treatments-xml/data/39/49/4C/39494C26FFE09F2BA2B9F8D9FEAA15C3.xml
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<document id="7F876A648581798C5C81E3417CBB73CD" ID-ISSN="1536-9307" ID-ZooBank="BE84880D-BA36-46FE-B3D4-FA85C52074A8" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638251163817" checkinUser="felipe" docAuthor="Trujillo, Rony E. &amp; de Armas, Luis F." docDate="2016" docId="39494C26FFE09F2BA2B9F8D9FEAA15C3" docLanguage="en" docName="Euscorpius.233.1-8.pdf" docOrigin="Euscorpius 233" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Centruroides ixil Trujillo &amp; de Armas 2016, sp. n." docType="treatment" docUuid="1C2CB6B8-46EA-45C3-BA07-D446D0D93F95" docUuidSource="ZooBank" docVersion="3" lastPageNumber="7" masterDocId="C570345EFFE29F23A148FFD3FFB61348" masterDocTitle="A new species of Centruroides (Scorpiones: Buthidae) from Quiché, northwestern Guatemala" masterLastPageNumber="8" masterPageNumber="1" pageNumber="1" updateTime="1699435380254" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="E0CC3146179A01E1EB279BF384565920">A new species of Centruroides (Scorpiones: Buthidae) from Quiché, northwestern Guatemala</mods:title>
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<mods:namePart id="EDA5CD2392FB1E976F8A001A2AE202B8">Trujillo, Rony E.</mods:namePart>
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<mods:namePart id="E031899F424CCFEC359DDE42F2794799">de Armas, Luis F.</mods:namePart>
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<mods:date id="7C7670A6E5BEB1E2943B6659604F639C">2016</mods:date>
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<subSubSection id="F9FAAEBBFFE09F21A2B9F8D9FB58146A" box="[1009,1262,1802,1826]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="B15FFD30FFE09F21A2B9F8D9FB58146A" blockId="2.[831,1440,1802,1918]" box="[1009,1262,1802,1826]" pageId="2" pageNumber="1">
<heading id="EA174A5CFFE09F21A2B9F8D9FB58146A" box="[1009,1262,1802,1826]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1" reason="2">
<taxonomicName id="76E086B3FFE09F21A2B9F8D9FB1C146A" authority="Trujillo &amp; de Armas, 2016" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[1009,1194,1802,1826]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="ixil" status="sp. nov.">Centruroides ixil</taxonomicName>
<taxonomicNameLabel id="98A79C59FFE09F21A5F9F8D8FB58146A" box="[1201,1262,1803,1826]" pageId="2" pageNumber="1" rank="species">
<emphasis id="83942122FFE09F21A5F9F8D8FB51146A" bold="true" box="[1201,1255,1803,1826]" pageId="2" pageNumber="1">sp. n</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F9FAAEBBFFE09F2BA2EAF8FAFEAA15C3" lastPageId="8" lastPageNumber="7" pageId="2" pageNumber="1" type="description">
<paragraph id="B15FFD30FFE09F21A2EAF8FAFA161417" blockId="2.[831,1440,1802,1918]" pageId="2" pageNumber="1">
<figureCitation id="29DBE1B5FFE09F21A2EAF8FAFBAA1408" box="[930,1052,1833,1856]" captionStart="Figure 1" captionStartId="3.[192,268,1350,1373]" captionTargetBox="[198,1440,189,1320]" captionTargetId="figure-6@3.[192,1440,188,1320]" captionTargetPageId="3" captionText="Figure 1: A, geographical distribution of Centruroides ixil sp. n. (red star) and C. caral (red circle). BC, male holotype of Centruroides ixil sp. n. in dorsal (B) and ventral (C) aspect." pageId="2" pageNumber="1">Fig. 1 AC</figureCitation>
,
<figureCitation id="29DBE1B5FFE09F21A56FF8FAFBC71408" box="[1063,1137,1833,1856]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="2" pageNumber="1">2 AG</figureCitation>
,
<figureCitation id="29DBE1B5FFE09F21A534F8FAFB701408" box="[1148,1222,1833,1856]" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="2" pageNumber="1">3 AG</figureCitation>
,
<figureCitation id="29DBE1B5FFE09F21A599F8FAFB561408" box="[1233,1248,1833,1856]" captionStart="Figure 4" captionStartId="7.[192,268,1276,1299]" captionTargetBox="[204,1429,167,1244]" captionTargetId="figure-6@7.[204,1428,168,1243]" captionTargetPageId="7" captionText="Figure 4: Habitat of Centruroides ixil sp. n. (photos by Andrea Martinez)." pageId="2" pageNumber="1">4</figureCitation>
,
<tableCitation id="FC62C88BFFE09F21A5A4F8FAFA8A1408" box="[1260,1340,1833,1856]" captionStart="Table 1" captionStartId="6.[192,257,834,857]" captionText="Table 1: Measurements (mm) of Centruroides ixil sp. n. D, depth; L, length; W, width." pageId="2" pageNumber="1">Table 1</tableCitation>
<emphasis id="83942122FFE09F21A277F89BFA161417" bold="true" box="[831,1440,1864,1887]" pageId="2" pageNumber="1">http://zoobank.org/urn:lsid:zoobank.org:act:1C2CB6</emphasis>
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<emphasis id="83942122FFE09F21A2DEF8B4FAFE1436" bold="true" box="[918,1352,1895,1918]" pageId="2" pageNumber="1">B8-46EA-45C3-BA07-D446D0D93F95</emphasis>
</paragraph>
<caption id="E59FADB8FFE19F20A188FA95FD4A163D" pageId="3" pageNumber="2" startId="3.[192,268,1350,1373]" targetBox="[198,1440,189,1320]" targetPageId="3">
<paragraph id="B15FFD30FFE19F20A188FA95FD4A163D" blockId="3.[192,1440,1350,1397]" pageId="3" pageNumber="2">
<emphasis id="83942122FFE19F20A188FA95FE9D1615" bold="true" box="[192,299,1350,1373]" pageId="3" pageNumber="2">Figure 1:</emphasis>
A, geographical distribution of
<taxonomicName id="76E086B3FFE19F20A324FA94FCA41614" authority="Trujillo &amp; de Armas, 2016" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[620,786,1351,1372]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="ixil" status="sp. nov.">
<emphasis id="83942122FFE19F20A324FA94FCA41614" box="[620,786,1351,1372]" italics="true" pageId="3" pageNumber="2">Centruroides ixil</emphasis>
</taxonomicName>
<emphasis id="83942122FFE19F20A252FA94FCE51614" bold="true" box="[794,851,1351,1372]" pageId="3" pageNumber="2">
<taxonomicNameLabel id="98A79C59FFE19F20A252FA94FCE51614" box="[794,851,1351,1372]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(red star) and
<taxonomicName id="76E086B3FFE19F20A2A1FA94FB8C1614" authorityName="Armas et Trujillo" authorityYear="2015" box="[1001,1082,1351,1372]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFE19F20A2A1FA94FB8C1614" box="[1001,1082,1351,1372]" italics="true" pageId="3" pageNumber="2">C. caral</emphasis>
</taxonomicName>
(red circle). BC, male holotype of
<taxonomicName id="76E086B3FFE19F20A188FAB3FED2163D" authority="Trujillo &amp; de Armas, 2016" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[192,356,1376,1397]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="ixil" status="sp. nov.">
<emphasis id="83942122FFE19F20A188FAB3FED2163D" box="[192,356,1376,1397]" italics="true" pageId="3" pageNumber="2">Centruroides ixil</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98A79C59FFE19F20A021FAB3FE2B163D" box="[361,413,1376,1397]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
in dorsal (B) and ventral (C) aspect.
</paragraph>
</caption>
<paragraph id="B15FFD30FFE19F20A1B8FA79FE901574" blockId="3.[192,802,1450,1903]" pageId="3" pageNumber="2">
<emphasis id="83942122FFE19F20A1B8FA79FEED1689" box="[240,347,1450,1473]" italics="true" pageId="3" pageNumber="2">Type data</emphasis>
.
<materialsCitation id="0188F76DFFE19F20A023FA79FE971574" collectingDate="2016-09-02" collectionCode="UV" collectorName="Luis A. Trujillo" country="Guatemala" county="Chajul Municipality" elevation="1173" latitude="15.61775" location="Male" longLatPrecision="1" longitude="-91.09745" municipality="Sotzil Village" pageId="3" pageNumber="2" specimenCount="2" stateProvince="Quiche" typeStatus="holotype">
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<typeStatus id="6E5B4392FFE19F20A0E2FA79FDBC1689" box="[426,522,1450,1473]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
, female
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(MHN),
<collectingMunicipality id="513B674AFFE19F20A188FA1AFEEE16A8" box="[192,344,1481,1504]" pageId="3" pageNumber="2">Sotzil Village</collectingMunicipality>
(
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, -
<geoCoordinate id="D4D49BF7FFE19F20A35AFA1AFD2916A8" box="[530,671,1481,1504]" degrees="91.09745" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="1" value="-91.09745">91.09745 W</geoCoordinate>
;
<elevation id="3ACD1A03FFE19F20A3E6FA1AFF6816B7" metricMagnitude="3" metricUnit="m" metricValue="1.173" pageId="3" pageNumber="2" unit="m" value="1173.0">
<quantity id="761850D5FFE19F20A3E6FA1AFCB216A8" box="[686,772,1481,1504]" metricMagnitude="3" metricUnit="m" metricValue="1.173" pageId="3" pageNumber="2" unit="m" value="1173.0">1173 m</quantity>
a. s.l.
</elevation>
),
<collectingCounty id="583E85BCFFE19F20A1B5FA3BFE6916B7" box="[253,479,1512,1535]" pageId="3" pageNumber="2">Chajul Municipality</collectingCounty>
,
<collectingRegion id="732433D2FFE19F20A0B8FA3BFD7B16B7" box="[496,717,1512,1535]" country="Guatemala" name="Quiche" pageId="3" pageNumber="2">Quiché Department</collectingRegion>
,
<collectingCountry id="C9F7BDA0FFE19F20A3A4FA3BFEBC1555" name="Guatemala" pageId="3" pageNumber="2">Guatemala</collectingCountry>
,
<date id="C55EDBF0FFE19F20A052F9D5FE461555" box="[282,496,1542,1565]" pageId="3" pageNumber="2" value="2016-09-02">
<collectingDate id="D51A2218FFE19F20A052F9D5FE461555" box="[282,496,1542,1565]" pageId="3" pageNumber="2" value="2016-09-02">September 2, 2016</collectingDate>
</date>
,
<collectorName id="1C1598E6FFE19F20A348F9D5FD031555" box="[512,693,1542,1565]" pageId="3" pageNumber="2">Luis A. Trujillo</collectorName>
, at night,
<collectionCode id="D7F165F5FFE19F20A188F9F6FF501574" box="[192,230,1573,1596]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
light
</materialsCitation>
.
</paragraph>
<paragraph id="B15FFD30FFE19F20A1B8F997FE871531" blockId="3.[192,802,1450,1903]" pageId="3" pageNumber="2">
<emphasis id="83942122FFE19F20A1B8F997FEC41513" box="[240,370,1604,1627]" italics="true" pageId="3" pageNumber="2">Distribution</emphasis>
. Only known from the
<typeStatus id="6E5B4392FFE19F20A3DDF997FD751513" box="[661,707,1604,1627]" pageId="3" pageNumber="2">type</typeStatus>
locality (
<figureCitation id="29DBE1B5FFE19F20A180F9B1FE931531" box="[200,293,1634,1657]" captionStart="Figure 1" captionStartId="3.[192,268,1350,1373]" captionTargetBox="[198,1440,189,1320]" captionTargetId="figure-6@3.[192,1440,188,1320]" captionTargetPageId="3" captionText="Figure 1: A, geographical distribution of Centruroides ixil sp. n. (red star) and C. caral (red circle). BC, male holotype of Centruroides ixil sp. n. in dorsal (B) and ventral (C) aspect." pageId="3" pageNumber="2">Fig. 1 A</figureCitation>
).
</paragraph>
<paragraph id="B15FFD30FFE19F20A1B8F952FDDE159D" blockId="3.[192,802,1450,1903]" pageId="3" pageNumber="2">
<emphasis id="83942122FFE19F20A1B8F952FED415D0" box="[240,354,1665,1688]" italics="true" pageId="3" pageNumber="2">Etymology</emphasis>
. The specific name is a noun in apposition, designating the predominant Mayan ethnic group in the region where this new species lives.
</paragraph>
<paragraph id="B15FFD30FFE19F25A1B8F90EFD2310B0" blockId="3.[192,802,1450,1903]" lastBlockId="6.[192,802,901,1917]" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2">
<emphasis id="83942122FFE19F20A1B8F90EFEEA15BC" box="[240,348,1757,1780]" italics="true" pageId="3" pageNumber="2">Diagnosis</emphasis>
.
<materialsCitation id="0188F76DFFE19F25A038F90EFF6210F2" collectionCode="VI" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
Male
<quantity id="761850D5FFE19F20A0FBF90EFDAE15BC" box="[435,536,1757,1780]" metricMagnitude="-2" metricUnit="m" metricValue="6.85" pageId="3" pageNumber="2" unit="mm" value="68.5">68.5 mm</quantity>
in total length; female
<quantity id="761850D5FFE19F20A188F92FFE94145B" box="[192,290,1788,1811]" metricMagnitude="-2" metricUnit="m" metricValue="5.66" pageId="3" pageNumber="2" unit="mm" value="56.6">56.6 mm</quantity>
. Base color light brownish; darker on last metasomal segments and posterior margin of tergites I
<collectionCode id="D7F165F5FFE19F20A248F8C9FCA81479" box="[768,798,1818,1841]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="2">VI</collectionCode>
; cheliceral manus with dark brown reticulations all over; pedipalp chelae with light yellowish manus and blackish fingers; sternites III
<collectionCode id="D7F165F5FFE19F20A564FA79FBFE1689" box="[1068,1096,1450,1473]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="2">VI</collectionCode>
brown yellowish in the male (darker on the female). Carapace finely granular, with moderate granules on the interocular triangle; anteromedian notch V-shaped. Pedipalp manus slightly wider (
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,
<figureCitation id="29DBE1B5FFE19F20A227F9F6FC621574" box="[879,980,1573,1596]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="3" pageNumber="2">Fig. 2 B</figureCitation>
) or slender (
<specimenCount id="A7E636B9FFE19F20A539F9F6FB331574" box="[1137,1157,1573,1596]" pageId="3" pageNumber="2" type="female"></specimenCount>
,
<figureCitation id="29DBE1B5FFE19F20A5D2F9F6FAB61574" box="[1178,1280,1573,1596]" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="3" pageNumber="2">Fig. 3 D</figureCitation>
) than patella; movable finger with moderately developed basal lobe. Pectines with 19 (
<specimenCount id="A7E636B9FFE19F20A54BF9B1FBA11531" box="[1027,1047,1634,1657]" pageId="3" pageNumber="2" type="female"></specimenCount>
,
<figureCitation id="29DBE1B5FFE19F20A56EF9B1FBC91531" box="[1062,1151,1634,1657]" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="3" pageNumber="2">Fig. 3 E</figureCitation>
) or 20 (
<specimenCount id="A7E636B9FFE19F20A591F9B1FB5B1531" box="[1241,1261,1634,1657]" pageId="3" pageNumber="2" type="male"></specimenCount>
,
<figureCitation id="29DBE1B5FFE19F20A5B4F9B1FAE31531" box="[1276,1365,1634,1657]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="3" pageNumber="2">Fig. 2 E</figureCitation>
) teeth; basal plate rectangular, wider than long. Metasoma: segments IIV with dorsolateral carinae, lateral supramedian carinae, ventrolateral carinae and ventral submedian carinae moderately developed, granular to slightly serrate; segments IIIV with two pairs of ventrolateral macrochaetae; intercarinal spaces IIV coriaceous; vesicle deeper than wide; subaculear tubercle strong, spine-like, with tip pointing towards the apex of the aculeus. Metasomal segment III length/width ratio:
<specimenCount id="A7E636B9FFE49F25A188FC70FF6210F2" box="[192,212,931,954]" pageId="6" pageNumber="5" type="female"></specimenCount>
</materialsCitation>
= 1.86,
<specimenCount id="A7E636B9FFE49F25A067FC70FEF510F2" box="[303,323,931,954]" pageId="6" pageNumber="5" type="male"></specimenCount>
=
<materialsCitation id="0188F76DFFE49F25A028FC70FEAE1091" collectionCode="V" pageId="6" pageNumber="5" specimenCount="1" specimenCount-female="1">
2.76; metasomal segment
<collectionCode id="D7F165F5FFE49F25A331FC70FD2610F2" box="[633,656,931,954]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="5" type="Museum">V</collectionCode>
: length/width ratio:
<specimenCount id="A7E636B9FFE49F25A04CFC11FEAE1091" box="[260,280,962,985]" pageId="6" pageNumber="5" type="female"></specimenCount>
</materialsCitation>
= 2.43,
<specimenCount id="A7E636B9FFE49F25A037FC11FE251091" box="[383,403,962,985]" pageId="6" pageNumber="5" type="male"></specimenCount>
=
<materialsCitation id="0188F76DFFE49F25A0F2FC11FE6010B0" collectionCode="V" pageId="6" pageNumber="5" specimenCount="1" specimenCount-female="1">
2.78; carapace length/metasomal segment
<collectionCode id="D7F165F5FFE49F25A068FC32FE8510B0" box="[288,307,993,1016]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="5" type="Museum">V</collectionCode>
length ratio:
<specimenCount id="A7E636B9FFE49F25A08AFC32FE6010B0" box="[450,470,993,1016]" pageId="6" pageNumber="5" type="female"></specimenCount>
</materialsCitation>
= 0.88,
<specimenCount id="A7E636B9FFE49F25A366FC32FDF410B0" box="[558,578,993,1016]" pageId="6" pageNumber="5" type="male"></specimenCount>
= 0.68.
</paragraph>
<caption id="E59FADB8FFE69F27A188F892FA9B1439" pageId="4" pageNumber="3" startId="4.[192,268,1857,1880]" targetBox="[251,1393,191,1828]" targetPageId="4">
<paragraph id="B15FFD30FFE69F27A188F892FA9B1439" blockId="4.[192,1440,1857,1905]" pageId="4" pageNumber="3">
<emphasis id="83942122FFE69F27A188F892FE9A1410" bold="true" box="[192,300,1857,1880]" pageId="4" pageNumber="3">Figure 2:</emphasis>
<taxonomicName id="76E086B3FFE69F27A07DF890FE6D1410" authority="Trujillo &amp; de Armas, 2016" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[309,475,1859,1880]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="ixil" status="sp. nov.">
<emphasis id="83942122FFE69F27A07DF890FE6D1410" box="[309,475,1859,1880]" italics="true" pageId="4" pageNumber="3">Centruroides ixil</emphasis>
</taxonomicName>
<emphasis id="83942122FFE69F27A0ACF890FDAB1410" bold="true" box="[484,541,1859,1880]" pageId="4" pageNumber="3">
<taxonomicNameLabel id="98A79C59FFE69F27A0ACF890FDAB1410" box="[484,541,1859,1880]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect.
</paragraph>
</caption>
<caption id="E59FADB8FFE79F26A188F8E5FEEA1436" pageId="5" pageNumber="4" startId="5.[192,268,1846,1869]" targetBox="[191,1440,191,1816]" targetPageId="5">
<paragraph id="B15FFD30FFE79F26A188F8E5FEEA1436" blockId="5.[192,1440,1846,1918]" pageId="5" pageNumber="4">
<emphasis id="83942122FFE79F26A188F8E5FE9F1405" bold="true" box="[192,297,1846,1869]" pageId="5" pageNumber="4">Figure 3:</emphasis>
<taxonomicName id="76E086B3FFE79F26A067F8E4FE651404" authority="Trujillo &amp; de Armas, 2016" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[303,467,1847,1868]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="ixil" status="sp. nov.">
<emphasis id="83942122FFE79F26A067F8E4FE651404" box="[303,467,1847,1868]" italics="true" pageId="5" pageNumber="4">Centruroides ixil</emphasis>
</taxonomicName>
<emphasis id="83942122FFE79F26A091F8E4FDB91404" bold="true" box="[473,527,1847,1868]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="98A79C59FFE79F26A091F8E4FDB91404" box="[473,527,1847,1868]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI.
</paragraph>
</caption>
<paragraph id="B15FFD30FFE49F25A085FF1AFBD4106B" pageId="6" pageNumber="5">
<table id="C3E00F90FFE460DCA0D9FF1AFB24106B" box="[401,1170,201,803]" gridcols="3" gridrows="19" pageId="6" pageNumber="5">
<tr id="0FD0FF72FFE460DCA0D9FF1AFB2413AB" box="[401,1170,201,227]" gridrow="0" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FF1AFD2E13AB" box="[401,664,201,227]" gridcol="0" gridrow="0" pageId="6" pageNumber="5">
<emphasis id="83942122FFE49F25A085FF1AFDEF13AB" bold="true" box="[461,601,201,227]" pageId="6" pageNumber="5">Characters</emphasis>
</th>
<th id="4C01960EFFE460DCA3BFFF1AFC3013AB" box="[759,902,201,227]" gridcol="1" gridrow="0" pageId="6" pageNumber="5">
<emphasis id="83942122FFE49F25A25CFF1AFC3013AB" bold="true" box="[788,902,201,227]" pageId="6" pageNumber="5">Holotype</emphasis>
</th>
<th id="4C01960EFFE460DCA54AFF1AFB2413AB" box="[1026,1170,201,227]" gridcol="2" gridrow="0" pageId="6" pageNumber="5">
<emphasis id="83942122FFE49F25A568FF1AFB2413AB" bold="true" box="[1056,1170,201,227]" pageId="6" pageNumber="5">Paratype</emphasis>
</th>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FF38FB24124A" box="[401,1170,235,258]" gridrow="1" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FF38FD2E124A" box="[401,664,235,258]" gridcol="0" gridrow="1" pageId="6" pageNumber="5">Carapace, L/posterior W</th>
<td id="4C01960EFFE460DCA3BFFF38FC30124A" box="[759,902,235,258]" gridcol="1" gridrow="1" pageId="6" pageNumber="5">6.1/5.9</td>
<td id="4C01960EFFE460DCA54AFF38FB24124A" box="[1026,1170,235,258]" gridcol="2" gridrow="1" pageId="6" pageNumber="5">6.0/6.0</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FED8FB24126A" box="[401,1170,267,290]" gridrow="2" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FED8FD2E126A" box="[401,664,267,290]" gridcol="0" gridrow="2" pageId="6" pageNumber="5">Pedipalp, L</th>
<td id="4C01960EFFE460DCA3BFFED8FC30126A" box="[759,902,267,290]" gridcol="1" gridrow="2" pageId="6" pageNumber="5">25.15</td>
<td id="4C01960EFFE460DCA54AFED8FB24126A" box="[1026,1170,267,290]" gridcol="2" gridrow="2" pageId="6" pageNumber="5">21.9</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FEF8FB24120A" box="[401,1170,299,322]" gridrow="3" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FEF8FD2E120A" box="[401,664,299,322]" gridcol="0" gridrow="3" pageId="6" pageNumber="5">Femur, L/W</th>
<td id="4C01960EFFE460DCA3BFFEF8FC30120A" box="[759,902,299,322]" gridcol="1" gridrow="3" pageId="6" pageNumber="5">6.75/1.5</td>
<td id="4C01960EFFE460DCA54AFEF8FB24120A" box="[1026,1170,299,322]" gridcol="2" gridrow="3" pageId="6" pageNumber="5">5.85/1.5</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FE98FB24122A" box="[401,1170,331,354]" gridrow="4" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FE98FD2E122A" box="[401,664,331,354]" gridcol="0" gridrow="4" pageId="6" pageNumber="5">Patella, L/W</th>
<td id="4C01960EFFE460DCA3BFFE98FC30122A" box="[759,902,331,354]" gridcol="1" gridrow="4" pageId="6" pageNumber="5">7.1/2.2</td>
<td id="4C01960EFFE460DCA54AFE98FB24122A" box="[1026,1170,331,354]" gridcol="2" gridrow="4" pageId="6" pageNumber="5">6.0/2.1</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FEB8FB2412CA" box="[401,1170,363,386]" gridrow="5" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FEB8FD2E12CA" box="[401,664,363,386]" gridcol="0" gridrow="5" pageId="6" pageNumber="5">Chela, L/W/D</th>
<td id="4C01960EFFE460DCA3BFFEB8FC3012CA" box="[759,902,363,386]" gridcol="1" gridrow="5" pageId="6" pageNumber="5">11.3/2.9/2.8</td>
<td id="4C01960EFFE460DCA54AFEB8FB2412CA" box="[1026,1170,363,386]" gridcol="2" gridrow="5" pageId="6" pageNumber="5">10.05/2.0/1.9</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FE58FB2412EA" box="[401,1170,395,418]" gridrow="6" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FE58FD2E12EA" box="[401,664,395,418]" gridcol="0" gridrow="6" pageId="6" pageNumber="5">Underhand, L</th>
<td id="4C01960EFFE460DCA3BFFE58FC3012EA" box="[759,902,395,418]" gridcol="1" gridrow="6" pageId="6" pageNumber="5">4.5</td>
<td id="4C01960EFFE460DCA54AFE58FB2412EA" box="[1026,1170,395,418]" gridcol="2" gridrow="6" pageId="6" pageNumber="5">3.5</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FE78FB24128A" box="[401,1170,427,450]" gridrow="7" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FE78FD2E128A" box="[401,664,427,450]" gridcol="0" gridrow="7" pageId="6" pageNumber="5">Movable finger, L</th>
<td id="4C01960EFFE460DCA3BFFE78FC30128A" box="[759,902,427,450]" gridcol="1" gridrow="7" pageId="6" pageNumber="5">6.8</td>
<td id="4C01960EFFE460DCA54AFE78FB24128A" box="[1026,1170,427,450]" gridcol="2" gridrow="7" pageId="6" pageNumber="5">6.55</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FE18FB2412AA" box="[401,1170,459,482]" gridrow="8" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FE18FD2E12AA" box="[401,664,459,482]" gridcol="0" gridrow="8" pageId="6" pageNumber="5">Mesosoma, L</th>
<td id="4C01960EFFE460DCA3BFFE18FC3012AA" box="[759,902,459,482]" gridcol="1" gridrow="8" pageId="6" pageNumber="5">17.5</td>
<td id="4C01960EFFE460DCA54AFE18FB2412AA" box="[1026,1170,459,482]" gridcol="2" gridrow="8" pageId="6" pageNumber="5">17.3</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FE38FB24114A" box="[401,1170,491,514]" gridrow="9" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FE38FD2E114A" box="[401,664,491,514]" gridcol="0" gridrow="9" pageId="6" pageNumber="5">Tergite VII, L/W</th>
<td id="4C01960EFFE460DCA3BFFE38FC30114A" box="[759,902,491,514]" gridcol="1" gridrow="9" pageId="6" pageNumber="5">5.0/5.9</td>
<td id="4C01960EFFE460DCA54AFE38FB24114A" box="[1026,1170,491,514]" gridcol="2" gridrow="9" pageId="6" pageNumber="5">4.5/6.4</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FDD8FB24116A" box="[401,1170,523,546]" gridrow="10" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FDD8FD2E116A" box="[401,664,523,546]" gridcol="0" gridrow="10" pageId="6" pageNumber="5">Metasoma, L</th>
<td id="4C01960EFFE460DCA3BFFDD8FC30116A" box="[759,902,523,546]" gridcol="1" gridrow="10" pageId="6" pageNumber="5">44.9</td>
<td id="4C01960EFFE460DCA54AFDD8FB24116A" box="[1026,1170,523,546]" gridcol="2" gridrow="10" pageId="6" pageNumber="5">33.3</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FDF8FB24110A" box="[401,1170,555,578]" gridrow="11" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FDF8FD2E110A" box="[401,664,555,578]" gridcol="0" gridrow="11" pageId="6" pageNumber="5">I, L/W/D</th>
<td id="4C01960EFFE460DCA3BFFDF8FC30110A" box="[759,902,555,578]" gridcol="1" gridrow="11" pageId="6" pageNumber="5">5.8/3.1/2.8</td>
<td id="4C01960EFFE460DCA54AFDF8FB24110A" box="[1026,1170,555,578]" gridcol="2" gridrow="11" pageId="6" pageNumber="5">4.3/3.4/2.8</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FD98FB24112A" box="[401,1170,587,610]" gridrow="12" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FD98FD2E112A" box="[401,664,587,610]" gridcol="0" gridrow="12" pageId="6" pageNumber="5">II, L/W</th>
<td id="4C01960EFFE460DCA3BFFD98FC30112A" box="[759,902,587,610]" gridcol="1" gridrow="12" pageId="6" pageNumber="5">7.2/3.0</td>
<td id="4C01960EFFE460DCA54AFD98FB24112A" box="[1026,1170,587,610]" gridcol="2" gridrow="12" pageId="6" pageNumber="5">5.2/3.2</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FDB8FB2411CA" box="[401,1170,619,642]" gridrow="13" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FDB8FD2E11CA" box="[401,664,619,642]" gridcol="0" gridrow="13" pageId="6" pageNumber="5">III, L/W</th>
<td id="4C01960EFFE460DCA3BFFDB8FC3011CA" box="[759,902,619,642]" gridcol="1" gridrow="13" pageId="6" pageNumber="5">8.0/2.9</td>
<td id="4C01960EFFE460DCA54AFDB8FB2411CA" box="[1026,1170,619,642]" gridcol="2" gridrow="13" pageId="6" pageNumber="5">5.6/3.0</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FD58FB2411EA" box="[401,1170,651,674]" gridrow="14" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FD58FD2E11EA" box="[401,664,651,674]" gridcol="0" gridrow="14" pageId="6" pageNumber="5">IV, L/W</th>
<td id="4C01960EFFE460DCA3BFFD58FC3011EA" box="[759,902,651,674]" gridcol="1" gridrow="14" pageId="6" pageNumber="5">8.3/2.8</td>
<td id="4C01960EFFE460DCA54AFD58FB2411EA" box="[1026,1170,651,674]" gridcol="2" gridrow="14" pageId="6" pageNumber="5">6.0/2.9</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FD78FB24118A" box="[401,1170,683,706]" gridrow="15" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FD78FD2E118A" box="[401,664,683,706]" gridcol="0" gridrow="15" pageId="6" pageNumber="5">V, L/W/D</th>
<td id="4C01960EFFE460DCA3BFFD78FC30118A" box="[759,902,683,706]" gridcol="1" gridrow="15" pageId="6" pageNumber="5">8.9/3.2/3.3</td>
<td id="4C01960EFFE460DCA54AFD78FB24118A" box="[1026,1170,683,706]" gridcol="2" gridrow="15" pageId="6" pageNumber="5">6.8/2.8/2.8</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FD18FB2411AA" box="[401,1170,715,738]" gridrow="16" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FD18FD2E11AA" box="[401,664,715,738]" gridcol="0" gridrow="16" pageId="6" pageNumber="5">Telson, L</th>
<td id="4C01960EFFE460DCA3BFFD18FC3011AA" box="[759,902,715,738]" gridcol="1" gridrow="16" pageId="6" pageNumber="5">6.7</td>
<td id="4C01960EFFE460DCA54AFD18FB2411AA" box="[1026,1170,715,738]" gridcol="2" gridrow="16" pageId="6" pageNumber="5">5.4</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FD38FB24104A" box="[401,1170,747,770]" gridrow="17" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FD38FD2E104A" box="[401,664,747,770]" gridcol="0" gridrow="17" pageId="6" pageNumber="5">vesicle, L/W/D</th>
<td id="4C01960EFFE460DCA3BFFD38FC30104A" box="[759,902,747,770]" gridcol="1" gridrow="17" pageId="6" pageNumber="5">4.4/2.5/2.3</td>
<td id="4C01960EFFE460DCA54AFD38FB24104A" box="[1026,1170,747,770]" gridcol="2" gridrow="17" pageId="6" pageNumber="5">3.3/2.1/2.0</td>
</tr>
<tr id="0FD0FF72FFE460DCA0D9FCDFFB24106B" box="[401,1170,780,803]" gridrow="18" pageId="6" pageNumber="5">
<th id="4C01960EFFE460DCA0D9FCDFFD2E106B" box="[401,664,780,803]" gridcol="0" gridrow="18" pageId="6" pageNumber="5">Total L</th>
<td id="4C01960EFFE460DCA3BFFCDFFC30106B" box="[759,902,780,803]" gridcol="1" gridrow="18" pageId="6" pageNumber="5">68.5</td>
<td id="4C01960EFFE460DCA54AFCDFFB24106B" box="[1026,1170,780,803]" gridcol="2" gridrow="18" pageId="6" pageNumber="5">56.6</td>
</tr>
</table>
</paragraph>
<caption id="E59FADB8FFE49F25A188FC91FBA41011" box="[192,1042,834,857]" pageId="6" pageNumber="5" startId="6.[192,257,834,857]">
<paragraph id="B15FFD30FFE49F25A188FC91FBA41011" blockId="6.[192,1042,834,857]" box="[192,1042,834,857]" pageId="6" pageNumber="5">
<emphasis id="83942122FFE49F25A188FC91FEA81011" bold="true" box="[192,286,834,857]" pageId="6" pageNumber="5">Table 1:</emphasis>
Measurements (mm) of
<taxonomicName id="76E086B3FFE49F25A343FC97FD191011" authority="Trujillo &amp; de Armas, 2016" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[523,687,836,857]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="ixil" status="sp. nov.">
<emphasis id="83942122FFE49F25A343FC97FD191011" box="[523,687,836,857]" italics="true" pageId="6" pageNumber="5">Centruroides ixil</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98A79C59FFE49F25A3FDFC97FD5F1011" box="[693,745,836,857]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
D, depth; L, length; W, width.
</paragraph>
</caption>
<paragraph id="B15FFD30FFE49F25A1B8FC2CFDF51628" blockId="6.[192,802,901,1917]" pageId="6" pageNumber="5">
<emphasis id="83942122FFE49F25A1B8FC2CFDE5175E" box="[240,595,1023,1046]" italics="true" pageId="6" pageNumber="5">
Description of the male
<typeStatus id="6E5B4392FFE49F25A0BFFC2CFDE5175E" box="[503,595,1023,1046]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="29DBE1B5FFE49F25A32BFC2CFD69175E" box="[611,735,1023,1046]" captionStart="Figure 1" captionStartId="3.[192,268,1350,1373]" captionTargetBox="[198,1440,189,1320]" captionTargetId="figure-6@3.[192,1440,188,1320]" captionTargetPageId="3" captionText="Figure 1: A, geographical distribution of Centruroides ixil sp. n. (red star) and C. caral (red circle). BC, male holotype of Centruroides ixil sp. n. in dorsal (B) and ventral (C) aspect." pageId="6" pageNumber="5">Fig. 1 BC</figureCitation>
,
<figureCitation id="29DBE1B5FFE49F25A3A4FC2CFF64177C" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">2 A F</figureCitation>
). Base color light brown. Carapace with posterior margins infuscate and distinct dusky marbling throughout (
<figureCitation id="29DBE1B5FFE49F25A1BAFB8AFEE71738" box="[242,337,1113,1136]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 A</figureCitation>
); interocular triangle darker. Tergites IVI with dusky band along posterior margins (
<figureCitation id="29DBE1B5FFE49F25A3E4FBA4FCA217C6" box="[684,788,1143,1166]" captionStart="Figure 1" captionStartId="3.[192,268,1350,1373]" captionTargetBox="[198,1440,189,1320]" captionTargetId="figure-6@3.[192,1440,188,1320]" captionTargetPageId="3" captionText="Figure 1: A, geographical distribution of Centruroides ixil sp. n. (red star) and C. caral (red circle). BC, male holotype of Centruroides ixil sp. n. in dorsal (B) and ventral (C) aspect." pageId="6" pageNumber="5">Fig. 1 B</figureCitation>
). Metasomal segments IIV light brown; V and telson slightly darker. Cheliceral manus with dark brown reticulations all over (
<figureCitation id="29DBE1B5FFE49F25A0C3FB02FE5F17A0" box="[395,489,1233,1256]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 B</figureCitation>
). Pedipalp femur and patella brownish, with small yellowish areas around the trichobothria; manus light yellowish, with blackish fingers. Legs yellowish with pale brown marks. Sternites IIIVI brown yellowish; VII darker.
</paragraph>
<paragraph id="B15FFD30FFE49F25A1B8FAB4FD211518" blockId="6.[192,802,901,1917]" pageId="6" pageNumber="5">
Carapace (
<figureCitation id="29DBE1B5FFE49F25A02FFAB4FE7D1636" box="[359,459,1383,1406]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 A</figureCitation>
) 1.03 times longer than wide; anteromedian notch V-shaped, reaching the posterior margin level of the second pair of lateral eyes; interocular triangle with moderate granules, the rest finely and sparsely granular. Anterior median carinae vestigial, with disperse moderate granules; superciliary carinae strong, subgranulose; posterior median carinae moderate, subgranulose, other carinae indistinct.
</paragraph>
<paragraph id="B15FFD30FFE49F25A1B8F984FBC11725" blockId="6.[192,802,901,1917]" lastBlockId="6.[830,1441,900,1915]" pageId="6" pageNumber="5">
Pedipalps orthobothriotaxic
<typeStatus id="6E5B4392FFE49F25A36CF984FDED1526" box="[548,603,1623,1646]" pageId="6" pageNumber="5">Type</typeStatus>
A. Femur (
<figureCitation id="29DBE1B5FFE49F25A3A8F984FF6315C4" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 B</figureCitation>
): internal surface with large granules, all other surfaces finely granular; dorsointernal, dorsoexternal and ventrointernal carinae moderate, granular; ventroexternal carina strong, serrate. Patella (
<figureCitation id="29DBE1B5FFE49F25A329F91CFD0A15AE" box="[609,700,1743,1766]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 B</figureCitation>
): internal surface with 56 large conical granules; dorsal, external and ventral surfaces finely granular; dorsointernal and dorsoexternal carinae moderate, granular; dorsomedian carina weak, granular; ventrointernal carina weak, granular. Manus (
<figureCitation id="29DBE1B5FFE49F25A011F8B5FE531435" box="[345,485,1894,1917]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 B, D</figureCitation>
): globose, 1.32 times wider than patella; intercarinal surfaces almost smooth, dorsomarginal, dorsal secondary and digital carinae weak to moderate, minutely granular to subgranular; ventroexternal carina strong, smooth; other carinae vestigial. Fixed fingers with eight oblique rows of denticles; trichobothrium
<emphasis id="83942122FFE49F25A2A5FBC9FBB61779" box="[1005,1024,1050,1073]" italics="true" pageId="6" pageNumber="5">et</emphasis>
basal to
<emphasis id="83942122FFE49F25A531FBC9FB221779" box="[1145,1172,1050,1073]" italics="true" pageId="6" pageNumber="5">db</emphasis>
. Movable fingers with eight principal oblique rows of denticles, plus a distal short one with four denticles.
</paragraph>
<paragraph id="B15FFD30FFE49F25A226FBA6FAF117E3" blockId="6.[830,1441,900,1915]" pageId="6" pageNumber="5">
Pectines (
<figureCitation id="29DBE1B5FFE49F25A295FBA6FBF417C4" box="[989,1090,1141,1164]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 E</figureCitation>
) with 20/20 teeth; basal plate rectangular in shape, 2.14 times wider than long.
</paragraph>
<paragraph id="B15FFD30FFE49F25A226FB60FC2916EE" blockId="6.[830,1441,900,1915]" pageId="6" pageNumber="5">
Mesosoma. Tergites IVII finely granular, with some larger granules on the posterior one-half of IVI (
<figureCitation id="29DBE1B5FFE49F25A20EFB21FC1F1641" box="[838,937,1266,1289]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 C</figureCitation>
); longitudinal median carina IVII moderate, subgranular. Tergite VII lateral and submedian carinae moderate, granulose. Sternites IIIVI mostly smooth, except on lateral submargins; VII mostly smooth, with lateral and submedian carinae moderate to feeble, subgranular.
</paragraph>
<paragraph id="B15FFD30FFE49F25A226FA7DFA371433" blockId="6.[830,1441,900,1915]" pageId="6" pageNumber="5">
Metasoma (
<figureCitation id="29DBE1B5FFE49F25A2A5FA7DFBF2168D" box="[1005,1092,1454,1477]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 F</figureCitation>
). Intercarinal spaces IIV coriaceous, with sparse minute granules. Segments IIIV with two pairs of ventrolateral macrochaetae. Segment I, ten carinae; IIIV, eight carinae; V with five carinae; on II, the lateral inframedian carina is only represented by 68 distal granules, being the distal ones the largest; IIV with dorsolateral, lateral supramedian, ventrolateral and ventrosubmedian carinae moderately developed, granular to slightly serrate; dorsolateral carinae with a larger terminal granule. Segment V: intercarinal spaces finely granulate; dorsolateral, ventrolateral and ventromedian carinae moderate, granular. Vesicle with ventral and lateral surfaces finely granular, 1.08 times as wide as deep; subaculear tubercle strong, spine-like, with tip pointing towards the apex of the aculeus (
<figureCitation id="29DBE1B5FFE49F25A5B4F8B7FAC31433" box="[1276,1397,1892,1915]" captionStart="Figure 2" captionStartId="4.[192,268,1857,1880]" captionTargetBox="[251,1393,191,1828]" captionTargetId="figure-8@4.[252,1393,192,1827]" captionTargetPageId="4" captionText="Figure 2: Centruroides ixil sp. n. Male holotype. A, carapace; B, left pedipalp, dorsal aspect; C, tergites IVI; D, pedipalp chela, dorsal lateral aspect; E, pectines; F, metasomal segments IIIV + telson, lateral aspect; G, telson, lateral aspect." pageId="6" pageNumber="5">Fig. 2 F, G</figureCitation>
).
</paragraph>
<caption id="E59FADB8FFE59F24A188FB2FFC2F165B" box="[192,921,1276,1299]" pageId="7" pageNumber="6" startId="7.[192,268,1276,1299]" targetBox="[204,1429,167,1244]" targetPageId="7">
<paragraph id="B15FFD30FFE59F24A188FB2FFC2F165B" blockId="7.[192,921,1276,1299]" box="[192,921,1276,1299]" pageId="7" pageNumber="6">
<emphasis id="83942122FFE59F24A188FB2FFE9F165B" bold="true" box="[192,297,1276,1299]" pageId="7" pageNumber="6">Figure 4:</emphasis>
Habitat of
<taxonomicName id="76E086B3FFE59F24A0DDFB2DFD8F165B" authority="Trujillo &amp; de Armas, 2016" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[405,569,1278,1299]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ixil" status="sp. nov.">
<emphasis id="83942122FFE59F24A0DDFB2DFD8F165B" box="[405,569,1278,1299]" italics="true" pageId="7" pageNumber="6">Centruroides ixil</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98A79C59FFE59F24A377FB2DFDC2165B" box="[575,628,1278,1299]" pageId="7" pageNumber="6" rank="species">
<emphasis id="83942122FFE59F24A377FB2DFDD8165B" bold="true" box="[575,622,1278,1299]" pageId="7" pageNumber="6">sp. n</emphasis>
.
</taxonomicNameLabel>
(photos by Andrea Martinez).
</paragraph>
</caption>
<paragraph id="B15FFD30FFE59F24A1B8FA9BFE3B16D5" blockId="7.[192,802,1352,1743]" pageId="7" pageNumber="6">Legs with femora finely granular on the prolateral surface. Telotarsi moderately covered by fine setae on the ventral surface.</paragraph>
<paragraph id="B15FFD30FFE59F24A1B8FA77FE161587" blockId="7.[192,802,1352,1743]" pageId="7" pageNumber="6">
<emphasis id="83942122FFE59F24A1B8FA77FEF616F3" box="[240,320,1444,1467]" italics="true" pageId="7" pageNumber="6">Female</emphasis>
(
<figureCitation id="29DBE1B5FFE59F24A01CFA77FE6B16F3" box="[340,477,1444,1467]" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="7" pageNumber="6">Fig. 3 AG</figureCitation>
). The only examined female differs from the male
<typeStatus id="6E5B4392FFE59F24A087FA10FD9A1692" box="[463,556,1475,1498]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
by having a darker pattern, mainly on carapace and tergites (
<figureCitation id="29DBE1B5FFE59F24A3DAFA31FCA016B1" box="[658,790,1506,1529]" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="7" pageNumber="6">Fig. 3 AB</figureCitation>
), small size (
<tableCitation id="FC62C88BFFE59F24A00BF9D3FE2C155F" box="[323,410,1536,1559]" captionStart="Table 1" captionStartId="6.[192,257,834,857]" captionText="Table 1: Measurements (mm) of Centruroides ixil sp. n. D, depth; L, length; W, width." pageId="7" pageNumber="6">Table 1</tableCitation>
), carapace (
<figureCitation id="29DBE1B5FFE59F24A360F9D3FD3A155F" box="[552,652,1536,1559]" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="7" pageNumber="6">Fig. 3 C</figureCitation>
) and tergites most strongly granulate, pedipalp chelae with slender manus (
<figureCitation id="29DBE1B5FFE59F24A050F9EDFE29151D" box="[280,415,1598,1621]" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="7" pageNumber="6">Fig. 3 A, D</figureCitation>
), pectines with 19/19 teeth (
<figureCitation id="29DBE1B5FFE59F24A397F9EDFF65153B" captionStart="Figure 3" captionStartId="5.[192,268,1846,1869]" captionTargetBox="[191,1440,191,1816]" captionTargetId="figure-6@5.[192,1439,192,1815]" captionTargetPageId="5" captionText="Figure 3: Centruroides ixil sp. n. AG, female paratype: AB, dorsal (A) and ventral (B) aspect. C, metasomal segments III V + telson, lateral aspect. D, left pedipalp, dorsal aspect. E, pectines. F, metasomal segments IIIV + telson, lateral aspect; G, sternites IVVI." pageId="7" pageNumber="6">Fig. 3 E</figureCitation>
), metasoma not attenuate (segment III length/width ratio = 1.86 vs 2.76; segment V length/width ratio = 0.88 vs 0.68; carapace length/metasomal segment V length ratio = 2.43 vs 2.78).
</paragraph>
<paragraph id="B15FFD30FFE59F24A188F924FEC3145B" blockId="7.[192,373,1783,1811]" box="[192,373,1783,1811]" pageId="7" pageNumber="6">
<emphasis id="83942122FFE59F24A188F924FEC3145B" bold="true" box="[192,373,1783,1811]" pageId="7" pageNumber="6">Comparisons</emphasis>
</paragraph>
<paragraph id="B15FFD30FFE59F24A1B8F8E0FBED151D" blockId="7.[192,802,1843,1897]" lastBlockId="7.[830,1440,1352,1897]" pageId="7" pageNumber="6">
The male
<typeStatus id="6E5B4392FFE59F24A02EF8E0FE721402" box="[358,452,1843,1866]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
of this new species resembles that of
<taxonomicName id="76E086B3FFE59F24A05AF881FDCB1421" authority="Armas et Trujillo, 2013" authorityName="Armas et Trujillo" authorityYear="2013" box="[274,637,1874,1897]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFE59F24A05AF881FEDA1421" box="[274,364,1874,1897]" italics="true" pageId="7" pageNumber="6">C. caral</emphasis>
Armas et Trujillo, 2013
</taxonomicName>
, from which it differs by having a most globose pedipalp manus, telson wider than its height, and more attenuate metasoma (length/width ratio: segments IIIV = 2.4, 2.7 and 1.4, respectively; 1.7, 2.2 and
<quantity id="761850D5FFE59F24A510FA77FB2016F3" box="[1112,1174,1444,1467]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" pageId="7" pageNumber="6" unit="in" value="2.5">2.5 in</quantity>
the male
<typeStatus id="6E5B4392FFE59F24A44BFA77FAD616F3" box="[1283,1376,1444,1467]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
of
<taxonomicName id="76E086B3FFE59F24A4C0FA77FCC01692" authorityName="Armas et Trujillo" authorityYear="2013" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFE59F24A4C0FA77FCC01692" italics="true" pageId="7" pageNumber="6">C. caral</emphasis>
</taxonomicName>
, only known specimen of this taxon). Also, general pattern differs in both species: base color light brownish in
<taxonomicName id="76E086B3FFE59F24A229F9D3FC1C155F" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[865,938,1536,1559]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ixil">
<emphasis id="83942122FFE59F24A229F9D3FC1C155F" box="[865,938,1536,1559]" italics="true" pageId="7" pageNumber="6">C. ixil</emphasis>
</taxonomicName>
, but light yellow in
<taxonomicName id="76E086B3FFE59F24A5E0F9D3FAB1155F" authorityName="Armas et Trujillo" authorityYear="2013" box="[1192,1287,1536,1559]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFE59F24A5E0F9D3FAB1155F" box="[1192,1287,1536,1559]" italics="true" pageId="7" pageNumber="6">C. caral</emphasis>
</taxonomicName>
; each tergite bearing a narrow yellow median longitudinal line in
<taxonomicName id="76E086B3FFE59F24A4CFF9CCFCC0151D" authorityName="Armas et Trujillo" authorityYear="2013" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFE59F24A4CFF9CCFCC0151D" italics="true" pageId="7" pageNumber="6">C. caral</emphasis>
</taxonomicName>
, but absent in
<taxonomicName id="76E086B3FFE59F24A55BF9EDFBE2151D" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[1043,1108,1598,1621]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ixil">
<emphasis id="83942122FFE59F24A55BF9EDFBE2151D" box="[1043,1108,1598,1621]" italics="true" pageId="7" pageNumber="6">C. ixil</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B15FFD30FFE59F24A226F98FFCC81421" blockId="7.[830,1440,1352,1897]" pageId="7" pageNumber="6">
Although male scorpions of the same species that reach the adulthood from different stadia show significant morphometric differences, those observed between the respective males of
<taxonomicName id="76E086B3FFE59F24A5F2F96BFAA11587" authorityName="Armas et Trujillo" authorityYear="2013" box="[1210,1303,1720,1743]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFE59F24A5F2F96BFAA11587" box="[1210,1303,1720,1743]" italics="true" pageId="7" pageNumber="6">C. caral</emphasis>
</taxonomicName>
and
<taxonomicName id="76E086B3FFE59F24A411F96BFA161587" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[1369,1440,1720,1743]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ixil">
<emphasis id="83942122FFE59F24A411F96BFA161587" box="[1369,1440,1720,1743]" italics="true" pageId="7" pageNumber="6">C. ixil</emphasis>
</taxonomicName>
seem to be not caused by that phenomenon. Unfortunately, the female of the former is unknown, but the differences between the males of these two species strongly suggest that
<taxonomicName id="76E086B3FFE59F24A563F8E0FBD91402" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[1067,1135,1843,1866]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ixil">
<emphasis id="83942122FFE59F24A563F8E0FBD91402" box="[1067,1135,1843,1866]" italics="true" pageId="7" pageNumber="6">C. ixil</emphasis>
</taxonomicName>
is not the large male of
<taxonomicName id="76E086B3FFE59F24A4CFF8E0FCC01421" authorityName="Armas et Trujillo" authorityYear="2013" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFE59F24A4CFF8E0FCC01421" italics="true" pageId="7" pageNumber="6">C. caral</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B15FFD30FFEA9F2BA188FF1AFE2C13AD" blockId="8.[192,410,201,229]" box="[192,410,201,229]" pageId="8" pageNumber="7">
<emphasis id="83942122FFEA9F2BA188FF1AFE2C13AD" bold="true" box="[192,410,201,229]" pageId="8" pageNumber="7">Natural History</emphasis>
</paragraph>
<paragraph id="B15FFD30FFEA9F2BA1B8FED9FE1F115F" blockId="8.[192,802,266,872]" pageId="8" pageNumber="7">
The
<typeStatus id="6E5B4392FFEA9F2BA066FED9FE3D1269" box="[302,395,266,289]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
male and
<typeStatus id="6E5B4392FFEA9F2BA36AFED9FDC81269" box="[546,638,266,289]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
female were collected on a rocky wall partly covered by vegetation in a mountainous subtropical wet forest,
<quantity id="761850D5FFEA9F2BA338FE9BFD701217" box="[624,710,328,351]" metricMagnitude="3" metricUnit="m" metricValue="1.173" pageId="8" pageNumber="7" unit="m" value="1173.0">1173 m</quantity>
a.s.l., a few kilometers west from Visís Cabá Biosphere Reserve. The male and female were separate
<quantity id="761850D5FFEA9F2BA3F5FE56FCB412D4" box="[701,770,389,412]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="7" unit="cm" value="30.0">30 cm</quantity>
to each other, about
<quantity id="761850D5FFEA9F2BA036FE77FE1012F3" box="[382,422,420,443]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="7" unit="m" value="3.0">3 m</quantity>
over the soil, in a place covered by leaf litter. The type locality is in the margin of a nonpavement road, approximately at
<quantity id="761850D5FFEA9F2BA373FE32FD3312B0" box="[571,645,481,504]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="7" unit="m" value="300.0">300 m</quantity>
from a small river cascade (
<figureCitation id="29DBE1B5FFEA9F2BA012FDD3FE2A115F" box="[346,412,512,535]" captionStart="Figure 4" captionStartId="7.[192,268,1276,1299]" captionTargetBox="[204,1429,167,1244]" captionTargetId="figure-6@7.[204,1428,168,1243]" captionTargetPageId="7" captionText="Figure 4: Habitat of Centruroides ixil sp. n. (photos by Andrea Martinez)." pageId="8" pageNumber="7">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="B15FFD30FFEA9F2BA1B8FDCDFE971020" blockId="8.[192,802,266,872]" pageId="8" pageNumber="7">
In
<collectingCountry id="C9F7BDA0FFEA9F2BA050FDCDFE38117D" box="[280,398,542,565]" name="Guatemala" pageId="8" pageNumber="7">Guatemala</collectingCountry>
, the mountainous subtropical wet forest is located in parts of Los Cuchumatanes, Chamá, Chuacús and Las Minas mountain systems. This forest is characterized by a complex vegetation structure over an irregular landscape, with high levels of humidity and precipitation and a great diversity of habitats, supporting high levels of endemism (
<bibRefCitation id="D57180C1FFEA9F2BA0AFFD05FD1411A5" author="CASTANEDA" box="[487,674,726,749]" pageId="8" pageNumber="7" refId="ref3223" refString="CASTANEDA, C. 2008. Diversidad de Ecosistemas de Guatemala. In: CONAP (eds.), Guatemala y su biodiversidad: Un enfoque historico, cultural, biologico y economico. Consejo Nacional de Areas Protegidas, Oficina Tecnica de Biodiversidad. Guatemala, 630 pp." type="book" year="2008">Castañeda, 2008</bibRefCitation>
). This ecosystem is sometimes known as “mix forest” due to the association of conifers and broad-leaf species. Epiphytes, mosses and ferns are also abundant in this
<typeStatus id="6E5B4392FFEA9F2BA3BCFCE1FC941001" box="[756,802,818,841]" pageId="8" pageNumber="7">type</typeStatus>
of forest.
</paragraph>
<paragraph id="B15FFD30FFEA9F2BA188FC43FD9710E4" blockId="8.[192,545,912,940]" box="[192,545,912,940]" pageId="8" pageNumber="7">
<emphasis id="83942122FFEA9F2BA188FC43FD9710E4" bold="true" box="[192,545,912,940]" pageId="8" pageNumber="7">Biogeographic Comments</emphasis>
</paragraph>
<paragraph id="B15FFD30FFEA9F2BA1B8FC01FDCC169C" blockId="8.[192,803,978,1675]" pageId="8" pageNumber="7">
The type locality of
<taxonomicName id="76E086B3FFEA9F2BA099FC01FDA310A1" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[465,533,978,1001]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="ixil">
<emphasis id="83942122FFEA9F2BA099FC01FDA310A1" box="[465,533,978,1001]" italics="true" pageId="8" pageNumber="7">C. ixil</emphasis>
</taxonomicName>
belongs to Sierra de los Cuchumatanes, a continuous of the
<collectingRegion id="732433D2FFEA9F2BA338FC22FD701740" box="[624,710,1009,1032]" country="Mexico" name="Chiapas" pageId="8" pageNumber="7">Chiapas</collectingRegion>
Range. According with several authors (for details see
<bibRefCitation id="D57180C1FFEA9F2BA3F5FBDCFF4F170D" author="MORRONE" pageId="8" pageNumber="7" pagination="1 - 148" refId="ref3357" refString="MORRONE, J. J. 2001. Biogeografia de America Latina y el Caribe. Manuales &amp; Tesis SEA, 3: 1 - 148." type="journal article" year="2001">Morrone, 2001</bibRefCitation>
), this orographic area belongs to the
<collectingRegion id="732433D2FFEA9F2BA384FBFDFE96172C" country="Mexico" name="Chiapas" pageId="8" pageNumber="7">Chiapas province</collectingRegion>
, a biogeographic unity that shows a strong relationship with the neighboring province of Eastern Central America. To the later belongs the
<typeStatus id="6E5B4392FFEA9F2BA3D2FB59FD7E17E9" box="[666,712,1162,1185]" pageId="8" pageNumber="7">type</typeStatus>
locality of
<taxonomicName id="76E086B3FFEA9F2BA1AAFB7BFE8917F7" authorityName="Armas et Trujillo" authorityYear="2013" box="[226,319,1192,1215]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFEA9F2BA1AAFB7BFE8917F7" box="[226,319,1192,1215]" italics="true" pageId="8" pageNumber="7">C. caral</emphasis>
</taxonomicName>
, a species geographically and ecologically disjointed from
<taxonomicName id="76E086B3FFEA9F2BA03CFB14FE0C1796" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[372,442,1223,1246]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="ixil">
<emphasis id="83942122FFEA9F2BA03CFB14FE0C1796" box="[372,442,1223,1246]" italics="true" pageId="8" pageNumber="7">C. ixil</emphasis>
</taxonomicName>
by the valleys of Motagua and Chixoy, two areas with xeromorphic vegetation. A similar distribution is shown by the beetles
<emphasis id="83942122FFEA9F2BA3DBFAD7FEBC1672" italics="true" pageId="8" pageNumber="7">Crysina purulensis</emphasis>
(Monzón et Warner, 1993),
<taxonomicName id="76E086B3FFEA9F2BA319FAF0FEB71611" authority="(Ohaus, 1912)" baseAuthorityName="Ohaus" baseAuthorityYear="1912" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="83942122FFEA9F2BA319FAF0FD721672" box="[593,708,1315,1338]" italics="true" pageId="8" pageNumber="7">C. diversa</emphasis>
(Ohaus, 1912)
</taxonomicName>
,
<taxonomicName id="76E086B3FFEA9F2BA05DFA91FD881611" authority="(Moran, 1990)" baseAuthorityName="Moran" baseAuthorityYear="1990" box="[277,574,1346,1369]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="quiche">
<emphasis id="83942122FFEA9F2BA05DFA91FE331611" box="[277,389,1346,1369]" italics="true" pageId="8" pageNumber="7">C. quiche</emphasis>
(Moran, 1990)
</taxonomicName>
, and
<taxonomicName id="76E086B3FFEA9F2BA3C2FA91FECE163F" authority="(Boucard, 1878)" baseAuthorityName="Boucard" baseAuthorityYear="1878" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="rodriguezi">
<emphasis id="83942122FFEA9F2BA3C2FA91FC941611" box="[650,802,1346,1369]" italics="true" pageId="8" pageNumber="7">C. rodriguezi</emphasis>
(Boucard, 1878)
</taxonomicName>
, all them restricted to Cuchumatanes Range, whereas
<taxonomicName id="76E086B3FFEA9F2BA0CCFAACFD7716DE" authority="(Ohaus, 1924)" baseAuthorityName="Ohaus" baseAuthorityYear="1924" box="[388,705,1407,1430]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="strasseni">
<emphasis id="83942122FFEA9F2BA0CCFAACFDBB16DE" box="[388,525,1407,1430]" italics="true" pageId="8" pageNumber="7">C. strasseni</emphasis>
(Ohaus, 1924)
</taxonomicName>
and
<taxonomicName id="76E086B3FFEA9F2BA241FAACFDF916FD" authority="(Monzon et Cano, 1999)" baseAuthorityName="Monzon et Cano" baseAuthorityYear="1999" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="ericsmithi">
<emphasis id="83942122FFEA9F2BA241FAACFE9D16FD" italics="true" pageId="8" pageNumber="7">C. ericsmithi</emphasis>
(Monzón et Cano, 1999)
</taxonomicName>
are endemic from Merendón Range (
<bibRefCitation id="D57180C1FFEA9F2BA0CEFA6EFDD9169C" author="MONZON SIERRA" box="[390,623,1469,1492]" pageId="8" pageNumber="7" pagination="393 - 401" refId="ref3303" refString="MONZON SIERRA, J. 2006. El genero Chrysina Kirby (Coleoptera: Scarabaeidae) en Guatemala. Pp. 393 - 401 in: Cano, E. B. (ed.), Bioiversidad de Guatemala. Universidad del Valle de Guatemala, Guatemala, Guatemala. 674 pp." type="book chapter" year="2006">Monzón Sierra, 2006</bibRefCitation>
).
</paragraph>
<paragraph id="B15FFD30FFEA9F2BA1B8FA08FEAA15C3" blockId="8.[192,803,978,1675]" pageId="8" pageNumber="7">
On this respect,
<taxonomicName id="76E086B3FFEA9F2BA0E5FA08FE4216BA" authorityName="Trujillo &amp; de Armas" authorityYear="2016" box="[429,500,1499,1522]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="ixil">
<emphasis id="83942122FFEA9F2BA0E5FA08FE4216BA" box="[429,500,1499,1522]" italics="true" pageId="8" pageNumber="7">C. ixil</emphasis>
</taxonomicName>
and
<taxonomicName id="76E086B3FFEA9F2BA37AFA08FD3816BA" authorityName="Armas et Trujillo" authorityYear="2013" box="[562,654,1499,1522]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="caral">
<emphasis id="83942122FFEA9F2BA37AFA08FD3816BA" box="[562,654,1499,1522]" italics="true" pageId="8" pageNumber="7">C. caral</emphasis>
</taxonomicName>
seems to be closely related species, perhaps originated as result of a vicariance event during the Upper Pliocene, and pronounced during the last two millions of years, when the valleys of Motagua and Chixoy became xeric ecosystems.
</paragraph>
</subSubSection>
</treatment>
</document>