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<document id="B2C5525462169E77FB752C6E5A5AF54C" ID-CLB-Dataset="46677" ID-DOI="10.5281/zenodo.207506" ID-GBIF-Dataset="dcce60fc-6550-4824-8dbb-b16bf1545895" ID-ISSN="1175-5326" ID-Zenodo-Dep="207506" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460368299503" checkinUser="plazi" docAuthor="Topkara, Tarik, Jäch, Manfred &amp; Lu, Ahmet Kasapo Ğ" docDate="2011" docId="03F187D6155D8D31FF35FBEE9476F842" docLanguage="en" docName="zt02913p062.pdf" docOrigin="Zootaxa 2913" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochthebius ustaoglui Topkara, Jäch &amp; Lu, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="60" masterDocId="FFC8FFAE155D8D30FFA2FFDD971AFF96" masterDocTitle="Ochthebius ustaoglui sp. nov. (Coleoptera: Hydraenidae), a new species of the O. metallescens group from Turkey" masterLastPageNumber="62" masterPageNumber="59" pageNumber="59" updateTime="1698295445416" updateUser="ExternalLinkService">
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<mods:title id="87626A2D47A47F76254D07E0EF0CE331">Ochthebius ustaoglui sp. nov. (Coleoptera: Hydraenidae), a new species of the O. metallescens group from Turkey</mods:title>
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<mods:namePart id="0E4A3AC6842F55E6DFC42C86EC5BC277">Topkara, Tarik</mods:namePart>
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<mods:namePart id="E24EFDAB140287DB074C905D891FAEB8">Jäch, Manfred</mods:namePart>
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<mods:namePart id="69D31304B396B7D3C244CF467AC59B70">Lu, Ahmet Kasapo Ğ</mods:namePart>
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<mods:date id="78D6CD00C06D82C808EEEE8FD5F2CE89">2011</mods:date>
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<treatment id="03F187D6155D8D31FF35FBEE9476F842" ID-DOI="http://doi.org/10.5281/zenodo.6189346" ID-GBIF-Taxon="119411226" ID-Zenodo-Dep="6189346" LSID="urn:lsid:plazi:treatment:03F187D6155D8D31FF35FBEE9476F842" httpUri="http://treatment.plazi.org/id/03F187D6155D8D31FF35FBEE9476F842" lastPageId="1" lastPageNumber="60" pageId="0" pageNumber="59">
<subSubSection id="C342654B155D8D30FF35FBEE96C4FBDD" box="[151,478,1075,1099]" pageId="0" pageNumber="59" type="nomenclature">
<paragraph id="8BE736C0155D8D30FF35FBEE96C4FBDD" blockId="0.[151,478,1075,1099]" box="[151,478,1075,1099]" pageId="0" pageNumber="59">
<heading id="D0AF81AC155D8D30FF35FBEE96C4FBDD" bold="true" box="[151,478,1075,1099]" fontSize="10" level="2" pageId="0" pageNumber="59" reason="2">
<emphasis id="B92CEAD2155D8D30FF35FBEE96C4FBDD" bold="true" box="[151,478,1075,1099]" pageId="0" pageNumber="59">
<taxonomicName id="4C584D43155D8D30FF35FBEE969BFBDD" ID-CoL="48FCY" box="[151,385,1075,1099]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="ustaoglui" status="sp. nov.">
<emphasis id="B92CEAD2155D8D30FF35FBEE969BFBDD" bold="true" box="[151,385,1075,1099]" italics="true" pageId="0" pageNumber="59">Ochthebius ustaoglui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F57A9155D8D30FE2AFBEE96C4FBDD" box="[392,478,1075,1099]" pageId="0" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C342654B155D8D30FF35FBA49539FA8D" pageId="0" pageNumber="59" type="materials_examined">
<paragraph id="8BE736C0155D8D30FF35FBA496C4FB25" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
<emphasis id="B92CEAD2155D8D30FF35FBA4962BFB07" bold="true" box="[151,305,1145,1169]" pageId="0" pageNumber="59">
<typeStatus id="54E38862155D8D30FF35FBA497D5FB07" box="[151,207,1145,1169]" pageId="0" pageNumber="59">Type</typeStatus>
locality:
</emphasis>
Small stream, ca.
<quantity id="4CA09B25155D8D30FE57FBA4952DFB06" box="[501,567,1145,1168]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="59" unit="cm" value="50.0">50 cm</quantity>
wide,
<geoCoordinate id="EE6C5007155D8D30FDDFFBA495E0FB06" box="[637,762,1145,1168]" direction="north" orientation="latitude" pageId="0" pageNumber="59" precision="15" value="40.58583">40°35'09''N</geoCoordinate>
<geoCoordinate id="EE6C5007155D8D30FCA2FBA49461FB06" box="[768,891,1145,1168]" direction="east" orientation="longitude" pageId="0" pageNumber="59" precision="15" value="40.863052">40°51'47''E</geoCoordinate>
, ca.
<quantity id="4CA09B25155D8D30FC09FBA494E1FB06" box="[939,1019,1145,1168]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="0" pageNumber="59" unit="m" value="2200.0">2200 m</quantity>
a.s.l., Ovit Pass, ca.
<quantity id="4CA09B25155D8D30FB74FBA49203FB06" box="[1238,1305,1145,1168]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="59" unit="km" value="30.0">30 km</quantity>
NNW İspir, Erzurum, northeastern
<collectingCountry id="F34F7650155D8D30FE2EFB4196C0FB25" box="[396,474,1180,1203]" name="Turkey" pageId="0" pageNumber="59">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="8BE736C0155D8D30FF65FB639539FA8D" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
<emphasis id="B92CEAD2155D8D30FF65FB6396B6FB40" bold="true" box="[199,428,1214,1238]" pageId="0" pageNumber="59">Material examined.</emphasis>
<typeStatus id="54E38862155D8D30FE1BFB639506FB43" box="[441,540,1214,1237]" pageId="0" pageNumber="59" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED49AE05155D8D30FDD1FB6395A4FB43" box="[627,702,1214,1237]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="59">NMW</collectionCode>
): “
<collectingCountry id="F34F7650155D8D30FD43FB639412FB43" box="[737,776,1214,1237]" name="Chile" pageId="0" pageNumber="59">TR</collectingCountry>
: ERZURUM Ovit Pass \ leg. Kasapoğlu 0
<date id="FFE61000155D8D30FB5EFB639294FB43" box="[1276,1422,1214,1237]" pageId="0" pageNumber="59" value="2000-07-03">3. VII. 2000</date>
”.
<typeStatus id="54E38862155D8D30FF35FB3C9619FB6E" box="[151,259,1249,1272]" pageId="0" pageNumber="59" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5EFD49155D8D30FEAEFB3C967BFB6E" box="[268,353,1249,1272]" pageId="0" pageNumber="59" type="male">5 males</specimenCount>
,
<specimenCount id="9D5EFD49155D8D30FEC9FB3C96C8FB6E" box="[363,466,1249,1272]" pageId="0" pageNumber="59" type="female">6 females</specimenCount>
(
<collectionCode id="ED49AE05155D8D30FE42FB3C9530FB6E" box="[480,554,1249,1272]" httpUri="http://grbio.org/cool/vqgc-v6vf" name="Museum of Faculty of Fisheries, Ege University" pageId="0" pageNumber="59">ESFM</collectionCode>
,
<collectionCode id="ED49AE05155D8D30FD96FB3C9564FB6E" box="[564,638,1249,1272]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="0" pageNumber="59">NMW</collectionCode>
): “
<collectingCountry id="F34F7650155D8D30FD3AFB3C9412FB6E" box="[664,776,1249,1272]" name="Turkey" pageId="0" pageNumber="59">TURKEY</collectingCountry>
: Prov. Denizli ca.
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SW of Beyağaç Lake Kartal,
<quantity id="4CA09B25155D8D30FAEEFB3C9286FB6E" box="[1356,1436,1249,1272]" metricMagnitude="3" metricUnit="m" metricValue="1.903" pageId="0" pageNumber="59" unit="m" value="1903.0">1903 m</quantity>
a.s.l.
<date id="FFE61000155D8D30FF6DFAD9964BFA8D" box="[207,337,1284,1307]" pageId="0" pageNumber="59" value="2010-07-16">16.VII.2010</date>
leg. E.T. Topkara”.
</paragraph>
</subSubSection>
<subSubSection id="C342654B155D8D30FF65FAFB929BF81D" pageId="0" pageNumber="59" type="description">
<paragraph id="8BE736C0155D8D30FF65FAFB9401FA15" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
<emphasis id="B92CEAD2155D8D30FF65FAFB9648FAA8" bold="true" box="[199,338,1318,1342]" pageId="0" pageNumber="59">Description.</emphasis>
<quantity id="4CA09B25155D8D30FEF8FAFB96ECFAAB" box="[346,502,1318,1341]" metricMagnitude="-3" metricUnit="m" metricValue="1.925" metricValueMax="2.1" metricValueMin="1.75" pageId="0" pageNumber="59" unit="mm" value="1.925" valueMax="2.1" valueMin="1.75">1.752.10 mm</quantity>
long. Black, legs sometimes slightly paler dark brown. Anterior corners of pronotum acute, postocular excision and postocular tooth usually well developed; lateral margins of pronotal ears straight, slightly convergent anteriad. Metaventrite posteromedially glabrous.
</paragraph>
<paragraph id="8BE736C0155D8D30FF65FA5393B1FA7D" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
<emphasis id="B92CEAD2155D8D30FF65FA5396B9FA30" bold="true" box="[199,419,1422,1446]" pageId="0" pageNumber="59">Sexual dimophism:</emphasis>
Weakly pronounced. Explanate margin of elytra of females very slightly wider than in males, tips of elytra hardly noticeably more acuminate and more conjointly rounded. The shapes of labrum and protarsi are similar in both sexes. Posterior margin of terminal abdominal tergite of females with blunt spines.
</paragraph>
<paragraph id="8BE736C0155D8D30FF65FA2B9510F9C5" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
<emphasis id="B92CEAD2155D8D30FF65FA2B965CF998" bold="true" box="[199,326,1526,1550]" pageId="0" pageNumber="59">Variability.</emphasis>
Postocular emargination of pronotum and postocular tooth weakly or well developed. Pronotal disc moderately to densely punctate, glabrous. Length/width ratio of main part of aedeagal distal lobe varies considerably, from 1.73 to 2.16 (
<typeStatus id="54E38862155D8D30FEFDF9E196D8F9C5" box="[351,450,1596,1619]" pageId="0" pageNumber="59" type="holotype">holotype</typeStatus>
: 1.86).
</paragraph>
<paragraph id="8BE736C0155D8D30FF65F983951BF92D" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
<emphasis id="B92CEAD2155D8D30FF65F98396DBF9E0" bold="true" box="[199,449,1630,1654]" pageId="0" pageNumber="59">Differential diagnosis.</emphasis>
Due to the high variability, we have not been able to detect significant external characters to distinguish the new species from
<taxonomicName id="4C584D43155D8D30FE5FF95C9407F90E" box="[509,797,1665,1688]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="hofratvukovitsi">
<emphasis id="B92CEAD2155D8D30FE5FF95C9407F90E" box="[509,797,1665,1688]" italics="true" pageId="0" pageNumber="59">Ochthebius hofratvukovitsi</emphasis>
</taxonomicName>
. The new species is, however, sufficiently characterized by its unusual aedeagus morphology.
</paragraph>
<paragraph id="8BE736C0155D8D30FF65F91B929BF81D" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
Aedeagus (
<figureCitation id="13632A45155D8D30FEE3F91B9698F94B" box="[321,386,1734,1757]" captionStart="FIGURE 1" captionStartId="1.[151,250,1499,1522]" captionTargetBox="[151,1436,195,1475]" captionTargetId="figure@1.[151,1436,195,1475]" captionTargetPageId="1" captionText="FIGURE 1. Ochthebius ustaoglui sp. nov., lateral view of the aedeagus, paratype (from Kartal Lake)." httpUri="https://zenodo.org/record/207507/files/figure.png" pageId="0" pageNumber="59">Fig. 1</figureCitation>
): Main piece ca. 320360 µm long, almost evenly curved (lateral view), slightly attenuate near middle, apex obliquely truncate. Distal lobe very conspicuously shaped: main part moderately wide, subcylindrical, subrectangular, about twice as long as wide, baso-ventrally and apically widely rounded; ventral margin at about apical third with a somewhat ax blade-shaped appendage, more or less compressed laterally, its distal apex widely rounded, basal apex acuminate and directed towards tip of main piece. Parameres more or less symmetrical, right one slightly longer than left one, close to main piece, inserted ventrally near basal 0.3 of main piece; apices with a few short setae.
</paragraph>
</subSubSection>
<subSubSection id="C342654B155D8D31FF65F84B9468F8DF" lastPageId="1" lastPageNumber="60" pageId="0" pageNumber="59" type="discussion">
<paragraph id="8BE736C0155D8D30FF65F84B9259F865" blockId="0.[151,1437,1141,2035]" pageId="0" pageNumber="59">
<emphasis id="B92CEAD2155D8D30FF65F84B965CF838" bold="true" box="[199,326,1942,1966]" pageId="0" pageNumber="59">Discussion.</emphasis>
Despite the considerable linear distance between the two populations of about
<quantity id="4CA09B25155D8D30FB00F84B921EF83B" box="[1186,1284,1942,1965]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="59" unit="km" value="1000.0">1000 km</quantity>
, we could not detect significant aedeagal differences. However, examination of more material and molecular data will be necessary to bring forth conclusive evidence that the populations from Denizli and Erzurum really belong to a single species.
</paragraph>
<caption id="DF276648155C8D31FF35FA0693BBFA67" box="[151,1185,1499,1522]" httpUri="https://zenodo.org/record/207507/files/figure.png" pageId="1" pageNumber="60" targetBox="[151,1436,195,1475]" targetPageId="1">
<paragraph id="8BE736C0155C8D31FF35FA0693BBFA67" blockId="1.[151,1185,1499,1522]" box="[151,1185,1499,1522]" pageId="1" pageNumber="60">
<emphasis id="B92CEAD2155C8D31FF35FA069609FA67" bold="true" box="[151,275,1499,1522]" pageId="1" pageNumber="60">FIGURE 1.</emphasis>
<taxonomicName id="4C584D43155C8D31FEBBFA0196EBFA64" box="[281,497,1500,1522]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="ustaoglui" status="sp. nov.">
<emphasis id="B92CEAD2155C8D31FEBBFA0196EBFA64" box="[281,497,1500,1522]" italics="true" pageId="1" pageNumber="60">Ochthebius ustaoglui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F57A9155C8D31FE55FA019552FA67" box="[503,584,1500,1521]" pageId="1" pageNumber="60" rank="species">
<emphasis id="B92CEAD2155C8D31FE55FA019552FA67" bold="true" box="[503,584,1500,1521]" pageId="1" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
, lateral view of the aedeagus, paratype (from Kartal Lake).
</paragraph>
</caption>
<paragraph id="8BE736C0155C8D31FF65F9C0942BF929" blockId="1.[151,1437,1565,2004]" pageId="1" pageNumber="60">
Based on external similarities, the new species is obviously related to
<taxonomicName id="4C584D43155C8D31FC1AF9C093C3F9A2" box="[952,1241,1565,1588]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="hofratvukovitsi">
<emphasis id="B92CEAD2155C8D31FC1AF9C093C3F9A2" box="[952,1241,1565,1588]" italics="true" pageId="1" pageNumber="60">Ochthebius hofratvukovitsi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C584D43155C8D31FB44F9C09609F9C1" authority="Ferro, 1982" authorityName="Ferro" authorityYear="1982" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="khuzestanicus">
<emphasis id="B92CEAD2155C8D31FB44F9C09281F9A2" box="[1254,1435,1565,1588]" italics="true" pageId="1" pageNumber="60">
<collectingCountry id="F34F7650155C8D31FB44F9C093E6F9A2" box="[1254,1276,1565,1588]" name="Iceland" pageId="1" pageNumber="60">O</collectingCountry>
. khuzestanicus
</emphasis>
Ferro, 1982
</taxonomicName>
and
<taxonomicName id="4C584D43155C8D31FEE5F99D9579F9C1" authority="Reitter, 1885" authorityName="Reitter" authorityYear="1885" box="[327,611,1600,1623]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="puberulus">
<emphasis id="B92CEAD2155C8D31FEE5F99D96C8F9C1" box="[327,466,1600,1623]" italics="true" pageId="1" pageNumber="60">
<collectingCountry id="F34F7650155C8D31FEE5F99D9647F9C1" box="[327,349,1600,1623]" name="Iceland" pageId="1" pageNumber="60">O</collectingCountry>
. puberulus
</emphasis>
Reitter, 1885
</taxonomicName>
(see
<bibRefCitation id="EFC94B31155C8D31FD39F99D9413F9C1" author="Jach" box="[667,777,1600,1623]" pageId="1" pageNumber="60" refString="Jach, M. A. (1989) Revision of the Palearctic species of the genus Ochthebius Leach III. The metallescens - group (Hydraenidae, Coleoptera), Linzer Biologische Beitrage, 21, 351 - 390." type="journal article" year="1989">Jäch 1989</bibRefCitation>
,
<bibRefCitation id="EFC94B31155C8D31FCB1F99D9456F9C1" author="Jach" box="[787,844,1600,1623]" pageId="1" pageNumber="60" refString="Jach, M. A. (1994) Revision der palaarktischen Arten der Gattung Ochthebius Leach. XIII. Beschreibung neuer Arten aus Spanien, der Turkei, Rußland und China (Insecta: Coleoptera: Hydraenidae), Annalen des Naturhistorischen Museums Wien, B, 96, 199 - 208." type="journal article" year="1994">1994</bibRefCitation>
). No other species of this genus has an ax blade-shaped aedeagal apex. Externally,
<taxonomicName id="4C584D43155C8D31FE63F9BF9560F9EF" box="[449,634,1634,1657]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="khuzestanicus">
<emphasis id="B92CEAD2155C8D31FE63F9BF9560F9EF" box="[449,634,1634,1657]" italics="true" pageId="1" pageNumber="60">
<collectingCountry id="F34F7650155C8D31FE63F9BF96CDF9EF" box="[449,471,1634,1657]" name="Iceland" pageId="1" pageNumber="60">O</collectingCountry>
. khuzestanicus
</emphasis>
</taxonomicName>
deviates from the other three species in the presence of distinct elytral granules. Due to aedeagal similarities,
<taxonomicName id="4C584D43155C8D31FD9DF95894A2F90A" authority="Jach, 1989" authorityName="Jach" authorityYear="1989" box="[575,952,1669,1692]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="serpentinus">
<emphasis id="B92CEAD2155C8D31FD9DF9589426F90A" box="[575,828,1669,1692]" italics="true" pageId="1" pageNumber="60">Ochthebius serpentinus</emphasis>
<bibRefCitation id="EFC94B31155C8D31FCE6F95894A2F90A" author="Jach" box="[836,952,1669,1692]" pageId="1" pageNumber="60" refString="Jach, M. A. (1989) Revision of the Palearctic species of the genus Ochthebius Leach III. The metallescens - group (Hydraenidae, Coleoptera), Linzer Biologische Beitrage, 21, 351 - 390." type="journal article" year="1989">Jäch, 1989</bibRefCitation>
</taxonomicName>
might also belong to this cluster of species. However, its elytra are usually more ovoid and more glabrous.
</paragraph>
<paragraph id="8BE736C0155C8D31FF65F9179468F8DF" blockId="1.[151,1437,1565,2004]" pageId="1" pageNumber="60">
<emphasis id="B92CEAD2155C8D31FF65F9179673F974" bold="true" box="[199,361,1738,1762]" pageId="1" pageNumber="60">Habitat notes.</emphasis>
The
<typeStatus id="54E38862155C8D31FE06F917951BF977" box="[420,513,1738,1761]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
was collected from a small stream with cold and clear water, whereas the
<typeStatus id="54E38862155C8D31FA94F9179286F977" box="[1334,1436,1738,1761]" pageId="1" pageNumber="60" type="paratype">paratypes</typeStatus>
were sampled in small ponds, formed by melting snow near a lake shore. However, near the
<typeStatus id="54E38862155C8D31FBD2F9309384F892" box="[1136,1182,1773,1796]" pageId="1" pageNumber="60">type</typeStatus>
locality there also were some small pools, which may in fact represent the original habitat of the population. Further field work will be necessary to reveal the habitat preferences of
<taxonomicName id="4C584D43155C8D31FDADF8EF95E8F8DF" box="[527,754,1842,1865]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="ustaoglui">
<emphasis id="B92CEAD2155C8D31FDADF8EF95E8F8DF" box="[527,754,1842,1865]" italics="true" pageId="1" pageNumber="60">Ochthebius ustaoglui</emphasis>
</taxonomicName>
(see fig. 2).
</paragraph>
</subSubSection>
<subSubSection id="C342654B155C8D31FF65F889961DF819" pageId="1" pageNumber="60" type="etymology">
<paragraph id="8BE736C0155C8D31FF65F889961DF819" blockId="1.[151,1437,1565,2004]" pageId="1" pageNumber="60">
<emphasis id="B92CEAD2155C8D31FF65F889965DF8FA" bold="true" box="[199,327,1876,1900]" pageId="1" pageNumber="60">Etymology.</emphasis>
The new species is named in honour of Prof. Dr. M. Ruşen Ustaoğlu (Ege University, Faculty of Fisheries).
</paragraph>
</subSubSection>
<subSubSection id="C342654B155C8D31FF65F8479476F842" pageId="1" pageNumber="60" type="distribution">
<paragraph id="8BE736C0155C8D31FF65F8479476F842" blockId="1.[151,1437,1565,2004]" pageId="1" pageNumber="60">
<emphasis id="B92CEAD2155C8D31FF65F8479643F824" bold="true" box="[199,345,1946,1970]" pageId="1" pageNumber="60">Distribution.</emphasis>
The new species is only known from southwestern
<collectingCountry id="F34F7650155C8D31FC07F84794EBF827" box="[933,1009,1946,1969]" name="Turkey" pageId="1" pageNumber="60">Turkey</collectingCountry>
(Denizli,
<geoCoordinate id="EE6C5007155C8D31FBC6F84793C5F827" box="[1124,1247,1946,1969]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="15" value="37.09611">37º05'46&quot;N</geoCoordinate>
<geoCoordinate id="EE6C5007155C8D31FB4BF8479279F827" box="[1257,1379,1946,1969]" direction="east" orientation="longitude" pageId="1" pageNumber="60" precision="15" value="28.850555">28º51'02&quot;E</geoCoordinate>
) and northeastern
<collectingCountry id="F34F7650155C8D31FE83F8609677F842" box="[289,365,1981,2004]" name="Turkey" pageId="1" pageNumber="60">Turkey</collectingCountry>
(Erzurum,
<geoCoordinate id="EE6C5007155C8D31FE4AF860957EF842" box="[488,612,1981,2004]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="15" value="40.58583">40°35'09''N</geoCoordinate>
<geoCoordinate id="EE6C5007155C8D31FDC8F86095FFF842" box="[618,741,1981,2004]" direction="east" orientation="longitude" pageId="1" pageNumber="60" precision="15" value="40.863052">40°51'47''E</geoCoordinate>
) (see fig. 3).
</paragraph>
</subSubSection>
</treatment>
</document>