254 lines
28 KiB
XML
254 lines
28 KiB
XML
<document id="920A9E182A4342235114FC72CB16A4F0" ID="10.11646/zootaxa.4125.1.1" ID-CLB-Dataset="38807" ID-DOI="10.11646/zootaxa.4125.1.1" ID-GBIF-Dataset="1af13aec-3c54-4a92-abdf-e4e29b97e82f" ID-ISSN="1175-5326" ID-Zenodo-Dep="271704" ID-ZooBank="118F4865-D89E-45EA-A210-8D61946CC37F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465971881907" checkinUser="plazi" docAuthor="Matz, Jess & Brower, Andrew V. Z." docDate="2016" docId="03F187D7FF9B846DFF11FD84FB32BC32" docLanguage="en" docName="zootaxa.4125.1.1.pdf" docOrigin="Zootaxa 4125 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argyrophorus Blanchard 1852" docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="FFC8FFAFFF8B847DFF86FFA0FFD9B918" masterDocTitle="The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes" masterLastPageNumber="108" masterPageNumber="1" pageNumber="17" updateTime="1698662999642" updateUser="ExternalLinkService">
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<mods:mods id="1BB76275F35E41937CF746AE2A085667" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="62D6B503E30634A6DB1508E90AE650FA">
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<mods:title id="AEBB6603E2B17078F47F433C2D7B9532">The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes</mods:title>
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<mods:name id="8D1C17B8D4123EC904C2D52268001E30" type="personal">
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<mods:roleTerm id="F27E9F6888D191022F3BB24B558F315E">Author</mods:roleTerm>
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<mods:namePart id="F4F055F9017022E298F8BC5D8D960143">Matz, Jess</mods:namePart>
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<mods:name id="7D31B2A3B0B4A7961E70DF710736F6E6" type="personal">
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<mods:roleTerm id="555C3514F91E3DC1249C837BAC2EB2E0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6A7EC02472B8CF541791C5AAD0C08752">Brower, Andrew V. Z.</mods:namePart>
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<mods:typeOfResource id="95C2EB2BED99DD78327386710B0499EE">text</mods:typeOfResource>
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<mods:title id="3857B5EED325F91D5C1FBFEB6D1822E2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="821C45341D7D996C2F0865BDE50AB0CB">
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<mods:date id="5513D9724A6C5E9EF71DADC6ABFE116B">2016</mods:date>
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<mods:detail id="EC19A96795BB6493D83F7D7902735AAE" type="volume">
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<mods:number id="B0B5780F14CE2CD089B31D983DD75FEA">4125</mods:number>
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<mods:detail id="DA69585618AB0E4A93E42484B6FE0ACC" type="issue">
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<mods:number id="5C8F3F313D87596167A543DFA34815F7">1</mods:number>
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<mods:classification id="7A8CE9F932BAB9BFEE4DF70839C80379">journal article</mods:classification>
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<mods:identifier id="5197521A119C20B4F2B9C430EABA1B0C" type="CLB-Dataset">38807</mods:identifier>
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<mods:identifier id="1457E31C41D30BD6E0021084BF05A040" type="DOI">10.11646/zootaxa.4125.1.1</mods:identifier>
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<mods:identifier id="2F4D7A7D2FABC21AC7BDD7CAFD9FF3D6" type="GBIF-Dataset">1af13aec-3c54-4a92-abdf-e4e29b97e82f</mods:identifier>
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<mods:identifier id="F7866D976970B2F697F80AC4343D6361" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BA3FCF9986E60D0402B87679086C434B" type="Zenodo-Dep">271704</mods:identifier>
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<treatment id="03F187D7FF9B846DFF11FD84FB32BC32" ID-DOI="http://doi.org/10.5281/zenodo.6069999" ID-GBIF-Taxon="120828497" ID-Zenodo-Dep="6069999" LSID="urn:lsid:plazi:treatment:03F187D7FF9B846DFF11FD84FB32BC32" httpUri="http://treatment.plazi.org/id/03F187D7FF9B846DFF11FD84FB32BC32" lastPageNumber="17" pageId="16" pageNumber="17">
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<subSubSection id="C342654AFF9B846DFF11FD84FDCCBB26" box="[151,533,548,574]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8BE736C1FF9B846DFF11FD84FDCCBB26" blockId="16.[151,533,548,574]" box="[151,533,548,574]" pageId="16" pageNumber="17">
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<heading id="D0AF81ADFF9B846DFF11FD84FDCCBB26" bold="true" box="[151,533,548,574]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<taxonomicName id="4C584D42FF9B846DFF11FD84FDCCBB26" ID-CoL="922WK" authority="Blanchard, 1852" authorityName="Blanchard" authorityYear="1852" box="[151,533,548,574]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFF11FD84FDCCBB26" bold="true" box="[151,533,548,574]" pageId="16" pageNumber="17">
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<emphasis id="B92CEAD3FF9B846DFF11FD84FE99BB26" bold="true" box="[151,320,548,574]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
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<bibRefCitation id="EFC94B30FF9B846DFEC1FD84FDCCBB26" author="Blanchard" box="[327,533,548,574]" pageId="16" pageNumber="17" refString="Blanchard, C. E. (1852) Orden VI: Lepidopteros. In: Gay, C. (Ed.), Historia fisica y politica de Chile segun documentos adquiridos durante doce anos de residencia en ella y publicada bajo los auspicios del supremo Gobierno. Zoologia. C. Gay, Paris, pp. 1 - 112, pls. 1 - 7 (1854)" type="journal article" year="1852">Blanchard, 1852</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFF9B846DFF11FDCAFD16BB98" box="[151,719,618,640]" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="8BE736C1FF9B846DFF11FDCAFD16BB98" blockId="16.[151,719,618,640]" box="[151,719,618,640]" pageId="16" pageNumber="17">
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<emphasis id="B92CEAD3FF9B846DFF11FDCAFEFCBB98" bold="true" box="[151,293,618,640]" pageId="16" pageNumber="17">
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<typeStatus id="54E38863FF9B846DFF11FDCAFF15BB98" box="[151,204,618,640]" pageId="16" pageNumber="17">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C584D42FF9B846DFEADFDCAFD16BB98" authority="Blanchard, 1852" authorityName="Blanchard" authorityYear="1852" box="[299,719,618,640]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="argenteus">
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<emphasis id="B92CEAD3FF9B846DFEADFDCAFDF8BB98" box="[299,545,618,640]" italics="true" pageId="16" pageNumber="17">Argyrophorus argenteus</emphasis>
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<bibRefCitation id="EFC94B30FF9B846DFDA1FDCAFD16BB98" author="Blanchard" box="[551,719,618,640]" pageId="16" pageNumber="17" refString="Blanchard, C. E. (1852) Orden VI: Lepidopteros. In: Gay, C. (Ed.), Historia fisica y politica de Chile segun documentos adquiridos durante doce anos de residencia en ella y publicada bajo los auspicios del supremo Gobierno. Zoologia. C. Gay, Paris, pp. 1 - 112, pls. 1 - 7 (1854)" type="journal article" year="1852">Blanchard, 1852</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFF9B846DFF11FD0DFCA4BA16" pageId="16" pageNumber="17" type="diagnosis">
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<paragraph id="8BE736C1FF9B846DFF11FD0DFCA4BA16" blockId="16.[151,1437,685,1322]" pageId="16" pageNumber="17">
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<emphasis id="B92CEAD3FF9B846DFF11FD0DFED3BBDE" bold="true" box="[151,266,685,710]" pageId="16" pageNumber="17">Diagnosis</emphasis>
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. The monobasic
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<taxonomicName id="4C584D42FF9B846DFE52FD0DFDB6BBDE" box="[468,623,685,710]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFE52FD0DFDB6BBDE" box="[468,623,685,710]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
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</taxonomicName>
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as defined here is distinguished from
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<taxonomicName id="4C584D42FF9B846DFBB4FD0EFB19BBDD" box="[1074,1216,686,709]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFBB4FD0EFB19BBDD" box="[1074,1216,686,709]" italics="true" pageId="16" pageNumber="17">Punargentus</emphasis>
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</taxonomicName>
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most notably by a singly pupillated M1-M3 VFW ocellus, valvae that are are rounder and wider at the distal end than those in
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<emphasis id="B92CEAD3FF9B846DFF11FD56FEF5BA15" box="[151,300,758,781]" italics="true" pageId="16" pageNumber="17">
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<taxonomicName id="4C584D42FF9B846DFF11FD56FEF1BA15" box="[151,296,758,781]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Punargentus</taxonomicName>
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,
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</emphasis>
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a narrower saccus, and aedeagus with medial wings.
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFF9B846DFF41FCB9FB32BC32" pageId="16" pageNumber="17" type="discussion">
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<paragraph id="8BE736C1FF9B846DFF41FCB9FB32BC32" blockId="16.[151,1437,685,1322]" pageId="16" pageNumber="17">
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<emphasis id="B92CEAD3FF9B846DFF41FCB9FEEDBA2A" bold="true" box="[199,308,793,818]" pageId="16" pageNumber="17">Remarks</emphasis>
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.
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<bibRefCitation id="EFC94B30FF9B846DFECFFCBAFD5DBA2A" author="Pyrcz" box="[329,644,793,818]" pageId="16" pageNumber="17" refString="Pyrcz, T. W. & Wojtusiak, J. (2010) A new species of Argyrophorus Blanchard from northern Peru and considerations on the value of wing venation as a source of synapomorphies in some temperate Neotropical Satyrinae (Lepidoptera: Nymphalidae). Genus, 21, 605 - 613." type="journal article" year="2010">Pyrcz & Wojtusiak (2010)</bibRefCitation>
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redefined
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<taxonomicName id="4C584D42FF9B846DFC8CFCB9FC7CBA2A" box="[778,933,793,818]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFC8CFCB9FC7CBA2A" box="[778,933,793,818]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
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</taxonomicName>
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using head morphology, wing patterning, venation, wing shape, and male genitalic characters, apparently to be inclusive of
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<taxonomicName id="4C584D42FF9B846DFB0FFC9DFAC1BA4E" box="[1161,1304,829,854]" class="Insecta" family="Nymphalidae" genus="Etcheverrius" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFB0FFC9DFAC1BA4E" box="[1161,1304,829,854]" italics="true" pageId="16" pageNumber="17">Etcheverrius</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C584D42FF9B846DFAA8FC9DFA4CBA4E" box="[1326,1429,829,854]" class="Insecta" family="Nymphalidae" genus="Palmaris" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFAA8FC9DFA4CBA4E" box="[1326,1429,829,854]" italics="true" pageId="16" pageNumber="17">Palmaris</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C584D42FF9B846DFF11FCC2FEEEBA61" box="[151,311,866,889]" class="Insecta" family="Nymphalidae" genus="Pampasatyrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFF11FCC2FEEEBA61" box="[151,311,866,889]" italics="true" pageId="16" pageNumber="17">Pampasatyrus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C584D42FF9B846DFEF4FCC2FDD9BA61" box="[370,512,866,889]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFEF4FCC2FDD9BA61" box="[370,512,866,889]" italics="true" pageId="16" pageNumber="17">Punargentus</emphasis>
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</taxonomicName>
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(although they neglected to provide a list of which species are included in their circumscription of the genus, and were not clear as to whether
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<taxonomicName id="4C584D42FF9B846DFCF0FC26FB87BA85" box="[886,1118,902,925]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="poaoeneis">
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<emphasis id="B92CEAD3FF9B846DFCF0FC26FB87BA85" box="[886,1118,902,925]" italics="true" pageId="16" pageNumber="17">Pamperis poaoeneis</emphasis>
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</taxonomicName>
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is or is not included). This situation was clarified somewhat by
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<bibRefCitation id="EFC94B30FF9B846DFDBBFC09FCECBADA" box="[573,821,937,962]" pageId="16" pageNumber="17" refString="Cerdena, J. A., Pyrcz, T. W. & Zacca, T. (2014) High Andean butterflies from southern Peru, I. Dry puna Satyrinae, with the description of two new taxa and three new records from Peru (Lepidoptera: Nymphalidae). Revista Peruana de Biologia, 21, 213 - 222." type="journal article">
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Cerdeña
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<emphasis id="B92CEAD3FF9B846DFD22FC0AFD0FBADA" box="[676,726,937,962]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
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. (2014)
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</bibRefCitation>
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, who reported that Pyrcz (2010) synonymized “nine” genera under
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<taxonomicName id="4C584D42FF9B846DFEBEFC6DFE0ABAFE" box="[312,467,973,998]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFEBEFC6DFE0ABAFE" box="[312,467,973,998]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
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</taxonomicName>
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(they listed seven:
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<taxonomicName id="4C584D42FF9B846DFD46FC6DFC96BAFE" box="[704,847,973,998]" class="Insecta" family="Nymphalidae" genus="Etcheverrius" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFD46FC6DFC96BAFE" box="[704,847,973,998]" italics="true" pageId="16" pageNumber="17">Etcheverrius</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C584D42FF9B846DFCE4FC6DFC32BAFE" box="[866,1003,973,998]" class="Insecta" family="Nymphalidae" genus="Neomaniola" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFCE4FC6DFC32BAFE" box="[866,1003,973,998]" italics="true" pageId="16" pageNumber="17">Neomaniola</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C584D42FF9B846DFC7BFC6DFBBABAFE" box="[1021,1123,973,998]" class="Insecta" family="Nymphalidae" genus="Palmaris" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFC7BFC6DFBBABAFE" box="[1021,1123,973,998]" italics="true" pageId="16" pageNumber="17">Palmaris</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C584D42FF9B846DFBF1FC6EFACEBAFD" box="[1143,1303,974,997]" class="Insecta" family="Nymphalidae" genus="Pampasatyrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFBF1FC6EFACEBAFD" box="[1143,1303,974,997]" italics="true" pageId="16" pageNumber="17">Pampasatyrus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C584D42FF9B846DFAACFC6EFA4CBAFD" box="[1322,1429,974,997]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFAACFC6EFA4CBAFD" box="[1322,1429,974,997]" italics="true" pageId="16" pageNumber="17">Pamperis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C584D42FF9B846DFF11FC52FEFCBD11" box="[151,293,1010,1033]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFF11FC52FEFCBD11" box="[151,293,1010,1033]" italics="true" pageId="16" pageNumber="17">Punargentus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C584D42FF9B846DFEDDFC51FDDBBD12" box="[347,514,1009,1034]" class="Insecta" family="Nymphalidae" genus="Stuardosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FF9B846DFEDDFC51FDDBBD12" box="[347,514,1009,1034]" italics="true" pageId="16" pageNumber="17">Stuardosatyrus</emphasis>
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</taxonomicName>
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). Subsequently,
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<bibRefCitation id="EFC94B30FF9B846DFD3BFC52FC97BD12" author="Pyrcz" box="[701,846,1010,1034]" pageId="16" pageNumber="17" refString="Pyrcz, T. W. (2012) A new species of satyrine butterfly from Patagonia in more than a century and revisional notes on the genus Faunula C. Felder & R. Felder (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa, 3342, 60 - 68." type="journal article" year="2012">Pyrcz (2012)</bibRefCitation>
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moved
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<taxonomicName id="4C584D42FF9B846DFC2EFC51FB79BD12" box="[936,1184,1009,1034]" class="Insecta" family="Nymphalidae" genus="Neomaniola" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="euripides">
|
|
<emphasis id="B92CEAD3FF9B846DFC2EFC51FB79BD12" box="[936,1184,1009,1034]" italics="true" pageId="16" pageNumber="17">Neomaniola euripides</emphasis>
|
|
</taxonomicName>
|
|
from
|
|
<taxonomicName id="4C584D42FF9B846DFB62FC51FAA6BD12" box="[1252,1407,1009,1034]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFB62FC51FAA6BD12" box="[1252,1407,1009,1034]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
|
|
</taxonomicName>
|
|
to
|
|
<taxonomicName id="4C584D42FF9B846DFF11FBB5FF2FBD36" box="[151,246,1045,1070]" class="Insecta" family="Nymphalidae" genus="Faunula" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFF11FBB5FF2FBD36" box="[151,246,1045,1070]" italics="true" pageId="16" pageNumber="17">Faunula</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<bibRefCitation id="EFC94B30FF9B846DFE82FBB5FE2ABD36" box="[260,499,1045,1070]" pageId="16" pageNumber="17" refString="Cerdena, J. A., Pyrcz, T. W. & Zacca, T. (2014) High Andean butterflies from southern Peru, I. Dry puna Satyrinae, with the description of two new taxa and three new records from Peru (Lepidoptera: Nymphalidae). Revista Peruana de Biologia, 21, 213 - 222." type="journal article">
|
|
Cerdeña
|
|
<emphasis id="B92CEAD3FF9B846DFEEFFBB6FE43BD36" box="[361,410,1045,1070]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
|
. (2014)
|
|
</bibRefCitation>
|
|
also noted that
|
|
<taxonomicName id="4C584D42FF9B846DFD23FBB6FC9CBD35" box="[677,837,1046,1069]" class="Insecta" family="Nymphalidae" genus="Pampasatyrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFD23FBB6FC9CBD35" box="[677,837,1046,1069]" italics="true" pageId="16" pageNumber="17">Pampasatyrus</emphasis>
|
|
</taxonomicName>
|
|
is closely-related to, but distinct from
|
|
<emphasis id="B92CEAD3FF9B846DFB7CFBB5FA44BD36" box="[1274,1437,1045,1070]" italics="true" pageId="16" pageNumber="17">
|
|
<taxonomicName id="4C584D42FF9B846DFB7CFBB5FA40BD36" box="[1274,1433,1045,1070]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Argyrophorus</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
They reported the latter to contain 11 species—presumably the species previously included by
|
|
<bibRefCitation id="EFC94B30FF9B846DFB5DFB9AFF38BD6E" author="Lamas" pageId="16" pageNumber="17" refString="Lamas, G. & Viloria, A. (2004) Subtribes Erebiina and Hypocystina. In: Lamas, G. (Ed) Checklist: Part 4 A. Hesperioidea- Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Association for Tropical Lepidoptera, Inc. / Scientific Publishers, Gainesville, FL, pp. 215 - 217." type="book chapter" year="2004">Lamas & Viloria (2004)</bibRefCitation>
|
|
in
|
|
<taxonomicName id="4C584D42FF9B846DFE8FFBFDFE7DBD6E" box="[265,420,1117,1142]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFE8FFBFDFE7DBD6E" box="[265,420,1117,1142]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
|
|
</taxonomicName>
|
|
(1 + 1 subsequently described by
|
|
<bibRefCitation id="EFC94B30FF9B846DFCB4FBFEFB8EBD6E" author="Pyrcz" box="[818,1111,1117,1142]" pageId="16" pageNumber="17" refString="Pyrcz, T. W. & Wojtusiak, J. (2010) A new species of Argyrophorus Blanchard from northern Peru and considerations on the value of wing venation as a source of synapomorphies in some temperate Neotropical Satyrinae (Lepidoptera: Nymphalidae). Genus, 21, 605 - 613." type="journal article" year="2010">Pyrcz & Wojtusiak, 2010</bibRefCitation>
|
|
),
|
|
<taxonomicName id="4C584D42FF9B846DFBEAFBFDFB22BD6E" box="[1132,1275,1117,1142]" class="Insecta" family="Nymphalidae" genus="Etcheverrius" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFBEAFBFDFB22BD6E" box="[1132,1275,1117,1142]" italics="true" pageId="16" pageNumber="17">Etcheverrius</emphasis>
|
|
</taxonomicName>
|
|
(2),
|
|
<taxonomicName id="4C584D42FF9B846DFAB3FBFDFA45BD6E" box="[1333,1436,1117,1142]" class="Insecta" family="Nymphalidae" genus="Palmaris" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFAB3FBFDFA45BD6E" box="[1333,1436,1117,1142]" italics="true" pageId="16" pageNumber="17">Palmaris</emphasis>
|
|
</taxonomicName>
|
|
(4),
|
|
<taxonomicName id="4C584D42FF9B846DFF40FB22FEEBBD81" box="[198,306,1154,1177]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFF40FB22FEEBBD81" box="[198,306,1154,1177]" italics="true" pageId="16" pageNumber="17">Pamperis</emphasis>
|
|
</taxonomicName>
|
|
(1) and
|
|
<taxonomicName id="4C584D42FF9B846DFE15FB22FDF8BD81" box="[403,545,1154,1177]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFE15FB22FDF8BD81" box="[403,545,1154,1177]" italics="true" pageId="16" pageNumber="17">Punargentus</emphasis>
|
|
</taxonomicName>
|
|
(2). According to our cladogram (
|
|
<figureCitation id="13632A44FF9B846DFC2EFB21FC33BD82" box="[936,1002,1153,1178]" captionStart="FIGURE 1" captionStartId="78.[151,250,1782,1804]" captionTargetBox="[187,1390,212,1741]" captionTargetId="figure@78.[180,1408,195,1758]" captionTargetPageId="78" captionText="FIGURE 1. One of six equally parsimonious trees for Neosatyriti and selected pronophiline outgroups, based on 4071 bp of concatenated DNA sequence from mtDNA COI-COII, elongation factor 1 - alpha and wingless. Length = 8955 steps, CIx = 0.2435; RI = 0.4729." httpUri="https://zenodo.org/record/271705/files/figure.png" pageId="16" pageNumber="17">Fig. 1</figureCitation>
|
|
), that circumscription is polyphyletic:
|
|
<taxonomicName id="4C584D42FF9B846DFF11FB05FEEBBDA6" box="[151,306,1189,1214]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFF11FB05FEEBBDA6" box="[151,306,1189,1214]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C584D42FF9B846DFEEDFB06FE0FBDA5" box="[363,470,1190,1213]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFEEDFB06FE0FBDA5" box="[363,470,1190,1213]" italics="true" pageId="16" pageNumber="17">Pamperis</emphasis>
|
|
</taxonomicName>
|
|
are phylogenetically distinct from
|
|
<taxonomicName id="4C584D42FF9B846DFCE0FB06FC2DBDA5" box="[870,1012,1190,1213]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFCE0FB06FC2DBDA5" box="[870,1012,1190,1213]" italics="true" pageId="16" pageNumber="17">Punargentus</emphasis>
|
|
</taxonomicName>
|
|
(here circumscribed more broadly to include
|
|
<taxonomicName id="4C584D42FF9B846DFF76FB69FEA6BDFA" box="[240,383,1225,1250]" class="Insecta" family="Nymphalidae" genus="Etcheverrius" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFF76FB69FEA6BDFA" box="[240,383,1225,1250]" italics="true" pageId="16" pageNumber="17">Etcheverrius</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C584D42FF9B846DFE32FB69FDC2BDFA" box="[436,539,1225,1250]" class="Insecta" family="Nymphalidae" genus="Palmaris" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFE32FB69FDC2BDFA" box="[436,539,1225,1250]" italics="true" pageId="16" pageNumber="17">Palmaris</emphasis>
|
|
</taxonomicName>
|
|
) and its sister genus,
|
|
<taxonomicName id="4C584D42FF9B846DFC8DFB6AFC72BDF9" box="[779,939,1226,1249]" class="Insecta" family="Nymphalidae" genus="Pampasatyrus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFC8DFB6AFC72BDF9" box="[779,939,1226,1249]" italics="true" pageId="16" pageNumber="17">Pampasatyrus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<bibRefCitation id="EFC94B30FF9B846DFC3FFB6AFB07BDFA" author="Pyrcz" box="[953,1246,1225,1250]" pageId="16" pageNumber="17" refString="Pyrcz, T. W. & Wojtusiak, J. (2010) A new species of Argyrophorus Blanchard from northern Peru and considerations on the value of wing venation as a source of synapomorphies in some temperate Neotropical Satyrinae (Lepidoptera: Nymphalidae). Genus, 21, 605 - 613." type="journal article" year="2010">Pyrcz & Wojtusiak (2010)</bibRefCitation>
|
|
placed their new species
|
|
<taxonomicName id="4C584D42FF9B846DFF76FB4DFEB0BC1E" box="[240,361,1261,1286]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="blanchardi">
|
|
<emphasis id="B92CEAD3FF9B846DFF76FB4DFEB0BC1E" box="[240,361,1261,1286]" italics="true" pageId="16" pageNumber="17">blanchardi</emphasis>
|
|
</taxonomicName>
|
|
in
|
|
<taxonomicName id="4C584D42FF9B846DFE16FB4DFDF2BC1E" box="[400,555,1261,1286]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B92CEAD3FF9B846DFE16FB4DFDF2BC1E" box="[400,555,1261,1286]" italics="true" pageId="16" pageNumber="17">Argyrophorus</emphasis>
|
|
</taxonomicName>
|
|
using their broader definition, but the similarities in the VFW M1-M3 ocelli, and the lack of medial appendages on the aedeagus suggest that it belongs in
|
|
<emphasis id="B92CEAD3FF9B846DFC71FAB2FB55BC31" box="[1015,1164,1298,1321]" italics="true" pageId="16" pageNumber="17">
|
|
<taxonomicName id="4C584D42FF9B846DFC71FAB2FB51BC31" box="[1015,1160,1298,1321]" class="Insecta" family="Nymphalidae" genus="Punargentus" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Punargentus</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
instead.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |