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<document id="68DBFF4E55A9BE408937DDD143BA9B4C" ID="10.11646/zootaxa.4169.1.5" ID-DOI="10.11646/zootaxa.4169.1.5" ID-GBIF-Dataset="e35dbd92-b156-4ae3-9f90-9996c91ee626" ID-ISSN="1175-5326" ID-Zenodo-Dep="265168" ID-ZooBank="62419F0E-128A-4E60-932C-DD746BA57906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474268893273" checkinUser="plazi" docAuthor="Thoni, R. J., Gurung, D. B. &amp; Mayden, R. L." docDate="2016" docId="03F187D8FFB2FF8CFF1BFA08FD3EF828" docLanguage="en" docName="zootaxa.4169.1.5.pdf" docOrigin="Zootaxa 4169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra gotyla Gray 1830" docType="treatment" docVersion="7" lastPageNumber="123" masterDocId="FFC8FFA0FFBAFF84FF8CFFB8FFA3FF9D" masterDocTitle="A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)" masterLastPageNumber="132" masterPageNumber="115" pageNumber="123" updateTime="1698386207380" updateUser="plazi">
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<mods:title id="EC4040D13560A843716F54C65A8FAA75">A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)</mods:title>
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<mods:namePart id="4D57EB718B7B78D32593248D72EE66F1">Thoni, R. J.</mods:namePart>
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<mods:namePart id="86B9D9AF10E8F1619B99830739BFBE23">Gurung, D. B.</mods:namePart>
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<mods:namePart id="25482AA4AF79B6A20DFBD42D53CBEF01">Mayden, R. L.</mods:namePart>
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<mods:title id="53305406602F839E7B0C8A829FD922E4">Zootaxa</mods:title>
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<mods:date id="629BC6C32F827F0948E7318444681732">2016</mods:date>
<mods:detail id="D4B51B250BC621420DE5D5D1B952BFB7" type="volume">
<mods:number id="D7668517B84B32FAE20E87274A5888E4">4169</mods:number>
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<mods:identifier id="E2B86D186E8C01DDCA447766398D26E2" type="DOI">10.11646/zootaxa.4169.1.5</mods:identifier>
<mods:identifier id="6E56D3C3AA2C77E5A149D86AB1B457C9" type="GBIF-Dataset">e35dbd92-b156-4ae3-9f90-9996c91ee626</mods:identifier>
<mods:identifier id="28F2A592045ECD948FC4FD58E0F45B71" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C7AE1D8A77F5697FE1BF7244A33FC076" type="Zenodo-Dep">265168</mods:identifier>
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<treatment id="03F187D8FFB2FF8CFF1BFA08FD3EF828" ID-DOI="http://doi.org/10.5281/zenodo.6087321" ID-GBIF-Taxon="124551997" ID-Zenodo-Dep="6087321" LSID="urn:lsid:plazi:treatment:03F187D8FFB2FF8CFF1BFA08FD3EF828" httpUri="http://treatment.plazi.org/id/03F187D8FFB2FF8CFF1BFA08FD3EF828" lastPageNumber="123" pageId="8" pageNumber="123">
<subSubSection id="C3426545FFB2FF8CFF1BFA08FD13F9B6" pageId="8" pageNumber="123" type="nomenclature">
<paragraph id="8BE736CEFFB2FF8CFF1BFA08FE77FA57" blockId="8.[151,468,1456,1482]" box="[151,468,1456,1482]" pageId="8" pageNumber="123">
<heading id="D0AF81A2FFB2FF8CFF1BFA08FE77FA57" bold="true" box="[151,468,1456,1482]" fontSize="11" level="1" pageId="8" pageNumber="123" reason="1">
