434 lines
63 KiB
XML
434 lines
63 KiB
XML
<document id="CB3C9FB0051C52ECB04646F1A0697236" ID-DOI="10.11646/zootaxa.4033.3.7" ID-GBIF-Dataset="a0232398-33ad-4502-a5a3-9f15a3f6e3be" ID-ISSN="1175-5326" ID-Zenodo-Dep="239105" ID-ZooBank="619026AF-B2D0-4C41-B6FF-D8CD8339B092" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461297700573" checkinUser="plazi" docAuthor="Sites, Robert W." docDate="2015" docId="03BE878487689238FF1456C6C18A8E86" docLanguage="en" docName="zt04033p438.pdf" docOrigin="Zootaxa 4033 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hygropetrocoris Sites, NEW GENUS" docType="treatment" docVersion="8" lastPageNumber="431" masterDocId="FF87FFFC8769923CFF835756C3218F05" masterDocTitle="New taxa of Cryphocricinae (Heteroptera: Naucoridae) from the Guiana Shield: Hygropetrocoris Sites, n. gen. and two new species of Ambrysus Stål" masterLastPageNumber="438" masterPageNumber="427" pageNumber="428" updateTime="1698616807018" updateUser="plazi">
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<mods:titleInfo id="67CABA463F5A66ABDC8381887CD066BD">
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<mods:title id="13C368AD47775FC83F18CF9A7E5DA727">New taxa of Cryphocricinae (Heteroptera: Naucoridae) from the Guiana Shield: Hygropetrocoris Sites, n. gen. and two new species of Ambrysus Stål</mods:title>
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<mods:namePart id="A5C4DE243A46D480E0B047F627946B4A">Sites, Robert W.</mods:namePart>
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<mods:title id="5C319941F2A4514D2C3EFB13500655A0">Zootaxa</mods:title>
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<mods:part id="49F8890E25CDE2B57551412DDFC9BFC1">
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<mods:date id="6C1344CE6A14FF001865DEF853482C2C">2015</mods:date>
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<mods:number id="5C90B15C56DF89F00D7E13F13BCBE986">4033</mods:number>
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<mods:identifier id="2B2E5B8E5F85BBF1C2C9A3D4C8EE434E" type="DOI">10.11646/zootaxa.4033.3.7</mods:identifier>
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<treatment id="03BE878487689238FF1456C6C18A8E86" ID-DOI="http://doi.org/10.5281/zenodo.6097799" ID-GBIF-Taxon="119643049" ID-Zenodo-Dep="6097799" LSID="urn:lsid:plazi:treatment:03BE878487689238FF1456C6C18A8E86" httpUri="http://treatment.plazi.org/id/03BE878487689238FF1456C6C18A8E86" lastPageId="4" lastPageNumber="431" pageId="1" pageNumber="428">
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<subSubSection id="C30D65198768923DFF1456C6C2218EC9" pageId="1" pageNumber="428" type="nomenclature">
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<paragraph id="8BA836928768923DFF1456C6C1768EAE" blockId="1.[151,599,400,461]" box="[151,599,400,427]" pageId="1" pageNumber="428">
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<heading id="D0E081FE8768923DFF1456C6C1768EAE" bold="true" box="[151,599,400,427]" fontSize="11" level="1" pageId="1" pageNumber="428" reason="1">
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<emphasis id="B963EA808768923DFF1456C6C1768EAE" bold="true" box="[151,599,400,427]" pageId="1" pageNumber="428">
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<taxonomicName id="4C174D118768923DFF1456C6C2BF8EAE" ID-CoL="4ZXZ" authority="Sites" authorityName="Sites" box="[151,414,400,427]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B963EA808768923DFF1456C6C27D8EAF" bold="true" box="[151,348,400,426]" italics="true" pageId="1" pageNumber="428">Hygropetrocoris</emphasis>
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Sites
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</taxonomicName>
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<taxonomicNameLabel id="A25057FB8768923DFE2656C7C1768EAE" box="[421,599,401,427]" pageId="1" pageNumber="428" rank="genus">NEW GENUS</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA836928768923DFF1456E2C2218EC9" blockId="1.[151,599,400,461]" box="[151,256,436,461]" pageId="1" pageNumber="428">
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<figureCitation id="132C2A178768923DFF1456E2C2218EC9" box="[151,256,436,461]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[199,298,1975,1997]" captionStartId-1="4.[151,250,1903,1925]" captionStartId-2="5.[151,250,1915,1937]" captionStartId-3="6.[151,250,1372,1394]" captionTargetBox-0="[151,1436,1125,1951]" captionTargetBox-1="[190,1395,874,1877]" captionTargetBox-2="[184,1389,731,1869]" captionTargetBox-3="[151,1436,772,1347]" captionTargetId-0="figure@1.[151,1436,1125,1951]" captionTargetId-1="figure@4.[190,1395,874,1877]" captionTargetId-2="figure@5.[184,1389,731,1870]" captionTargetId-3="figure@6.[151,1436,772,1348]" captionTargetPageId-0="1" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." captionText-1="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." captionText-2="FIGURE 3. Features of Hygropetrocoris guyana n. sp. (A) 6 th and 7 th abdominal terga; (B) 8 th abdominal tergum showing medial black pseudoparameres; (C) pygophore with phallosoma and parameres with (left) and without (right) terga IX and X (proctiger), size bar = 0.5 mm; (D) ventral surface of phallosoma." captionText-3="FIGURE 4. Type locality of Hygropetrocoris guyana n. gen. and Ambrysus brunneus n. sp. showing (A) general habitat and (B) specific habitat of H. guyana, which is in the center of the bedrock of Fig. 4 A. A. brunneus most likely was collected in small pools or rootmats of the vegetation on the left margin of the bedrock." httpUri-0="https://zenodo.org/record/239106/files/figure.png" httpUri-1="https://zenodo.org/record/239107/files/figure.png" httpUri-2="https://zenodo.org/record/239108/files/figure.png" httpUri-3="https://zenodo.org/record/239109/files/figure.png" pageId="1" pageNumber="428">Figs. 1–4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30D65198768923DFF1456ABC1DF8D17" box="[151,766,508,530]" pageId="1" pageNumber="428" type="materials_examined">
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<paragraph id="8BA836928768923DFF1456ABC1DF8D17" blockId="1.[151,766,508,530]" box="[151,766,508,530]" pageId="1" pageNumber="428">
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<typeStatus id="54AC88308768923DFF1456ABC3E88D17" box="[151,201,509,530]" pageId="1" pageNumber="428">Type</typeStatus>
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species.
