CFDC3F8A3" bold="true" box="[452,518,1834,1860]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="FCA257FA2849FFA25CD6095CFDC3F8A3" box="[452,518,1834,1860]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype).
<emphasis id="E791EA812849FFA25F98095AFD00F8A3" bold="true" box="[650,709,1836,1860]" pageId="13" pageNumber="14">C, D.</emphasis>
<taxonomicName id="12E54D102849FFA25FDE095AFCA2F8A4" authority="Hippa, 2006" authorityName="Hippa" authorityYear="2006" box="[716,871,1836,1859]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pollex" status="sp. nov.">
<emphasis id="E791EA812849FFA25FDE095AFCA2F8A4" box="[716,871,1836,1859]" italics="true" pageId="13" pageNumber="14">Manota pollex</emphasis>
</taxonomicName>
<emphasis id="E791EA812849FFA25E7C095CFC74F8A3" bold="true" box="[878,945,1834,1860]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="FCA257FA2849FFA25E7C095CFC74F8A3" box="[878,945,1834,1860]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). Scale for A and C 0.05 mm, for B and D 0.10 mm.
</paragraph>
</caption>
<paragraph id="D55A3693284AFFA15C2A0FCAFCDAFE31" blockId="14.[264,1323,284,470]" box="[312,799,444,470]" pageId="14" pageNumber="15">Female and preimaginal stages unknown.</paragraph>
<paragraph id="D55A3693284AFFA15C1A0C7FFE4CFDC5" blockId="14.[264,1323,521,747]" box="[264,393,521,546]" pageId="14" pageNumber="15">
<emphasis id="E791EA81284AFFA15C1A0C7FFE4CFDC5" box="[264,393,521,546]" italics="true" pageId="14" pageNumber="15">Discussion</emphasis>
</paragraph>
<paragraph id="D55A3693284AFFA15C2A0C47FC80FD0C" blockId="14.[264,1323,521,747]" pageId="14" pageNumber="15">
<taxonomicName id="12E54D10284AFFA15C2A0C47FE27FDAD" authorityName="Hippa" authorityYear="2006" box="[312,482,561,586]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pollex">
<emphasis id="E791EA81284AFFA15C2A0C47FE27FDAD" box="[312,482,561,586]" italics="true" pageId="14" pageNumber="15">Manota pollex</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="12E54D10284AFFA15F930C47FD2FFDAD" authorityName="Hippa" authorityYear="2006" box="[641,746,561,586]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="yongi">
<emphasis id="E791EA81284AFFA15F930C47FD2FFDAD" box="[641,746,561,586]" italics="true" pageId="14" pageNumber="15">M. yongi</emphasis>
</taxonomicName>
. It differs from the latter by having the group of stronger setae apicomesially on the ventral side of the gonocoxa on a small rounded lobe. Furthermore, the apical part of the gonostylus is less narrowed or pointed and the lobe dorsally on the gonocoxa in position I is narrower. Both species are rather similar to
<taxonomicName id="12E54D10284AFFA158190CDFFE80FD0D" authorityName="Hippa" authorityYear="2006" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roslii">
<emphasis id="E791EA81284AFFA158190CDFFE80FD0D" italics="true" pageId="14" pageNumber="15">M. roslii</emphasis>
</taxonomicName>
. For further discussion, see under
<taxonomicName id="12E54D10284AFFA15FC80CA7FCFBFD0D" authorityName="Hippa" authorityYear="2006" box="[730,830,721,746]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roslii">
<emphasis id="E791EA81284AFFA15FC80CA7FCFBFD0D" box="[730,830,721,746]" italics="true" pageId="14" pageNumber="15">M. roslii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D55A3693284AFFA15C1A0D57FE8FFCDD" blockId="14.[264,1323,801,987]" box="[264,330,801,826]" pageId="14" pageNumber="15">
<emphasis id="E791EA81284AFFA15C1A0D57FE8FFCDD" box="[264,330,801,826]" italics="true" pageId="14" pageNumber="15">Types</emphasis>
</paragraph>
<paragraph id="D55A3693284AFFA15C2A0D3FFE99FC3C" blockId="14.[264,1323,801,987]" pageId="14" pageNumber="15">
<materialsCitation id="658D3CCE284AFFA15C2A0D3FFE02FC54" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<emphasis id="E791EA81284AFFA15C2A0D3FFE66FC85" box="[312,419,841,866]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="0A5E8831284AFFA15C2A0D3FFE66FC85" box="[312,419,841,866]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCounty id="3C3B4E1F284AFFA15CA60D3FFE36FC84" box="[436,499,841,867]" pageId="14" pageNumber="15">Male</collectingCounty>
,
<collectingMunicipality id="353EACE9284AFFA15F100D3FFDF9FC84" box="[514,572,841,867]" pageId="14" pageNumber="15">West</collectingMunicipality>
<collectingCountry id="ADF27603284AFFA15F570D3FFD76FC84" box="[581,691,841,867]" name="Malaysia" pageId="14" pageNumber="15">Malaysia</collectingCountry>
,
<collectingRegion id="1721F871284AFFA15FD30D3FFCEEFC84" box="[705,811,841,867]" country="Malaysia" name="Selangor" pageId="14" pageNumber="15">Selangor</collectingRegion>
,
<location id="D03A6048284AFFA15E280D3FFC12FC84" LSID="urn:lsid:plazi:treatment:5D4C87852848FFA15C1A0B68FE99FC3C:D03A6048284AFFA15E280D3FFC12FC84" box="[826,983,841,867]" country="Malaysia" county="Male" municipality="West" name="Ulu Gombak" pageId="14" pageNumber="15" stateProvince="Selangor">Ulu Gombak</location>
,
<location id="D03A6048284AFFA15EF70D3FFE60FC6C" LSID="urn:lsid:plazi:treatment:5D4C87852848FFA15C1A0B68FE99FC3C:D03A6048284AFFA15EF70D3FFE60FC6C" country="Malaysia" county="Male" municipality="West" name="University of Malaya Field Study Centre" pageId="14" pageNumber="15" stateProvince="Selangor">University of Malaya Field Study Centre</location>
,
<quantity id="121D9B76284AFFA15CA00D07FE3EFC6C" box="[434,507,881,907]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="14" pageNumber="15" unit="ft" value="800.0">
<elevation id="5EC8D1A0284AFFA15CA00D07FE3EFC6C" box="[434,507,881,907]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="14" pageNumber="15" unit="ft" value="800.0">800 ft</elevation>
</quantity>
, Malaise in jungle,
<date id="A15B1053284AFFA15FF60D07FBD4FC6C" box="[740,1041,881,907]" pageId="14" pageNumber="15" value="1997-02-24" valueMax="1997-03-21" valueMin="1997-02-24">
<collectingDate id="B11FE9BB284AFFA15FF60D07FBD4FC6C" box="[740,1041,881,907]" pageId="14" pageNumber="15" value="1997-02-24" valueMax="1997-03-21" valueMin="1997-02-24">24. Feb.–21. March 1997</collectingDate>
</date>
,
<collectorName id="78105345284AFFA1590D0D07FB4BFC6C" box="[1055,1166,881,907]" pageId="14" pageNumber="15">H. Hippa</collectorName>
,
<collectorName id="78105345284AFFA159880D07FAEFFC6C" box="[1178,1322,881,907]" pageId="14" pageNumber="15">M. Jaschhof</collectorName>
and
<collectorName id="78105345284AFFA15C2C0DEFFE02FC54" box="[318,455,921,947]" pageId="14" pageNumber="15">B. Viklund</collectorName>
</materialsCitation>
.
<materialsCitation id="658D3CCE284AFFA15CC50DEFFE9DFC3C" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Selangor" typeStatus="paratype">
<typeStatus id="0A5E8831284AFFA15CC50DEFFD84FC54" box="[471,577,921,947]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
.
<specimenCount id="C3E3FD1A284AFFA15F430DEFFD61FC54" box="[593,676,921,947]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
with same data as holotype except for 22. Feb.–21. March
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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