<taxonomicName id="4C584D4DFFB2FF8CFF1BFA08FE77FA57" ID-CoL="3FBJH" authority="Gray, 1830" authorityName="Gray" authorityYear="1830" box="[151,468,1456,1482]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="123" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="B92CEADCFFB2FF8CFF1BFA08FE77FA57" bold="true" box="[151,468,1456,1482]" pageId="8" pageNumber="123">
<emphasis id="B92CEADCFFB2FF8CFF1BFA08FE93FA57" bold="true" box="[151,304,1456,1482]" italics="true" pageId="8" pageNumber="123">Garra gotyla</emphasis>
(Gray, 1830)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE736CEFFB2FF8CFF1BFA40FD13F9B6" blockId="8.[151,1436,1527,1579]" pageId="8" pageNumber="123">
<taxonomicName id="4C584D4DFFB2FF8CFF1BFA40FE12F990" authority="Gray 1830" authorityName="Gray" authorityYear="1830" box="[151,433,1527,1549]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="123" phylum="Chordata" rank="species" species="gotlya">
<emphasis id="B92CEADCFFB2FF8CFF1BFA40FE99F990" box="[151,314,1527,1549]" italics="true" pageId="8" pageNumber="123">Cyprinus gotlya</emphasis>
Gray 1830
</taxonomicName>
. Type locality: Mountain stream in Northern India. No Holotype. Neotype: MUMF4300; Neotype locality: India: Sikkim: Tista River and Rangpo.
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFB2FF8CFF1BF9E0FDF1F8B8" pageId="8" pageNumber="123" type="materials_examined">
<paragraph id="8BE736CEFFB2FF8CFF1BF9E0FDF1F8B8" blockId="8.[151,1437,1624,1973]" pageId="8" pageNumber="123">
<materialsCitation id="3B303C93FFB2FF8CFF1BF9E0FDEEF908" ID-GBIF-Occurrence="1313779485" collectingDate="2013-08-09" collectionCode="CNR" collectorName="Damdum, D. B. &amp; R. J. Thoni" country="Bhutan" location="Damdum" pageId="8" pageNumber="123" specimenCode="CNR 15023, 15026" specimenCount="2" stateProvince="Samtse">
<emphasis id="B92CEADCFFB2FF8CFF1BF9E0FE28F9EC" bold="true" box="[151,395,1624,1649]" pageId="8" pageNumber="123">Materials examined.</emphasis>
<specimenCount id="9D5EFD47FFB2FF8CFE1DF9E1FD81F9EC" box="[401,546,1624,1649]" pageId="8" pageNumber="123" type="generic">2 Specimens</specimenCount>
,
<specimenCode id="DBFE9EB5FFB2FF8CFDA1F9E1FCAAF9EC" box="[557,777,1624,1649]" pageId="8" pageNumber="123">
<collectionCode id="ED49AE0BFFB2FF8CFDA1F9E1FDC4F9EC" box="[557,615,1625,1649]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="8" pageNumber="123">CNR</collectionCode>
15023, 15026
</specimenCode>
;
<collectingCountry id="F34F765EFFB2FF8CFC99F9E0FCC9F9EC" box="[789,874,1624,1649]" name="Bhutan" pageId="8" pageNumber="123">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFB2FF8CFCF8F9E1FC66F9EC" box="[884,965,1625,1649]" country="Bhutan" name="Samchi" pageId="8" pageNumber="123">Samtse</collectingRegion>
Dzongkhag:
<quantity id="4CA09B2BFFB2FF8CFBD7F9E0FB30F9EC" box="[1115,1171,1624,1649]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="123" unit="km" value="6.0">6 km</quantity>
west of
<collectorName id="26AD5318FFB2FF8CFB7EF9E0FA3FF9ED" box="[1266,1436,1624,1649]" pageId="8" pageNumber="123">
<location id="8E876015FFB2FF8CFB7EF9E0FAFEF9EC" LSID="urn:lsid:plazi:treatment:03F187D8FFB2FF8CFF1BFA08FD3EF828:8E876015FFB2FF8CFB7EF9E0FAFEF9EC" box="[1266,1373,1624,1649]" country="Bhutan" name="Damdum" pageId="8" pageNumber="123" stateProvince="Samtse">Damdum</location>
, D.B.
</collectorName>
Gurung &amp;
<collectorName id="26AD5318FFB2FF8CFE98F9C5FE2DF908" box="[276,398,1660,1685]" pageId="8" pageNumber="123">R.J. Thoni</collectorName>
,
<collectingDate id="EFA2E9E6FFB2FF8CFE16F9C5FDEEF908" box="[410,589,1661,1685]" pageId="8" pageNumber="123" value="2013-08-09">
<date id="FFE6100EFFB2FF8CFE16F9C5FDEEF908" box="[410,589,1661,1685]" pageId="8" pageNumber="123" value="2013-08-09">August 9, 2013</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B303C93FFB2FF8CFDD5F9C5FD03F924" ID-GBIF-Occurrence="1313779477" collectingDate="2013-08-09" collectionCode="CNR" collectorName="Damdum, D. B. &amp; R. J. Thoni" country="Bhutan" location="Samtse" pageId="8" pageNumber="123" specimenCode="CNR 13012-13014" specimenCount="3" stateProvince="Samtse">
<specimenCount id="9D5EFD47FFB2FF8CFDD5F9C5FD48F908" box="[601,747,1660,1685]" pageId="8" pageNumber="123" type="generic">3 Specimens</specimenCount>
,
<specimenCode id="DBFE9EB5FFB2FF8CFD74F9C5FC77F908" box="[760,980,1661,1685]" collectionCode="CNR" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="8" pageNumber="123">
<collectionCode id="ED49AE0BFFB2FF8CFD74F9C5FC91F908" box="[760,818,1661,1685]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="8" pageNumber="123">CNR</collectionCode>
13012-13014
</specimenCode>
;
<collectingCountry id="F34F765EFFB2FF8CFC6CF9C4FB96F908" box="[992,1077,1660,1685]" name="Bhutan" pageId="8" pageNumber="123">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFB2FF8CFBCEF9C5FB30F908" box="[1090,1171,1661,1685]" country="Bhutan" name="Samchi" pageId="8" pageNumber="123">Samtse</collectingRegion>
Dzongkhag:
<collectorName id="26AD5318FFB2FF8CFAA1F9C4FF6FF925" pageId="8" pageNumber="123">Damdum, D.B.</collectorName>
Gurung &amp;
<collectorName id="26AD5318FFB2FF8CFED6F919FE74F924" box="[346,471,1696,1721]" pageId="8" pageNumber="123">R.J. Thoni</collectorName>
,
<collectingDate id="EFA2E9E6FFB2FF8CFE6BF919FD03F924" box="[487,672,1697,1721]" pageId="8" pageNumber="123" value="2013-08-09">
<date id="FFE6100EFFB2FF8CFE6BF919FD03F924" box="[487,672,1697,1721]" pageId="8" pageNumber="123" value="2013-08-09">August 9, 2013</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B303C93FFB2FF8CFD22F919FD41F940" ID-GBIF-Occurrence="1313779493" collectingDate="2013-08" collectionCode="CNR" collectorName="Chukkha Dzongkhag &amp; Above Pasakha, D. B. &amp; R. J. Thoni" country="Bhutan" location="Bhutan" pageId="8" pageNumber="123" specimenCount="1">
<specimenCount id="9D5EFD47FFB2FF8CFD22F919FC9AF924" box="[686,825,1696,1721]" pageId="8" pageNumber="123" type="generic">1 Specimen</specimenCount>
,
<collectionCode id="ED49AE0BFFB2FF8CFCC5F919FC20F924" box="[841,899,1697,1721]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="8" pageNumber="123">CNR</collectionCode>
15052;
<collectingCountry id="F34F765EFFB2FF8CFC64F918FB9EF924" box="[1000,1085,1696,1721]" name="Bhutan" pageId="8" pageNumber="123">Bhutan</collectingCountry>
:
<collectorName id="26AD5318FFB2FF8CFBC0F918FAE1F924" box="[1100,1346,1696,1721]" pageId="8" pageNumber="123">Chukkha Dzongkhag</collectorName>
:
<collectorName id="26AD5318FFB2FF8CFADEF918FE9BF941" pageId="8" pageNumber="123">Above Pasakha, D.B.</collectorName>
Gurung &amp;
<collectorName id="26AD5318FFB2FF8CFE4DF97DFD9EF940" box="[449,573,1732,1757]" pageId="8" pageNumber="123">R.J. Thoni</collectorName>
,
<collectingDate id="EFA2E9E6FFB2FF8CFDC7F97DFD41F940" box="[587,738,1733,1757]" pageId="8" pageNumber="123" value="2013-08">
<date id="FFE6100EFFB2FF8CFDC7F97DFD41F940" box="[587,738,1733,1757]" pageId="8" pageNumber="123" value="2013-08">August 2013</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B303C93FFB2FF8CFD63F97DFDF1F89C" ID-GBIF-Occurrence="1313779490" collectingDate="2013-10-09" collectionCode="CNR" collectorName="R. J. Thoni &amp; D. B. Gurung" country="Bhutan" location="Samtse" pageId="8" pageNumber="123" specimenCount="1" stateProvince="Samtse">
<specimenCount id="9D5EFD47FFB2FF8CFD63F97DFCDBF940" box="[751,888,1732,1757]" pageId="8" pageNumber="123" type="generic">1 Specimen</specimenCount>
,
<collectionCode id="ED49AE0BFFB2FF8CFC0AF97DFC63F940" box="[902,960,1733,1757]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="8" pageNumber="123">CNR</collectionCode>
15029;
<collectingCountry id="F34F765EFFB2FF8CFBADF97CFBD5F940" box="[1057,1142,1732,1757]" name="Bhutan" pageId="8" pageNumber="123">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFB2FF8CFB08F97DFB76F940" box="[1156,1237,1733,1757]" country="Bhutan" name="Samchi" pageId="8" pageNumber="123">Samtse</collectingRegion>
Dzongkhag,
<collectorName id="26AD5318FFB2FF8CFAFDF97DFF79F89C" pageId="8" pageNumber="123">R.J. Thoni</collectorName>
&amp;
<collectorName id="26AD5318FFB2FF8CFF70F951FE2CF89C" box="[252,399,1769,1793]" pageId="8" pageNumber="123">D.B. Gurung</collectorName>
,
<collectingDate id="EFA2E9E6FFB2FF8CFE16F950FDF1F89C" box="[410,594,1768,1793]" pageId="8" pageNumber="123" value="2013-10-09">
<date id="FFE6100EFFB2FF8CFE16F950FDF1F89C" box="[410,594,1768,1793]" pageId="8" pageNumber="123" value="2013-10-09">October 9, 2013</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B303C93FFB2FF8CFDD7F951FDEDF8B8" ID-GBIF-Occurrence="1313779482" collectingDate="2013-09" collectionCode="CNR" collectorName="Chungpatang, D. B. &amp; R. J. Thoni" country="Bhutan" location="Samtse" pageId="8" pageNumber="123" specimenCount="1" stateProvince="Samtse">
<specimenCount id="9D5EFD47FFB2FF8CFDD7F951FD43F89C" box="[603,736,1768,1793]" pageId="8" pageNumber="123" type="generic">1 Specimen</specimenCount>
,
<collectionCode id="ED49AE0BFFB2FF8CFD67F951FC86F89C" box="[747,805,1769,1793]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="8" pageNumber="123">CNR</collectionCode>
15015;
<collectingCountry id="F34F765EFFB2FF8CFCF3F950FC77F89C" box="[895,980,1768,1793]" name="Bhutan" pageId="8" pageNumber="123">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFB2FF8CFC52F951FB8CF89C" box="[990,1071,1769,1793]" country="Bhutan" name="Samchi" pageId="8" pageNumber="123">Samtse</collectingRegion>
Dzongkhag:
<collectorName id="26AD5318FFB2FF8CFB49F950FA3FF89D" box="[1221,1436,1768,1793]" pageId="8" pageNumber="123">Chungpatang, D.B.</collectorName>
Gurung &amp;
<collectorName id="26AD5318FFB2FF8CFE9CF8B5FE2AF8B8" box="[272,393,1804,1829]" pageId="8" pageNumber="123">R.J. Thoni</collectorName>
,
<collectingDate id="EFA2E9E6FFB2FF8CFE18F8B4FDEDF8B8" box="[404,590,1804,1829]" pageId="8" pageNumber="123" value="2013-09">
<date id="FFE6100EFFB2FF8CFE18F8B4FDEDF8B8" box="[404,590,1804,1829]" pageId="8" pageNumber="123" value="2013-09">September 2013</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFB2FF8CFF4BF888FD3EF828" pageId="8" pageNumber="123" type="discussion">
<paragraph id="8BE736CEFFB2FF8CFF4BF888FD3EF828" blockId="8.[151,1437,1624,1973]" pageId="8" pageNumber="123">
<emphasis id="B92CEADCFFB2FF8CFF4BF888FE98F8D4" bold="true" box="[199,315,1840,1865]" pageId="8" pageNumber="123">Remarks.</emphasis>
For a full discussion on the taxonomy of
<taxonomicName id="4C584D4DFFB2FF8CFCBEF889FC38F8D4" box="[818,923,1840,1865]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="123" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="B92CEADCFFB2FF8CFCBEF889FC38F8D4" box="[818,923,1840,1865]" italics="true" pageId="8" pageNumber="123">G. gotyla</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFC94B3FFFB2FF8CFC59F888FAF8F8D4" author="Nebeshwar" box="[981,1371,1840,1865]" pageId="8" pageNumber="123" refString="Nebeshwar &amp; Vishwanath (2013) Three new species of Garra (Pisces: Cyprinidae) from north-eastern India and redescription of G. gotyla. Ichthyological Exploration of Freshwaters, 24 (2) 97 - 120." type="journal article" year="2013">Nebeshwar &amp; Vishwanath (2013)</bibRefCitation>
. This species was encountered in Southwestern
<collectingCountry id="F34F765EFFB2FF8CFDE2F8ECFD61F8F0" box="[622,706,1876,1901]" name="Bhutan" pageId="8" pageNumber="123">Bhutan</collectingCountry>
, in the rivers of
<collectingRegion id="499CF82CFFB2FF8CFCF5F8EDFC6EF8F0" box="[889,973,1877,1901]" country="Bhutan" name="Samchi" pageId="8" pageNumber="123">Samtse</collectingRegion>
, Chukkha,
<collectingRegion id="499CF82CFFB2FF8CFBC7F8EDFB00F8F0" box="[1099,1187,1877,1901]" country="Bhutan" name="Daga" pageId="8" pageNumber="123">Dagana</collectingRegion>
,
<collectingRegion id="499CF82CFFB2FF8CFB23F8ECFAA6F8F0" box="[1199,1285,1876,1901]" country="Bhutan" name="Geylegphug" pageId="8" pageNumber="123">Tsirang</collectingRegion>
, and Sarpang Dzongkhags (vouchers not collected from all locations). This species was identified by the diagnostic criteria provided in
<bibRefCitation id="EFC94B3FFFB2FF8CFE92F824FD3AF828" author="Nebeshwar" box="[286,665,1948,1973]" pageId="8" pageNumber="123" refString="Nebeshwar &amp; Vishwanath (2013) Three new species of Garra (Pisces: Cyprinidae) from north-eastern India and redescription of G. gotyla. Ichthyological Exploration of Freshwaters, 24 (2) 97 - 120." type="journal article" year="2013">Nebeshwar &amp; Vishwanath (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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