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<taxonomicName id="4C174D118768923DFEA656AAC1758D17" authority="Sites" authorityName="Sites" box="[293,596,508,530]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="guyana" status="sp. nov.">
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<emphasis id="B963EA808768923DFEA656AAC13C8D14" box="[293,541,508,529]" italics="true" pageId="1" pageNumber="428">Hygropetrocoris guyana</emphasis>
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Sites
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</taxonomicName>
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<taxonomicNameLabel id="A25057FB8768923DFDD856ABC1DF8D17" box="[603,766,508,530]" pageId="1" pageNumber="428" rank="species">
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<collectionCode id="ED06AE578768923DFDD856ABC1B48D17" box="[603,661,509,530]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="1" pageNumber="428">NEW</collectionCode>
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<collectionCode id="ED06AE578768923DFD1856AAC1DF8D17" box="[667,766,508,530]" pageId="1" pageNumber="428">SPECIES</collectionCode>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30D65198768923EFF145568C7078AE0" lastPageId="2" lastPageNumber="429" pageId="1" pageNumber="428" type="description">
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<paragraph id="8BA836928768923DFF145568C0D88DE3" blockId="1.[151,1437,574,1070]" pageId="1" pageNumber="428">
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<emphasis id="B963EA808768923DFF145568C2088D52" bold="true" box="[151,297,574,599]" pageId="1" pageNumber="428">Description.</emphasis>
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Length 8.96–9.20 (mean = 9.08), maximum width 5.60–5.84 (mean = 5.72). Body ovate, parallelsided, broadly rounded anteriorly, narrowly rounded posteriorly, macropterous; widest across embolia; dorsally head and pronotum light colored but heavily speckled with dark brown punctures; scutellum patterned with light and dark brown and black, wings patterned with white, brown, and black; membrane black (
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<figureCitation id="132C2A178768923DFB1955FCC7FA8DC6" box="[1178,1243,682,707]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
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). Non-uniformly coarsely punctate throughout. Pronotum and scutellum irregularly tumescent.
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</paragraph>
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<paragraph id="8BA836928768923DFF4455A3C7F48B2B" blockId="1.[151,1437,574,1070]" pageId="1" pageNumber="428">
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<emphasis id="B963EA808768923DFF4455A3C2258C0B" box="[199,260,757,782]" italics="true" pageId="1" pageNumber="428">Head</emphasis>
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. Head declivent ca. 70o with respect to long axis of body (
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<figureCitation id="132C2A178768923DFC1355A3C0F58C0B" box="[912,980,757,782]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="1" pageNumber="428">Fig. 2</figureCitation>
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B); not continuous with lateral margin of pronotum (
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<figureCitation id="132C2A178768923DFE95544FC27A8C37" box="[278,347,793,818]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
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); compound eyes bulging, elevated above level of vertex (
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<figureCitation id="132C2A178768923DFB8F544FC77F8C37" box="[1036,1118,793,818]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="1" pageNumber="428">Figs. 2</figureCitation>
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B, C), inner margin convex when viewed from above (
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<figureCitation id="132C2A178768923DFE4E546BC1318C53" box="[461,528,829,854]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
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) and divergent when viewed from front (
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<figureCitation id="132C2A178768923DFC77546BC71B8C53" box="[1012,1082,829,854]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="1" pageNumber="428">Fig. 2</figureCitation>
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C); series of four setal rosettes paralleling inner margin of eye; posterior margin of head slightly convex between eyes (
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<figureCitation id="132C2A178768923DFB185437C7C18C7F" box="[1179,1248,865,890]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
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); layer of black recumbent hairs appressed to occiput at midline and continuing around side under eye; labrum broad, rounded, well-developed (
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<figureCitation id="132C2A178768923DFEDB54FFC28D8CC7" box="[344,428,937,962]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="1" pageNumber="428">Figs. 2</figureCitation>
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B, C); rostrum short, three visible segments, first visible segment partially concealed behind labrum, each subsequent segment progressively narrower (
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<figureCitation id="132C2A178768923DFC11549BC0F88CE3" box="[914,985,973,998]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="1" pageNumber="428">Fig. 2</figureCitation>
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B), reaching to near anterior margin of prothoracic coxae; maxillary plate impubis with distal margin nearly straight; antennae brown, segment one short and conspicuous, two and four subequal in length, three longest, segments three and four hirsute.
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</paragraph>
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<caption id="DF68661A8768923DFF4450E1C70188C8" box="[199,1056,1975,1997]" httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="1" pageNumber="428" targetBox="[151,1436,1125,1951]" targetPageId="1">
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<paragraph id="8BA836928768923DFF4450E1C70188C8" blockId="1.[199,1056,1975,1997]" box="[199,1056,1975,1997]" pageId="1" pageNumber="428">
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<emphasis id="B963EA808768923DFF4450E1C26288C9" bold="true" box="[199,323,1975,1997]" pageId="1" pageNumber="428">FIGURE 1.</emphasis>
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<taxonomicName id="4C174D118768923DFECA50EEC16188C8" box="[329,576,1976,1997]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="guyana" status="gen. nov.">
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<emphasis id="B963EA808768923DFECA50EEC16188C8" box="[329,576,1976,1997]" italics="true" pageId="1" pageNumber="428">Hygropetrocoris guyana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25057FB8768923DFDC450EEC1AF88C8" box="[583,654,1976,1997]" pageId="1" pageNumber="428" rank="genus">
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<emphasis id="B963EA808768923DFDC450EEC1AF88C8" bold="true" box="[583,654,1976,1997]" pageId="1" pageNumber="428">n. gen.</emphasis>
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</taxonomicNameLabel>
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dorsal (left) and ventral (right) habitus.
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</paragraph>
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</caption>
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<paragraph id="8BA83692876B923EFF4457C1C1A38DC8" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="429">
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<emphasis id="B963EA80876B923EFF4457C1C2398FB5" box="[199,280,151,176]" italics="true" pageId="2" pageNumber="429">Thorax</emphasis>
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. Pronotum nearly straight behind head (
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<figureCitation id="132C2A17876B923EFD7257C1C0178FB5" box="[753,822,151,176]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="2" pageNumber="429">Fig. 1</figureCitation>
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); transverse sulcus setting off band in posterior 1/4; transverse band ending at anterior end of clavus, end tumescent and with tuft of dark setae; anterior 3/4 with profuse dark punctation throughout; brush of elongate sensory setae on anterior surface behind eye; lateral margins smooth, convex; posterolateral corners broadly rounded; row of four to five clusters of stiff dark setae near lateral margin (
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<figureCitation id="132C2A17876B923EFF7A5671C26B8E45" box="[249,330,295,320]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="2" pageNumber="429">Figs. 1</figureCitation>
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,
|
||
<figureCitation id="132C2A17876B923EFEDF567EC24A8E45" box="[348,363,296,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="2" pageNumber="429">2</figureCitation>
|
||
B), sparse erect single erect setae scattered throughout lateral 1/3; posterior margin straight; pronounced tumescences in anterior 3/4 behind eye, midlateral, and on midline near transverse sulcus. Propleura meeting at midline (
|
||
<figureCitation id="132C2A17876B923EFEFE5639C2E18E8D" box="[381,448,367,392]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="2" pageNumber="429">Fig. 1</figureCitation>
|
||
), anterior margin joined to prosternum; prosternal midline with carina between coxae. Scutellum large, irregularly and coarsely punctate, raised above level of wings, broadly tumescent, erect curved black setae in longitudinal row of tufts, patterned with light and dark brown, lateral margins slightly sinuate (
|
||
<figureCitation id="132C2A17876B923EFAEC56E1C3858EF1" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="2" pageNumber="429">Fig. 1</figureCitation>
|
||
). Mesosternum flat, continuous with mesepisternum except for lightly pigmented line of fusion. Mesosternellum broadly pointed. Metasternellum (=metaxyphus) transverse; strongly acuminate at posterior midline; with strong, narrow, midventral ridge. Hemelytra finely punctate throughout, with distinct clavus and embolium (
|
||
<figureCitation id="132C2A17876B923EFAB35572C6598D39" box="[1328,1400,548,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="2" pageNumber="429">Fig. 2</figureCitation>
|
||
A); clavus with longitudinal intraclaval suture (
|
||
<figureCitation id="132C2A17876B923EFD025511C1E58D65" box="[641,708,583,608]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="2" pageNumber="429">Fig. 2</figureCitation>
|
||
A) between claval suture and scutellum, claval commissure with longitudinal ridge; embolium with lateral margin mostly straight, arcuate at ends; embolar suture sinuate; patterns of erect curved black setae in rows, tufts, and singular on hemelytra except membrane (
|
||
<figureCitation id="132C2A17876B923EFBEF55D9C7918DAD" box="[1132,1200,655,680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="2" pageNumber="429">Fig. 2</figureCitation>
|
||
A); membrane black, finely punctate. Hindwings well-developed.
|
||
</paragraph>
|
||
<paragraph id="8BA83692876B923EFF44558FC2EB8BC0" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="429">
|
||
<emphasis id="B963EA80876B923EFF44558FC3DD8DF5" box="[199,252,729,752]" italics="true" pageId="2" pageNumber="429">Legs</emphasis>
|
||
. Foreleg with femur broad, strongly widened posterobasally, distal half of anterior margin excavated, anterior margin with thick band of setae becoming paired in basal half; tibia narrow, with pronounced tooth in basal third; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular (
|
||
<figureCitation id="132C2A17876B923EFBE25449C7888C3D" box="[1121,1193,799,824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1925]" captionTargetBox="[190,1395,874,1877]" captionTargetId="figure@4.[190,1395,874,1877]" captionTargetPageId="4" captionText="FIGURE 2. Features of Hygropetrocoris guyana n. gen. (A) left hemelytron showing hair lines and hair patches on corium and embolium (yellow arrows); (B) lateral view showing 70 degree orientation of head with respect to the pronotum and marginal tufts of dark setae; (C) frontal view of head and prothoracic legs; (D) dorsal view of right prothoracic leg. cls = claval suture, ics = intraclaval suture, IV = abdominal segment IV." httpUri="https://zenodo.org/record/239107/files/figure.png" pageId="2" pageNumber="429">Fig. 2</figureCitation>
|
||
D). Middle and hind trochanters and femora flattened and glabrous, femora with sparse elongate pale hairs and row of minute reddish brown spines on posterior margin; middle femur slightly curved dorsad following curvature of venter; middle and hind tibiae with numerous stout reddish brown spines and continuous circlet of spines at distal end except on dorsal side. Middle tibia gradually widening distally; without pad of hairs on ventral surface; two rows of posteroventral heavy spines, multiple rows of anterior heavy spines, row of posterodorsal swimming hairs. Middle tarsus threesegmented, tarsomere one short, acuminate, and produced beneath base of tarsomere two.
|
||
<collectingCountry id="F3007602876B923EFB3E54A1C7D48B15" box="[1213,1269,1015,1040]" name="India" pageId="2" pageNumber="429">Hind</collectingCountry>
|
||
leg with tibia elongate, thin, multiple rows of spines on anterior and posterior margins, posterodorsal swimming hairs on tibia and tarsus; tarsus three-segmented, tarsomere one short, acuminate, and produced beneath base of tarsomere two, two long, three longer. Middle and hind pretarsi darkly colored; with short empodia; claws symmetrical, long, evenly curved, each with rounded basal tooth. Meso- and metathoracic coxae covered with ventromesal setae, becoming elongate distally.
|
||
</paragraph>
|
||
<paragraph id="8BA83692876B923EFF445399C7078AE0" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="429">
|
||
<emphasis id="B963EA80876B923EFF445399C2108BED" box="[199,305,1231,1256]" italics="true" pageId="2" pageNumber="429">Abdomen</emphasis>
|
||
. Dorsally dark brown to black medially, connexiva pale yellow. Connexiva with margins smooth; posterolateral corners of II–IV narrowly rounded, V–VI broadly rounded, VII broadly acute; posterior third of III– VII with tuft of black setae near margin, III–V with scattered black setae mesad of tuft, III–VIII with marginal fringe of pale setae. Accessory genitalic process of mediotergite VI absent (
|
||
<figureCitation id="132C2A17876B923EFBAA526AC7508A51" box="[1065,1137,1340,1365]" captionStart="FIGURE 3" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[184,1389,731,1869]" captionTargetId="figure@5.[184,1389,731,1870]" captionTargetPageId="5" captionText="FIGURE 3. Features of Hygropetrocoris guyana n. sp. (A) 6 th and 7 th abdominal terga; (B) 8 th abdominal tergum showing medial black pseudoparameres; (C) pygophore with phallosoma and parameres with (left) and without (right) terga IX and X (proctiger), size bar = 0.5 mm; (D) ventral surface of phallosoma." httpUri="https://zenodo.org/record/239108/files/figure.png" pageId="2" pageNumber="429">Fig. 3</figureCitation>
|
||
A). Medial lobes of VIII (pseudoparameres) large, black, angled posterolaterally, extending over lateral lobes (
|
||
<figureCitation id="132C2A17876B923EFBD55209C7B88A7D" box="[1110,1177,1375,1400]" captionStart="FIGURE 3" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[184,1389,731,1869]" captionTargetId="figure@5.[184,1389,731,1870]" captionTargetPageId="5" captionText="FIGURE 3. Features of Hygropetrocoris guyana n. sp. (A) 6 th and 7 th abdominal terga; (B) 8 th abdominal tergum showing medial black pseudoparameres; (C) pygophore with phallosoma and parameres with (left) and without (right) terga IX and X (proctiger), size bar = 0.5 mm; (D) ventral surface of phallosoma." httpUri="https://zenodo.org/record/239108/files/figure.png" pageId="2" pageNumber="429">Fig. 3</figureCitation>
|
||
B). Abdomen ventrally generally covered with fine, light brown setae; with glabrous pale yellow lateral band; posterior margin of V asymmetrically concave on left side (
|
||
<figureCitation id="132C2A17876B923EFDB452F1C1598AC5" box="[567,632,1447,1472]" captionStart="FIGURE 1" captionStartId="1.[199,298,1975,1997]" captionTargetBox="[151,1436,1125,1951]" captionTargetId="figure@1.[151,1436,1125,1951]" captionTargetPageId="1" captionText="FIGURE 1. Hygropetrocoris guyana n. gen. dorsal (left) and ventral (right) habitus." httpUri="https://zenodo.org/record/239106/files/figure.png" pageId="2" pageNumber="429">Fig. 1</figureCitation>
|
||
). Transverse glabrous patches on at least II-VI, patches paired on III–V. Spiracles III–VI evident between and slightly anterior to paired glabrous patches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6519876B923FFF4452B9C3CB8DA0" lastPageId="3" lastPageNumber="430" pageId="2" pageNumber="429" type="diagnosis">
|
||
<paragraph id="8BA83692876B923EFF4452B9C11C8970" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="429">
|
||
<emphasis id="B963EA80876B923EFF4452B9C260890D" bold="true" box="[199,321,1519,1544]" pageId="2" pageNumber="429">Diagnosis.</emphasis>
|
||
The most striking attributes of this insect are the (1) nearly vertical head is relatively small and apparently more mobile with respect to the prothorax than in most other naucorids, (2) bulging compound eyes, (3) tufts of setae around the perimeter of the insect, (4) tumescent pronotum and scutellum, and (5) pattern of black setae on the hemelytra and scutellum.
|
||
</paragraph>
|
||
<paragraph id="8BA83692876B923EFF445129C75188BC" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="429">
|
||
<emphasis id="B963EA80876B923EFF445129C262899D" bold="true" box="[199,323,1663,1688]" pageId="2" pageNumber="429">Taxonomy</emphasis>
|
||
. Five subfamilies of
|
||
<taxonomicName id="4C174D11876B923EFDBA5129C19B899D" box="[569,698,1663,1688]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="family">Naucoridae</taxonomicName>
|
||
currently are recognized.
|
||
<taxonomicName id="4C174D11876B923EFC6A51D7C783899D" box="[1001,1186,1665,1688]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EA80876B923EFC6A51D7C783899D" box="[1001,1186,1665,1688]" italics="true" pageId="2" pageNumber="429">Hygropetrocoris</emphasis>
|
||
</taxonomicName>
|
||
is hereby assigned to
|
||
<taxonomicName id="4C174D11876B923EFF1451F2C26689B8" box="[151,327,1700,1725]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
|
||
, Ambrysini. The insect does not have (1) the ventral thoracic carinae characteristic of
|
||
<taxonomicName id="4C174D11876B923EFF145191C21689E5" box="[151,311,1735,1760]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="subFamily" subFamily="Limnocorinae">Limnocorinae</taxonomicName>
|
||
, (2) the tibial pad of setae, mid-ventral abdominal band of setae, front of the head folded posteroventrally, and other features characteristic of
|
||
<taxonomicName id="4C174D11876B923EFC8F51BAC0878800" box="[780,934,1772,1797]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
|
||
, nor (3) the rostrum arising from a deep excavation set back from the anterior margin of the head characteristic of
|
||
<taxonomicName id="4C174D11876B923EFC525059C75C882D" box="[977,1149,1807,1832]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
|
||
, which is restricted to the Old World based on known taxa. This leaves the New World subfamily
|
||
<taxonomicName id="4C174D11876B923EFC6E5062C7BB8848" box="[1005,1178,1844,1869]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
|
||
and the cosmopolitan
|
||
<taxonomicName id="4C174D11876B923EFF145001C23D8875" box="[151,284,1879,1904]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
|
||
. The propleura wrap ventrally and extend over the prosternellum to touch at the midline, which is a characteristic found in Ambrysini, and the pseudoparameres are enlarged and directed posterolaterad which is found in
|
||
<emphasis id="B963EA80876B923EFF7F50F6C2B988BC" box="[252,408,1952,1977]" italics="true" pageId="2" pageNumber="429">Carvalhoiella</emphasis>
|
||
,
|
||
<taxonomicName id="4C174D11876B923EFE2650F6C03488BC" authority="La Rivers" authorityName="La Rivers" box="[421,789,1952,1977]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="montandoni">
|
||
<emphasis id="B963EA80876B923EFE2650F6C18088BC" box="[421,673,1952,1977]" italics="true" pageId="2" pageNumber="429">Ambrysus montandoni</emphasis>
|
||
La Rivers
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C174D11876B923EFCCF50F7C75188BC" authority="La Rivers." authorityName="La Rivers." box="[844,1136,1952,1977]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="maldonadus">
|
||
<emphasis id="B963EA80876B923EFCCF50F7C0D588BC" box="[844,1012,1952,1977]" italics="true" pageId="2" pageNumber="429">A. maldonadus</emphasis>
|
||
La Rivers.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BA83692876A923FFF4457C1C3CB8DA0" blockId="3.[151,1436,151,677]" pageId="3" pageNumber="430">
|
||
Despite its aberrant physiognomy,
|
||
<taxonomicName id="4C174D11876A923FFDC957CFC0228FB5" box="[586,771,153,176]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EA80876A923FFDC957CFC0228FB5" box="[586,771,153,176]" italics="true" pageId="3" pageNumber="430">Hygropetrocoris</emphasis>
|
||
</taxonomicName>
|
||
seems most closely related to two groups of Ambrysini: (1)
|
||
<taxonomicName id="4C174D11876A923FFF1457EFC2B38FD7" box="[151,402,185,210]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="montandoni">
|
||
<emphasis id="B963EA80876A923FFF1457EFC2B38FD7" box="[151,402,185,210]" italics="true" pageId="3" pageNumber="430">Ambrysus montandoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C174D11876A923FFE4F57ECC1548FD7" box="[460,629,185,210]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="maldonadus">
|
||
<emphasis id="B963EA80876A923FFE4F57ECC1548FD7" box="[460,629,185,210]" italics="true" pageId="3" pageNumber="430">A. maldonadus</emphasis>
|
||
</taxonomicName>
|
||
and (2)
|
||
<emphasis id="B963EA80876A923FFD5557EFC0538FD7" box="[726,882,185,210]" italics="true" pageId="3" pageNumber="430">Carvalhoiella</emphasis>
|
||
.
|
||
<taxonomicName id="4C174D11876A923FFC0257EFC75C8FD7" box="[897,1149,185,210]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="montandoni">
|
||
<emphasis id="B963EA80876A923FFC0257EFC75C8FD7" box="[897,1149,185,210]" italics="true" pageId="3" pageNumber="430">Ambrysus montandoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C174D11876A923FFB3557ECC67E8FD7" box="[1206,1375,185,210]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="maldonadus">
|
||
<emphasis id="B963EA80876A923FFB3557ECC67E8FD7" box="[1206,1375,185,210]" italics="true" pageId="3" pageNumber="430">A. maldonadus</emphasis>
|
||
</taxonomicName>
|
||
have features that distinguish them from congeners and will be removed from
|
||
<taxonomicName id="4C174D11876A923FFC6B578CC7778FF6" box="[1000,1110,218,243]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFC6B578CC7778FF6" box="[1000,1110,218,243]" italics="true" pageId="3" pageNumber="430">Ambrysus</emphasis>
|
||
</taxonomicName>
|
||
in an upcoming publication with Higor Rodrigues.
|
||
<emphasis id="B963EA80876A923FFE1E57AAC1188E10" box="[413,569,252,277]" italics="true" pageId="3" pageNumber="430">Carvalhoiella</emphasis>
|
||
originally was placed in
|
||
<taxonomicName id="4C174D11876A923FFCD957AAC0C28E10" box="[858,995,252,277]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
|
||
[subsequently ranked as a tribe within
|
||
<taxonomicName id="4C174D11876A923FFF14564BC2658E33" box="[151,324,285,310]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF864B63876A923FFED5564BC1738E33" author="Stys" box="[342,594,285,310]" pageId="3" pageNumber="430" refString="Stys, P. & Jansson, A. (1988) Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica, 50, 1 - 44." type="journal article" year="1988">Štys & Jansson 1988</bibRefCitation>
|
||
)].
|
||
<bibRefCitation id="EF864B63876A923FFDF2564BC02E8E33" author="Nieser" box="[625,783,285,310]" pageId="3" pageNumber="430" refString="Nieser, N. (1975) The water bugs (Heteroptera: Nepomorpha) of the Guyana Region. Studies on the Fauna of Suriname and other Guyanas, 59, 1 - 310, plates 1 - 24." type="journal article" year="1975">Nieser (1975)</bibRefCitation>
|
||
transferred
|
||
<emphasis id="B963EA80876A923FFC1E564BC7198E33" box="[925,1080,285,310]" italics="true" pageId="3" pageNumber="430">Carvalhoiella</emphasis>
|
||
to
|
||
<taxonomicName id="4C174D11876A923FFBE7564BC7C98E33" box="[1124,1256,285,310]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
|
||
, Naucorini, but later transferred it back to
|
||
<taxonomicName id="4C174D11876A923FFE4E5668C1778E52" box="[461,598,318,343]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
|
||
after examining specimens and describing
|
||
<taxonomicName id="4C174D11876A923FFBCF5669C2B18E7D" authority="Nieser, Pelli & Melo (Nieser et al. 1999)" authorityName="Nieser, Pelli & Melo (Nieser et al." authorityYear="1999" pageId="3" pageNumber="430" rank="species" species="stysi">
|
||
<emphasis id="B963EA80876A923FFBCF5669C7838E53" box="[1100,1186,319,342]" italics="true" pageId="3" pageNumber="430">C. stysi</emphasis>
|
||
<bibRefCitation id="EF864B63876A923FFB285668C2B18E7D" pageId="3" pageNumber="430" refString="Nieser, N., Pelli, A. & Melo, A. L. (1999) Two new Ambrysinae (Heteroptera: Naucoridae) from Minas Gerais, Brazil. Acta Societatis Zoologicae Bohemoslovacae, 63, 157 - 163." type="journal article">
|
||
Nieser, Pelli & Melo (Nieser
|
||
<emphasis id="B963EA80876A923FFF795637C21A8E7D" box="[250,315,351,376]" italics="true" pageId="3" pageNumber="430">et al.</emphasis>
|
||
1999)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B963EA80876A923FFE265637C0FF8E7D" box="[421,990,351,376]" italics="true" pageId="3" pageNumber="430">
|
||
<taxonomicName id="4C174D11876A923FFE265637C1408E7D" box="[421,609,353,376]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">Hygropetrocoris</taxonomicName>
|
||
, Carvalhoiella,
|
||
<taxonomicName id="4C174D11876A923FFCAB5636C0FB8E7D" box="[808,986,351,376]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="montandoni">A. montandoni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C174D11876A923FFBA45636C7F88E7D" box="[1063,1241,351,376]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="maldonadus">
|
||
<emphasis id="B963EA80876A923FFBA45636C7F88E7D" box="[1063,1241,351,376]" italics="true" pageId="3" pageNumber="430">A. maldonadus</emphasis>
|
||
</taxonomicName>
|
||
share enlarged, posterolaterally-directed pseudoparameres (median lobes of tergum 8), loss of setation on some mediosternites, and a very shallow or no invagination of the head into the anterior margin of the pronotum. Together with
|
||
<emphasis id="B963EA80876A923FFF145693C2608ED9" box="[151,321,453,476]" italics="true" pageId="3" pageNumber="430">Cataractocoris</emphasis>
|
||
, all share a wide, marginal, glabrous, ventral abdominal band. Autapomorphies of
|
||
<taxonomicName id="4C174D11876A923FFB605693C6BD8ED9" box="[1251,1436,453,476]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EA80876A923FFB605693C6BD8ED9" box="[1251,1436,453,476]" italics="true" pageId="3" pageNumber="430">Hygropetrocoris</emphasis>
|
||
</taxonomicName>
|
||
are the bulbous eyes, nearly-vertical head orientation, hemelytral hair lines and patches, stout marginal tufts of setae from the pronotum to the abdominal segments, and the profemur excavated distally with a corresponding tooth on the protibia (see Table 1). The excavated profemur and toothed protibia might be restricted to only males because similar modifications of the prothoracic legs is a sexually dimorphic character in many species of the Asian genera
|
||
<emphasis id="B963EA80876A923FFEB8553CC17F8D87" box="[315,606,618,643]" italics="true" pageId="3" pageNumber="430">
|
||
<taxonomicName id="4C174D11876A923FFEB8553CC17B8D87" authority="Coptocatus" authorityName="Coptocatus" box="[315,602,618,643]" class="Insecta" family="Naucoridae" genus="Cheirochela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">Cheirochela, Coptocatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B963EA80876A923FFD1C553CC03D8D86" box="[671,796,618,643]" italics="true" pageId="3" pageNumber="430">Gestroiella</emphasis>
|
||
(
|
||
<taxonomicName id="4C174D11876A923FFCB3553CC0C08D86" box="[816,993,618,643]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
|
||
). Females of
|
||
<taxonomicName id="4C174D11876A923FFB06553AC61F8D86" box="[1157,1342,620,643]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EA80876A923FFB06553AC61F8D86" box="[1157,1342,620,643]" italics="true" pageId="3" pageNumber="430">Hygropetrocoris</emphasis>
|
||
</taxonomicName>
|
||
are not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6519876A9238FF4452DCC0458E1D" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430" type="discussion">
|
||
<paragraph id="8BA83692876A9238FF4452DCC0458E1D" blockId="3.[151,1437,1418,2032]" lastBlockId="4.[151,1436,151,387]" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430">
|
||
<emphasis id="B963EA80876A923FFF4452DCC26D8AA6" bold="true" box="[199,332,1418,1443]" pageId="3" pageNumber="430">Discussion.</emphasis>
|
||
The bulbous eyes, sensory hairs, and negligible invagination of the head into the pronotum suggest the head is more freely movable with respect to the pronotum than in other naucorids in which the head is relatively fixed and mostly acts in concert with the prothorax. This along with its irregularly shaped body surface and mottled color pattern are better suited for a terrestrial existence rather than living underwater, and its general appearance is reminiscent of
|
||
<taxonomicName id="4C174D11876A923FFEC15142C2F08928" box="[322,465,1556,1581]" class="Insecta" family="Gelastocoridae" genus="Gelastocoris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFEC15142C2F08928" box="[322,465,1556,1581]" italics="true" pageId="3" pageNumber="430">Gelastocoris</emphasis>
|
||
</taxonomicName>
|
||
. The habitat in which these specimens were collected was on a seepage over granite bedrock with detritus, algae, and fallen leaves (
|
||
<figureCitation id="132C2A17876A923FFD315161C1D88955" box="[690,761,1591,1616]" captionStart="FIGURE 4" captionStartId="6.[151,250,1372,1394]" captionTargetBox="[151,1436,772,1347]" captionTargetId="figure@6.[151,1436,772,1348]" captionTargetPageId="6" captionText="FIGURE 4. Type locality of Hygropetrocoris guyana n. gen. and Ambrysus brunneus n. sp. showing (A) general habitat and (B) specific habitat of H. guyana, which is in the center of the bedrock of Fig. 4 A. A. brunneus most likely was collected in small pools or rootmats of the vegetation on the left margin of the bedrock." httpUri="https://zenodo.org/record/239109/files/figure.png" pageId="3" pageNumber="430">Fig. 4</figureCitation>
|
||
). It was collected with a diverse community of hygropetric beetles including
|
||
<taxonomicName id="4C174D11876A923FFEE2510CC2D88976" box="[353,505,1626,1651]" class="Insecta" family="Dytiscidae" genus="Anodocheilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFEE2510CC2D88976" box="[353,505,1626,1651]" italics="true" pageId="3" pageNumber="430">Anodocheilus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C174D11876A923FFDB0510CC1848976" box="[563,677,1626,1651]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFDB0510CC1848976" box="[563,677,1626,1651]" italics="true" pageId="3" pageNumber="430">Copelatus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C174D11876A923FFD5E510CC7358976" authority="Miller" authorityName="Miller" box="[733,1044,1626,1651]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B963EA80876A923FFD5E510CC0E38977" box="[733,962,1626,1651]" italics="true" pageId="3" pageNumber="430">Fontidessus ornatus</emphasis>
|
||
Miller
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFBA2510CC3DE8990" authority="Miller and Montano" authorityName="Miller and Montano" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="aquarupe">
|
||
<emphasis id="B963EA80876A923FFBA2510CC63A8977" box="[1057,1307,1626,1651]" italics="true" pageId="3" pageNumber="430">Fontidessus aquarupe</emphasis>
|
||
Miller and Montano
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFE93512BC2B88993" box="[272,409,1661,1686]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFE93512BC2B88993" box="[272,409,1661,1686]" italics="true" pageId="3" pageNumber="430">Laccophilus</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="4C174D11876A923FFE59512BC1798993" box="[474,600,1661,1686]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C174D11876A923FFDF2512BC1C08991" box="[625,737,1661,1684]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFDF2512BC1C08991" box="[625,737,1661,1684]" italics="true" pageId="3" pageNumber="430">Anacaena</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C174D11876A923FFCA15128C70E8993" authority="Oliva" authorityName="Oliva" box="[802,1071,1661,1686]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="avernus">
|
||
<emphasis id="B963EA80876A923FFCA15128C0C38990" box="[802,994,1662,1685]" italics="true" pageId="3" pageNumber="430">Berosus avernus</emphasis>
|
||
Oliva
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFBC2512BC78D8993" box="[1089,1196,1661,1686]" class="Insecta" family="Hydrophilidae" genus="Enochrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFBC2512BC78D8993" box="[1089,1196,1661,1686]" italics="true" pageId="3" pageNumber="430">Enochrus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C174D11876A923FFB6E5128C6498990" box="[1261,1384,1662,1685]" class="Insecta" family="Hydrophilidae" genus="Notionotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFB6E5128C6498990" box="[1261,1384,1662,1685]" italics="true" pageId="3" pageNumber="430">Notionotus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C174D11876A923FFF1451C9C10089BD" authority="Short and Garcia" authorityName="Short and Garcia" box="[151,545,1695,1720]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="floccus">
|
||
<emphasis id="B963EA80876A923FFF1451C9C27789BD" box="[151,342,1695,1720]" italics="true" pageId="3" pageNumber="430">Oocyclus floccus</emphasis>
|
||
Short and García
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFDAF51C9C08389BD" authority="Short and Garcia" authorityName="Short and Garcia" box="[556,930,1695,1720]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="petra">
|
||
<emphasis id="B963EA80876A923FFDAF51C9C1F689BD" box="[556,727,1695,1720]" italics="true" pageId="3" pageNumber="430">Oocyclus petra</emphasis>
|
||
Short and García
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFC2D51C9C60989BD" authority="Short and Kadosoe" authorityName="Short and Kadosoe" box="[942,1320,1695,1720]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="trio">
|
||
<emphasis id="B963EA80876A923FFC2D51C9C76689BD" box="[942,1095,1695,1720]" italics="true" pageId="3" pageNumber="430">Oocyclus trio</emphasis>
|
||
Short and Kadosoe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFAB751C9C2E889DE" authority="Short and Garcia" authorityName="Short and Garcia" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="coromoto">
|
||
<emphasis id="B963EA80876A923FFAB751C9C22389DF" italics="true" pageId="3" pageNumber="430">Oocyclus coromoto</emphasis>
|
||
Short and García
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFE505195C17089DF" box="[467,593,1731,1754]" class="Insecta" family="Hydrophilidae" genus="Paracymus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFE505195C17089DF" box="[467,593,1731,1754]" italics="true" pageId="3" pageNumber="430">Paracymus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C174D11876A923FFD065194C71189DE" authority="Short and Garcia" authorityName="Short and Garcia" box="[645,1072,1730,1755]" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="sulcatus">
|
||
<emphasis id="B963EA80876A923FFD065194C04889DE" box="[645,873,1730,1755]" italics="true" pageId="3" pageNumber="430">Tobochares sulcatus</emphasis>
|
||
Short and García
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFBB85195C6BD89DF" authority="Laporte" authorityName="Laporte" box="[1083,1436,1730,1755]" class="Insecta" family="Hydrophilidae" genus="Tropisternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="chalybeus">
|
||
<emphasis id="B963EA80876A923FFBB85195C61F89DE" box="[1083,1342,1730,1755]" italics="true" pageId="3" pageNumber="430">Tropisternus chalybeus</emphasis>
|
||
Laporte
|
||
</taxonomicName>
|
||
de Castelnau (
|
||
<taxonomicName id="4C174D11876A923FFEBC51B3C2C689FB" box="[319,487,1765,1790]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
|
||
), and
|
||
<taxonomicName id="4C174D11876A923FFDB051B3C78B89FB" authority="J. Balfour-Browne" authorityName="J. Balfour-Browne" box="[563,1194,1765,1790]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B963EA80876A923FFDB051B3C00B89FB" box="[563,810,1765,1790]" italics="true" pageId="3" pageNumber="430">Liocanthydrus clayae</emphasis>
|
||
(J. Balfour-Browne) (Noteridae)
|
||
</taxonomicName>
|
||
(A.E.Z. Short, pers. com., field notes and database at http://creac.kubiodiversityinstitute.org/collections). Other naucorid genera with similar habitat associations have been described recently. Specifically, in Indochina,
|
||
<taxonomicName id="4C174D11876A923FFBE3507CC7DC8846" box="[1120,1277,1834,1859]" class="Insecta" family="Naucoridae" genus="Namtokocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFBE3507CC7DC8846" box="[1120,1277,1834,1859]" italics="true" pageId="3" pageNumber="430">Namtokocoris</emphasis>
|
||
</taxonomicName>
|
||
can be found completely immersed beneath sheeting water of waterfalls but also on wet rock surfaces that only are sprayed with mist from the waterfall (
|
||
<bibRefCitation id="EF864B63876A923FFE335039C1C1888D" author="Sites" box="[432,736,1903,1928]" pageId="3" pageNumber="430" refString="Sites, R. W. & Vitheepradit, A. (2007) Namtokocoris, a new genus of Naucoridae (Hemiptera: Heteroptera) in waterfalls of Indochina, with descriptions of six new species. Zootaxa, 1588, 1 - 29." type="journal article" year="2007">Sites & Vitheepradit 2007</bibRefCitation>
|
||
). In
|
||
<collectingCountry id="F3007602876A923FFC965039C070888D" box="[789,849,1903,1928]" name="India" pageId="3" pageNumber="430">India</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C174D11876A923FFCDD5039C723888D" box="[862,1026,1903,1928]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFCDD5039C723888D" box="[862,1026,1903,1928]" italics="true" pageId="3" pageNumber="430">Diaphorocoris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C174D11876A923FFBBE5039C7CE888D" box="[1085,1263,1903,1928]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EA80876A923FFBBE5039C7CE888D" box="[1085,1263,1903,1928]" italics="true" pageId="3" pageNumber="430">Pogonocaudina</emphasis>
|
||
</taxonomicName>
|
||
were collected out of water on wet rock surfaces during the dry season (
|
||
<bibRefCitation id="EF864B63876A923FFCA250C4C0FF88AE" author="Sites" box="[801,990,1938,1963]" pageId="3" pageNumber="430" refString="Sites, R. W., Zettel, H. & Arunachalam, M. (2011) Waterfall-inhabiting Naucoridae (Hemiptera: Heteroptera) of southern India and Sri Lanka: Pogonocaudina indica n. gen. and n. sp., and a review of Diaphorocoris with descriptions of two new species. Zootaxa, 2760, 1 - 17." type="journal article" year="2011">
|
||
Sites
|
||
<emphasis id="B963EA80876A923FFCE250C5C0BA88AE" box="[865,923,1938,1963]" italics="true" pageId="3" pageNumber="430">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
), although they might also occur in the film of water of waterfalls at times of the year when more water is available. All of the known Old World genera that have adapted to waterfall or hygropetric habitats are in the subfamily
|
||
<taxonomicName id="4C174D11876A923FFBAF5081C7E688F5" box="[1068,1223,2007,2032]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
|
||
. In Mesoamerica,
|
||
<emphasis id="B963EA80876D9238FF1457CEC2608FAA" box="[151,321,152,175]" italics="true" pageId="4" pageNumber="431">Cataractocoris</emphasis>
|
||
of the subfamily
|
||
<taxonomicName id="4C174D11876D9238FD8457C1C1958FB5" box="[519,692,151,176]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
|
||
occurs in waterfalls and other torrential habitats.
|
||
<emphasis id="B963EA80876D9238FB6057C1C65E8FB5" box="[1251,1407,151,176]" italics="true" pageId="4" pageNumber="431">Carvalhoiella</emphasis>
|
||
in South
|
||
<collectingCountry id="F3007602876D9238FF6257ECC2638FD6" box="[225,322,186,211]" name="United States of America" pageId="4" pageNumber="431">America</collectingCountry>
|
||
has been recorded from crevices in rocks of waterfalls (
|
||
<bibRefCitation id="EF864B63876D9238FC4557ECC7B68FD6" author="Nieser" box="[966,1175,186,211]" pageId="4" pageNumber="431" refString="Nieser, N., Pelli, A. & Melo, A. L. (1999) Two new Ambrysinae (Heteroptera: Naucoridae) from Minas Gerais, Brazil. Acta Societatis Zoologicae Bohemoslovacae, 63, 157 - 163." type="journal article" year="1999">
|
||
Nieser
|
||
<emphasis id="B963EA80876D9238FB9A57EAC7728FD6" box="[1049,1107,186,211]" italics="true" pageId="4" pageNumber="431">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
), and Daniel Reynoso- Velasco and I have collected
|
||
<taxonomicName id="4C174D11876D9238FE5D5788C06B8FF3" authority="Usinger" authorityName="Usinger" box="[478,842,221,246]" class="Insecta" family="Noteridae" genus="Interocoris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B963EA80876D9238FE5D5788C1F78FF0" box="[478,726,222,245]" italics="true" pageId="4" pageNumber="431">Interocoris mexicanus</emphasis>
|
||
(Usinger)
|
||
</taxonomicName>
|
||
in similar habitats in Chiapas,
|
||
<collectingCountry id="F3007602876D9238FB24578BC7DE8FF3" box="[1191,1279,221,246]" name="Mexico" pageId="4" pageNumber="431">Mexico</collectingCountry>
|
||
, although this species was described from a muddy river bank (
|
||
<bibRefCitation id="EF864B63876D9238FD3957A9C0768E1D" author="Usinger" box="[698,855,255,280]" pageId="4" pageNumber="431" refString="Usinger, R. L. (1935) A second American species of the naucorid subfamily Laccocorinae (Hemiptera). Revista de Entomologia, 5 (2), 133 - 136." type="journal article" year="1935">Usinger 1935</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6519876D9238FF445674C0118E5A" pageId="4" pageNumber="431" type="etymology">
|
||
<paragraph id="8BA83692876D9238FF445674C0118E5A" blockId="4.[151,1436,151,387]" pageId="4" pageNumber="431">
|
||
<emphasis id="B963EA80876D9238FF445674C2668E3E" bold="true" box="[199,327,290,315]" pageId="4" pageNumber="431">Etymology</emphasis>
|
||
. The generic name "
|
||
<taxonomicName id="4C174D11876D9238FDAD5672C1C78E3E" box="[558,742,292,315]" class="Insecta" family="Naucoridae" genus="Hygropetrocoris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B963EA80876D9238FDAD5672C1C78E3E" box="[558,742,292,315]" italics="true" pageId="4" pageNumber="431">Hygropetrocoris</emphasis>
|
||
</taxonomicName>
|
||
" is in reference to the bug's wet rock habitat and derives from Greek with
|
||
<emphasis id="B963EA80876D9238FE985610C27A8E5A" box="[283,347,326,351]" italics="true" pageId="4" pageNumber="431">hygro</emphasis>
|
||
for wet,
|
||
<emphasis id="B963EA80876D9238FE435611C2DB8E5B" box="[448,506,327,350]" italics="true" pageId="4" pageNumber="431">petro</emphasis>
|
||
for rock, and
|
||
<emphasis id="B963EA80876D9238FD1B5611C1EE8E5B" box="[664,719,327,350]" italics="true" pageId="4" pageNumber="431">coris</emphasis>
|
||
for bug.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6519876D9238FF44563CC18A8E86" box="[199,683,362,387]" pageId="4" pageNumber="431" type="materials_examined">
|
||
<paragraph id="8BA83692876D9238FF44563CC18A8E86" blockId="4.[151,1436,151,387]" box="[199,683,362,387]" pageId="4" pageNumber="431">
|
||
<emphasis id="B963EA80876D9238FF44563CC2918E86" bold="true" box="[199,432,362,387]" pageId="4" pageNumber="431">Material examined.</emphasis>
|
||
See species localities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |