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<document id="E66B06CB09DC7F8102AE90DECC3BC786" ID-DOI="10.11646/zootaxa.2851.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5286934" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629721501414" checkinUser="felipe" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2011" docId="03BE8797007DFF8498B1FB0FFC16C438" docLanguage="en" docName="zt02851p075.pdf" docOrigin="Zootaxa 2851 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mortoniella alicula Blahnik &amp; Holzenthal 2011, new species" docType="treatment" docVersion="5" lastPageNumber="49" masterDocId="FF87FFEF0053FFB49826FF89FFBAC62F" masterDocTitle="Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851" masterLastPageNumber="75" masterPageNumber="1" pageNumber="47" updateTime="1699101759394" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="84E2564A50535DCC6A9AA55A20D89158">Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851</mods:title>
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<mods:namePart id="1BB8D016A9C06E80E08BD0F2D9EC0A75">Blahnik, Roger J.</mods:namePart>
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<mods:namePart id="D0B9E9A20A14632161743E144D8AE69D">Holzenthal, Ralph W.</mods:namePart>
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<mods:title id="26E44099AC4F3D0DFCEB86C73E5000D4">Zootaxa</mods:title>
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<mods:part id="E4B7FF0A9DD3E983A255D90F7D0AEAA9">
<mods:date id="6A2A7B6D879105121BDE11FB6A8BB13F">2011</mods:date>
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<mods:number id="CAAF01E0871BC2FAA894627321441065">2011-04-29</mods:number>
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<mods:detail id="D4192216D8F9DB18981ABDF352AC2C1C" type="volume">
<mods:number id="FFEC438AC69BF7D51B84FF727E24FBDD">2851</mods:number>
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<mods:number id="DF95EADFD81801E3E6462A547266279E">1</mods:number>
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<mods:start id="109BC41BB49099805518978E503A944A">1</mods:start>
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<mods:url id="6DCD0A2DE1053D987D79CCE21BFB579C">https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1</mods:url>
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<mods:classification id="BBB223C609DC865DB9D2616957E75D01">journal article</mods:classification>
<mods:identifier id="FA7C752399DF4E40C2D18912E083816C" type="DOI">10.11646/zootaxa.2851.1.1</mods:identifier>
<mods:identifier id="EECC555C117EAF88CA9631D9490E66C9" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="A92AEBFD66F982B4A9DD4422A85DBEB3" type="Zenodo-Dep">5286934</mods:identifier>
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<treatment id="03BE8797007DFF8498B1FB0FFC16C438" ID-DOI="http://doi.org/10.5281/zenodo.5293498" ID-Zenodo-Dep="5293498" LSID="urn:lsid:plazi:treatment:03BE8797007DFF8498B1FB0FFC16C438" httpUri="http://treatment.plazi.org/id/03BE8797007DFF8498B1FB0FFC16C438" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="C30D650A007DFF9A98B1FB0FFDA5C28F" box="[151,543,1158,1184]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BA83681007DFF9A98B1FB0FFDA5C28F" blockId="46.[151,543,1158,1217]" box="[151,543,1158,1184]" pageId="46" pageNumber="47">
<heading id="D0E081ED007DFF9A98B1FB0FFDA5C28F" bold="true" box="[151,543,1158,1184]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="B963EA93007DFF9A98B1FB0FFDA5C28F" bold="true" box="[151,543,1158,1184]" pageId="46" pageNumber="47">
<taxonomicName id="4C174D02007DFF9A98B1FB0FFE3BC28F" authority="Blahnik &amp; Holzenthal, 2011" authorityName="Blahnik &amp; Holzenthal" authorityYear="2011" box="[151,385,1158,1184]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="alicula" status="sp. nov.">
<emphasis id="B963EA93007DFF9A98B1FB0FFE3BC28F" bold="true" box="[151,385,1158,1184]" italics="true" pageId="46" pageNumber="47">Mortoniella alicula</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25057E8007DFF9A99A9FB0FFDA5C28F" box="[399,543,1158,1184]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D650A007DFF8498B1FB20FB04C7A8" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="8BA83681007DFF9A98B1FB20FF5CC2EE" blockId="46.[151,543,1158,1217]" box="[151,230,1193,1217]" pageId="46" pageNumber="47">
<figureCitation id="132C2A04007DFF9A98B1FB20FF5CC2EE" box="[151,230,1193,1217]" captionStart="FIGURE 23" captionStartId="47.[151,250,1982,2005]" captionTargetBox="[229,1363,183,1974]" captionTargetId="figure-46@47.[209,1378,167,2002]" captionTargetPageId="47" captionText="FIGURE 23. Mortoniella alicula, new species, male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5287023" httpUri="https://zenodo.org/record/5287023/files/figure.png" pageId="46" pageNumber="47">Fig. 23</figureCitation>
</paragraph>
<paragraph id="8BA83681007DFF9A98B1FB7BFCE4C3CE" blockId="46.[151,1436,1265,2010]" pageId="46" pageNumber="47">
From other members of the
<taxonomicName id="4C174D02007DFF9A99FFFB7BFDEBC326" baseAuthorityName="Mosely" baseAuthorityYear="1939" box="[473,593,1266,1289]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="ormina">
<emphasis id="B963EA93007DFF9A99FFFB7BFE4AC326" box="[473,496,1266,1289]" italics="true" pageId="46" pageNumber="47">M</emphasis>
.
<emphasis id="B963EA93007DFF9A9A26FB7BFDEBC326" box="[512,593,1266,1289]" italics="true" pageId="46" pageNumber="47">ormina</emphasis>
</taxonomicName>
species group, this species is notably distinguished by the structure of its dorsal phallic spine, which has pronounced T-shaped apicolateral expansions and an apex that is upturned and posteriorly recurved. The long narrow paramere appendages are also distinctive as compared to other species described from this region. Among described species,
<taxonomicName id="4C174D02007DFF9A9ADFFAD7FCD7C35A" authorityName="Blahnik &amp; Holzenthal" authorityYear="2011" box="[761,877,1373,1397]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="alicula">
<emphasis id="B963EA93007DFF9A9ADFFAD7FCD7C35A" box="[761,877,1373,1397]" italics="true" pageId="46" pageNumber="47">M. alicula</emphasis>
</taxonomicName>
is undoubtedly most closely related to
<taxonomicName id="4C174D02007DFF9A9D03FAD7FEFEC3B6" authority="(Mosely, 1939)" baseAuthorityName="Mosely" baseAuthorityYear="1939" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="ormina">
<emphasis id="B963EA93007DFF9A9D03FAD7FA26C35A" box="[1317,1436,1374,1397]" italics="true" pageId="46" pageNumber="47">M. ormina</emphasis>
(
<bibRefCitation id="EF864B70007DFF9A98B8FA08FE86C3B6" author="Mosely" box="[158,316,1409,1433]" firstAuthor="Mosely" pageId="46" pageNumber="47" pagination="217 - 239" refId="ref42135" refString="Mosely, M. E. (1939) The Brazilian Hydroptilidae (Trichoptera). Novitates Zoologicae, 41, 217 - 239." type="journal article" year="1939">Mosely, 1939</bibRefCitation>
)
</taxonomicName>
, which also has the apex of the dorsal phallic spine similarly narrowed and posteriorly bent. Diagnostic differences include the much longer paramere appendages of
<taxonomicName id="4C174D02007DFF9A9BABFA2FFBBAC392" authorityName="Blahnik &amp; Holzenthal" authorityYear="2011" box="[909,1024,1445,1469]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="alicula">
<emphasis id="B963EA93007DFF9A9BABFA2FFBBAC392" box="[909,1024,1445,1469]" italics="true" pageId="46" pageNumber="47">M. alicula</emphasis>
</taxonomicName>
, as well as the more greatly widened dorsal phallic spine and the different armature of the phallicata.
</paragraph>
<paragraph id="8BA83681007DFF9A98E1FA64FC88C05E" blockId="46.[151,1436,1265,2010]" pageId="46" pageNumber="47">
Adult. Length of forewing: male
<quantity id="4CEF9B64007DFF9A9A11FA64FD35C02A" box="[567,655,1517,1541]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="46" pageNumber="47" unit="mm" value="2.1">2.1 mm</quantity>
. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0:3:4 (preapical spur of mesotibia very small). Overall color medium brown. Legs yellowish, tibial spurs darker in color, contrasting with legs. Wing bar at anastamosis marked with white, contrasting setae, apices of forewings with small white setal spots at apices of veins.
</paragraph>
<paragraph id="8BA83681007DFF8498E1F9F4FB04C7A8" blockId="46.[151,1436,1265,2010]" lastBlockId="48.[151,1436,151,535]" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
Male genitalia. Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base; apex dorsoventrally compressed, acute as viewed laterally (
<figureCitation id="132C2A04007DFF9A9C3CF928FBC5C096" box="[1050,1151,1697,1721]" captionStart="FIGURE 23" captionStartId="47.[151,250,1982,2005]" captionTargetBox="[229,1363,183,1974]" captionTargetId="figure-46@47.[209,1378,167,2002]" captionTargetPageId="47" captionText="FIGURE 23. Mortoniella alicula, new species, male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5287023" httpUri="https://zenodo.org/record/5287023/files/figure.png" pageId="46" pageNumber="47">Fig. 23E</figureCitation>
), rounded as viewed ventrally. Segment IX rounded anterolaterally, length greatest in ventral 1/2, posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by more than 1/2 width of segment. Tergum X elongate, simple in structure, with very deep, narrow U-shaped mesal excision; apical lobes broadly rounded. Inferior appendages short, inconspicuous, without dorsal or ventromesal projections, more or less fused ventromesally. Mesal pockets of inferior appendages with prominent, posteriorlydirected, spine-like projections, apparently fused to ventral margin of inferior appendages. Paramere appendages very narrow, elongate, slightly widened preapically, apices acute; appendages with broad U-shaped curvature, curved downward at base and upward at apex. Dorsal phallic spine of very distinctive structure, laterally with rounded, depressed projections; apically, as viewed dorsally, greatly widened, with T-shaped lateral projections and narrow, acute, strongly posteriorly curved dorsomesal projection. Phallicata with 2 pairs of relatively short spinelike processes, dorsally with mesally-directed, attenuate processes, crossing each other mesally, laterally with shorter, laterally-directed processes. Base of endophallic membrane (or apex of phallicata) with rounded, membranous (or lightly sclerotized) lateral projections, apex of endophallic membrane with paired, rounded, membranous projections, each with needle-like, sclerotized phallotremal spine, mesally with additional short spine (possibly modified apex of phallotremal sclerite). Phallotremal sclerite evident as small rounded sclerite.
</paragraph>
</subSubSection>
<caption id="DF686609007CFF9B98B1F837FBF1C1DC" ID-DOI="http://doi.org/10.5281/zenodo.5287023" ID-Zenodo-Dep="5287023" httpUri="https://zenodo.org/record/5287023/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1982,2005]" targetBox="[229,1363,183,1974]" targetPageId="47">
<paragraph id="8BA83681007CFF9B98B1F837FBF1C1DC" blockId="47.[151,1436,1982,2035]" pageId="47" pageNumber="48">
<emphasis id="B963EA93007CFF9B98B1F837FE9BC1FA" bold="true" box="[151,289,1982,2005]" pageId="47" pageNumber="48">FIGURE 23.</emphasis>
<taxonomicName id="4C174D02007CFF9B990EF836FE54C1FA" authority="Blahnik &amp; Holzenthal, 2011" authorityName="Blahnik &amp; Holzenthal" authorityYear="2011" box="[296,494,1983,2005]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="alicula" status="sp. nov.">
<emphasis id="B963EA93007CFF9B990EF836FE54C1FA" box="[296,494,1983,2005]" italics="true" pageId="47" pageNumber="48">Mortoniella alicula</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25057E8007CFF9B99DAF836FDC3C1FA" box="[508,633,1983,2005]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
, male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral.
</paragraph>
</caption>
<subSubSection id="C30D650A0063FF8498E1FE1DFC16C438" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8BA836810063FF8498E1FE1DFB1AC7E0" blockId="48.[151,1436,151,535]" pageId="48" pageNumber="49">
<materialsCitation id="3B7F3CDC0063FF8498E1FE1DFB1AC7E0" collectingDate="2002-03-08" collectionCode="MZUSP" collectorName="de Cima &amp; Holzenthal &amp; Blahnik &amp; Paprocki &amp; Prather" country="Brazil" elevation="1000" latitude="-22.394722" location="Macae de Cima" longLatPrecision="21" longitude="-42.502224" municipality="Rio Macae" pageId="48" pageNumber="49" specimenCode="UMSP000087906" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="B963EA930063FF8498E1FE1DFD01C783" bold="true" box="[199,699,403,428]" pageId="48" pageNumber="49">
<typeStatus id="54AC88230063FF8498E1FE1DFE8EC783" box="[199,308,404,428]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F30076110063FF8499A0FE1AFE4CC783" box="[390,502,403,428]" name="Brazil" pageId="48" pageNumber="49">BRAZIL</collectingCountry>
:
<collectingRegion id="49D3F8630063FF849A24FE1AFD0CC783" box="[514,694,403,428]" country="Brazil" name="Rio de Janeiro" pageId="48" pageNumber="49">Rio de Janeiro</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BCCACFB0063FF849AE5FE1DFCFAC783" box="[707,832,404,428]" pageId="48" pageNumber="49">Rio Macaé</collectingMunicipality>
,
<location id="8EC8605A0063FF849B6BFE1DFBBBC783" LSID="urn:lsid:plazi:treatment:03BE8797007DFF8498B1FB0FFC16C438:8EC8605A0063FF849B6BFE1DFBBBC783" box="[845,1025,404,428]" country="Brazil" latitude="-22.394722" longLatPrecision="21" longitude="-42.502224" municipality="Rio Macae" name="Macae de Cima" pageId="48" pageNumber="49" stateProvince="Rio de Janeiro">
Macaé
<collectorName id="26E253570063FF849BB9FE1DFBBBC783" box="[927,1025,404,428]" pageId="48" pageNumber="49">de Cima</collectorName>
</location>
,
<geoCoordinate id="EE2350460063FF849C2BFE1DFB34C783" box="[1037,1166,404,428]" degrees="22" direction="south" minutes="23" orientation="latitude" pageId="48" pageNumber="49" precision="15" seconds="41" value="-22.394722">22°23'41&quot;S</geoCoordinate>
,
<geoCoordinate id="EE2350460063FF849CBDFE1DFA89C783" box="[1179,1331,404,428]" degrees="042" direction="west" minutes="30" orientation="longitude" pageId="48" pageNumber="49" precision="15" seconds="08" value="-42.502224">042°30'08&quot;W</geoCoordinate>
,
<quantity id="4CEF9B640063FF849D19FE1DFA2DC784" box="[1343,1431,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="1000.0">
<elevation id="003AD1B20063FF849D19FE1DFA2DC784" box="[1343,1431,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFA910410063FF8498B1FE3EFEBFC7E0" box="[151,261,439,463]" pageId="48" pageNumber="49" value="2002-03-08">
<collectingDate id="EFEDE9A90063FF8498B1FE3EFEBFC7E0" box="[151,261,439,463]" pageId="48" pageNumber="49" value="2002-03-08">8.iii.2002</collectingDate>
</date>
,
<collectorName id="26E253570063FF849936FE3EFE34C7E0" box="[272,398,439,463]" pageId="48" pageNumber="49">Holzenthal</collectorName>
,
<collectorName id="26E253570063FF8499BFFE3EFE4EC7E0" box="[409,500,439,463]" pageId="48" pageNumber="49">Blahnik</collectorName>
,
<collectorName id="26E253570063FF849A26FE3EFDD9C7E0" box="[512,611,439,463]" pageId="48" pageNumber="49">Paprocki</collectorName>
&amp;
<collectorName id="26E253570063FF849AA1FE3EFD6DC7E0" box="[647,727,439,463]" pageId="48" pageNumber="49">Prather</collectorName>
(
<specimenCode id="DBB19EFA0063FF849AC0FE3EFC0EC7E0" box="[742,948,439,463]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="48" pageNumber="49" type="University or college">UMSP000087906</specimenCode>
) (pinned) (
<collectionCode id="ED06AE440063FF849C16FE3EFB2EC7E0" box="[1072,1172,439,463]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="48" pageNumber="49">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA836810063FF8498E1FE55FC16C438" blockId="48.[151,1436,151,535]" pageId="48" pageNumber="49">
Etymology. This species is named
<taxonomicName id="4C174D020063FF849A6DFE54FD04C7DB" authorityName="Blahnik &amp; Holzenthal" authorityYear="2011" box="[587,702,476,500]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="alicula">
<emphasis id="B963EA930063FF849A6DFE54FD04C7DB" box="[587,702,476,500]" italics="true" pageId="48" pageNumber="49">M. alicula</emphasis>
</taxonomicName>
, from a Latin word for wing and referring to the wing-like lateral expansions of the apical part of the dorsal phallic spine of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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