599 lines
72 KiB
XML
599 lines
72 KiB
XML
<document id="D2390235E16BE0BFB75C75BAC786B1A1" ID-ISSN="1536-9307" ID-ZooBank="73665FA8-D9E4-4996-8176-E22BB56086BD" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638259957435" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme & Šťáhlavský, František" docDate="2018" docId="6974879002098F56FC3EF959FD41D3CE" docLanguage="en" docName="Euscorpius.268-1-27.pdf" docOrigin="Euscorpius 268" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Chaerilus neradorum Kovařík & Lowe & Šťáhlavský 2018, sp. n." docType="treatment" docUuid="21E8792C-9989-4747-A6BD-FD75CD80F989" docUuidSource="ZooBank" docVersion="3" lastPageNumber="16" masterDocId="954DFFE802018F42FFF2FFF1FFFCD574" masterDocTitle="Three new Chaerilus from Malaysia (Tioman Island) and Thailand (Scorpiones Chaerilidae) with a review of C cimrmani C sejnai and C tichyi" masterLastPageNumber="27" masterPageNumber="1" pageNumber="7" updateTime="1699436302183" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5F6CE9F400577F792A3647FEA611A816">Three new Chaerilus from Malaysia (Tioman Island) and Thailand (Scorpiones Chaerilidae) with a review of C cimrmani C sejnai and C tichyi</mods:title>
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<mods:namePart id="82BB0E284A28A523EEDBC7B051B21E5E">Kovařík, František</mods:namePart>
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<mods:namePart id="7C380FBBF43D8AABC3C649B5B733272E">Lowe, Graeme</mods:namePart>
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<mods:namePart id="B2BA218DFFE6D66473F87E12C5B19DFA">Šťáhlavský, František</mods:namePart>
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<mods:title id="49EDABA2C248BEC213CB22E5780B7F74">Euscorpius</mods:title>
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<mods:date id="93EB9BEE49F79E648BFB2DBE4B81CB4D">2018</mods:date>
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<mods:number id="E1566AF750339ECB6E1307D9EBD61A92">268</mods:number>
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<taxonomicName id="26DD4D0502098F4AFC3EF959FB2ED3B7" ID-CoL="T9ZK" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[972,1234,1704,1731]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">Chaerilus neradorum</taxonomicName>
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.
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.
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="A9C7650D02098F4AFC3AF93AFB10D20F" pageId="8" pageNumber="7" type="description">
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<paragraph id="E162368602098F4AFC3AF93AFA63D275" blockId="8.[841,1439,1704,1823]" pageId="8" pageNumber="7">
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(
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<figureCitation id="79E62A0302098F4AFC23F93AFBADD396" box="[977,1105,1739,1762]" captionStart-0="Figures 43–44" captionStart-1="Figures 45–46" captionStart-2="Figures 47–51" captionStart-3="Figures 52–66" captionStart-4="Figures 67–68" captionStartId-0="12.[192,278,1767,1790]" captionStartId-1="13.[192,278,1653,1676]" captionStartId-2="14.[192,278,1786,1809]" captionStartId-3="15.[192,278,1734,1757]" captionStartId-4="16.[192,278,1720,1743]" captionTargetBox-0="[191,1440,199,1729]" captionTargetBox-1="[208,1429,213,1619]" captionTargetBox-2="[191,1440,191,1757]" captionTargetBox-3="[204,1435,217,1702]" captionTargetBox-4="[191,1440,191,1690]" captionTargetId-0="figure-75@12.[192,1439,200,1728]" captionTargetId-1="figure-124@13.[192,1439,192,1623]" captionTargetId-2="figure-76@14.[192,1439,192,1756]" captionTargetId-3="figure-6@15.[192,1439,212,1704]" captionTargetId-4="figure-19@16.[192,1439,192,1690]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionText-0="Figures 43–44: Chaerilus, collection localities. Figure 43. C. majkusi sp. n., type locality. Figure 44. C. neradorum sp. n., type locality." captionText-1="Figures 45–46: Chaerilus neradorum sp. n., holotype male, dorsal (45) and ventral (46) views. Scale bar: 10 mm." captionText-2="Figures 47–51: Chaerilus neradorum sp. n., holotype male, carapace and tergites I–III (47), sternopectinal region and sternites (48), metasoma and telson, lateral (49), ventral (50), and dorsal (51) views. Scale bar: 5 mm (49–51)." captionText-3="Figures 52–66: Chaerilus neradorum sp. n., holotype male, pedipalp chela, dorsal (52), externodorsal (53), and ventrointernal (54) views. Pedipalp patella, dorsal (55), external (56) and ventral (57) views. Pedipalp femur and trochanter, dorsal (58) and ventral (59) views. Movable (60) and fixed (61) fingers. Left legs I–IV, retrolateral/ventral aspects (62–66). The trichobothrial pattern is indicated in Figures 52–58." captionText-4="Figures 67–68: Chaerilus neradorum sp. n., male holotype (67) and Chaerilus cimrmani Kovařík, 2012, male (68) in vivo habitus." pageId="8" pageNumber="7">Figs. 44–67</figureCitation>
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,
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<figureCitation id="79E62A0302098F4AFBAEF93AFB45D396" box="[1116,1209,1739,1762]" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="8" pageNumber="7">117–118</figureCitation>
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,
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<tableCitation id="AC5F033D02098F4AFB34F93AFAEBD396" box="[1222,1303,1739,1762]" captionStart="Table 1" captionStartId="17.[192,257,818,841]" captionText="Table 1: Comparative measurements of Chaerilus majkusi sp. n. and Chaerilus neradorum sp. n. types. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="8" pageNumber="7">Table 1</tableCitation>
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)
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<emphasis id="D3A9EA9402098F4AFCBBF91BFA63D275" bold="true" box="[841,1439,1770,1793]" pageId="8" pageNumber="7">http://zoobank.org/urn:lsid:zoobank.org:act:21E879</emphasis>
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<materialsCitation id="51B53CDB02098F4AFCBAF8B4FB6BD20F" collectionCode="T, YPE, LOCALITY, AND, REPOSITORY" country="Thailand" latitude="9.46555" location="Samui" longLatPrecision="1" longitude="99.98419" pageId="8" pageNumber="7" specimenCount="1" typeStatus="holotype">
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<collectionCode id="87CCAE4302098F4AFCAAF8B8FBBCD228" box="[856,1088,1865,1884]" pageId="8" pageNumber="7">YPE LOCALITY AND</collectionCode>
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.
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<collectingCountry id="99CA761602098F4AFAC9F8B4FA66D228" box="[1339,1434,1861,1884]" name="Thailand" pageId="8" pageNumber="7">Thailand</collectingCountry>
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,
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,
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; FKCP.
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</paragraph>
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</subSubSection>
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<caption id="B5A2660E02088F4BFF32F8D7FA6CD249" box="[192,1424,1830,1853]" pageId="9" pageNumber="8" startId="9.[192,278,1830,1853]" targetBox="[191,1440,212,1801]" targetPageId="9">
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<paragraph id="E162368602088F4BFF32F8D7FA6CD249" blockId="9.[192,1424,1830,1853]" box="[192,1424,1830,1853]" pageId="9" pageNumber="8">
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<emphasis id="D3A9EA9402088F4BFF32F8D7FE94D249" bold="true" box="[192,360,1830,1853]" pageId="9" pageNumber="8">Figures 37–38:</emphasis>
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<taxonomicName id="26DD4D0502088F4BFE9CF8D9FDE2D249" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[366,542,1832,1853]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
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<emphasis id="D3A9EA9402088F4BFE9CF8D9FDE2D249" box="[366,542,1832,1853]" italics="true" pageId="9" pageNumber="8">Chaerilus majkusi</emphasis>
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</taxonomicName>
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<emphasis id="D3A9EA9402088F4BFDD6F8D9FDC7D249" bold="true" box="[548,571,1832,1853]" pageId="9" pageNumber="8">sp</emphasis>
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.
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<emphasis id="D3A9EA9402088F4BFDB4F8D9FDAFD249" bold="true" box="[582,595,1832,1853]" pageId="9" pageNumber="8">n</emphasis>
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.
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</taxonomicNameLabel>
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, female paratype (37) and
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<taxonomicName id="26DD4D0502088F4BFCAEF8D9FB71D249" authority="Kovarik, 2005" authorityName="Kovarik" authorityYear="2005" box="[860,1165,1832,1853]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="sejnai">
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<emphasis id="D3A9EA9402088F4BFCAEF8D9FC06D249" box="[860,1018,1832,1853]" italics="true" pageId="9" pageNumber="8">Chaerilus sejnai</emphasis>
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Kovařík, 2005
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</taxonomicName>
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, male (38) in vivo habitus.
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</paragraph>
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</caption>
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<emphasis id="D3A9EA94020B8F48FF32F951FE94D3C3" bold="true" box="[192,360,1696,1719]" pageId="10" pageNumber="9">Figures 39–40:</emphasis>
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<taxonomicName id="26DD4D05020B8F48FE9CF950FDE2D3C2" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[366,542,1697,1718]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
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<emphasis id="D3A9EA94020B8F48FE9CF950FDE2D3C2" box="[366,542,1697,1718]" italics="true" pageId="10" pageNumber="9">Chaerilus majkusi</emphasis>
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</taxonomicName>
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<emphasis id="D3A9EA94020B8F48FDD6F953FDC7D3C3" bold="true" box="[548,571,1698,1719]" pageId="10" pageNumber="9">sp</emphasis>
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.
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<emphasis id="D3A9EA94020B8F48FDB4F953FDAFD3C3" bold="true" box="[582,595,1698,1719]" pageId="10" pageNumber="9">n</emphasis>
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.
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</taxonomicNameLabel>
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, male holotype (39) and female paratype with newborns (40) in vivo habitus.
|
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</paragraph>
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</caption>
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<materialsCitation id="51B53CDB020B8F48FF32F913FD6DD26C" collectionCode="MATERIAL" collectorName="L. Nerad" country="Thailand" latitude="9.46555" location="Samui" longLatPrecision="1" longitude="99.98419" pageId="10" pageNumber="9" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<collectingCountry id="99CA7616020B8F48FE67F913FE01D38D" box="[405,509,1762,1785]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
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</emphasis>
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,
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,
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(
|
||
<figureCitation id="79E62A03020B8F48FEDFF8F0FE85D26C" box="[301,377,1793,1816]" captionStart="Figures 43–44" captionStartId="12.[192,278,1767,1790]" captionTargetBox="[191,1440,199,1729]" captionTargetId="figure-75@12.[192,1439,200,1728]" captionTargetPageId="12" captionText="Figures 43–44: Chaerilus, collection localities. Figure 43. C. majkusi sp. n., type locality. Figure 44. C. neradorum sp. n., type locality." pageId="10" pageNumber="9">Fig. 44</figureCitation>
|
||
),
|
||
<specimenCount id="F7DBFD0F020B8F48FE7AF8F0FE56D263" box="[392,426,1793,1816]" pageId="10" pageNumber="9" type="male">1♂</specimenCount>
|
||
(1216), leg.
|
||
<collectorName id="4C285350020B8F48FDC1F8F0FD6DD26C" box="[563,657,1793,1816]" pageId="10" pageNumber="9">L. Nerad</collectorName>
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="87CCAE43020B8F48FD6DF8F0FD1ED26C" box="[671,738,1793,1816]" pageId="10" pageNumber="9">FKCP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9C7650D020B8F48FF32F8CFFBC5D26C" pageId="10" pageNumber="9" type="etymology">
|
||
<paragraph id="E1623686020B8F48FF32F8CFFBC5D26C" blockId="10.[192,802,1854,1908]" lastBlockId="10.[830,1440,1762,1816]" pageId="10" pageNumber="9">
|
||
ETYMOLOGY. The specific epithet honors Ladislav Nerad and his wife Hana (
|
||
<collectingCountry id="99CA7616020B8F48FE09F8ACFD53D200" box="[507,687,1885,1908]" name="Czech Republic" pageId="10" pageNumber="9">Czech Republic</collectingCountry>
|
||
) for their invaluable contributions to the knowledge of the scorpion fauna of
|
||
<collectingCountry id="99CA7616020B8F48FC21F8F0FBCED26C" box="[979,1074,1793,1816]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9C7650D020B8F4FFCCCF8CFFD4DD393" lastPageId="13" lastPageNumber="12" pageId="10" pageNumber="9" type="diagnosis">
|
||
<paragraph id="E1623686020B8F48FCCCF8CFFA5DD200" blockId="10.[830,1441,1854,1908]" pageId="10" pageNumber="9">
|
||
DIAGNOSIS. Total length of male
|
||
<typeStatus id="3E668824020B8F48FB4CF8CFFAE0D221" box="[1214,1308,1854,1877]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
<quantity id="26259B63020B8F48FADAF8CFFA65D221" box="[1320,1433,1854,1877]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" pageId="10" pageNumber="9" unit="mm" value="23.25">23.25 mm</quantity>
|
||
. Two pairs of lateral eyes and one pair of median eyes.
|
||
</paragraph>
|
||
<caption id="B5A2660E020A8F49FF32F915FB63D38F" box="[192,1183,1764,1787]" pageId="11" pageNumber="10" startId="11.[192,278,1764,1787]" targetBox="[191,1440,191,1734]" targetPageId="11">
|
||
<paragraph id="E1623686020A8F49FF32F915FB63D38F" blockId="11.[192,1183,1764,1787]" box="[192,1183,1764,1787]" pageId="11" pageNumber="10">
|
||
<emphasis id="D3A9EA94020A8F49FF32F915FE94D38F" bold="true" box="[192,360,1764,1787]" pageId="11" pageNumber="10">Figures 41–42:</emphasis>
|
||
<taxonomicName id="26DD4D05020A8F49FE9CF917FD65D38F" authority="Kovarik, 2000" authorityName="Kovarik" authorityYear="2000" box="[366,665,1766,1787]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="tichyi">
|
||
<emphasis id="D3A9EA94020A8F49FE9CF917FDF4D38F" box="[366,520,1766,1787]" italics="true" pageId="11" pageNumber="10">Chaerilus tichyi</emphasis>
|
||
Kovařík, 2000
|
||
</taxonomicName>
|
||
, male (41) and female (42) topotypes in vivo habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E1623686020A8F49FF32F8D1FA66D200" blockId="11.[192,794,1824,1908]" lastBlockId="11.[840,1441,1824,1908]" pageId="11" pageNumber="10">
|
||
Chela length/width ratio in male 2.4. Movable and fixed fingers of pedipalp with 11 imbricated granule rows. Fingers straight. Chela of pedipalp with 8 carinae. Pectinal teeth number
|
||
<quantity id="26259B63020A8F49FBD0F8D1FBADD243" box="[1058,1105,1824,1847]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="11" pageNumber="10" unit="in" value="6.0">6 in</quantity>
|
||
males. Carapace and tergites sparsely granulated. All sternites smooth without carinae and granules. First metasomal segment with 10
|
||
</paragraph>
|
||
<caption id="B5A2660E020D8F4EFF32F916FEC3D262" pageId="12" pageNumber="11" startId="12.[192,278,1767,1790]" targetBox="[191,1440,199,1729]" targetPageId="12">
|
||
<paragraph id="E1623686020D8F4EFF32F916FEC3D262" blockId="12.[192,1440,1767,1814]" pageId="12" pageNumber="11">
|
||
<emphasis id="D3A9EA94020D8F4EFF32F916FE96D38A" bold="true" box="[192,362,1767,1790]" pageId="12" pageNumber="11">Figures 43–44:</emphasis>
|
||
<taxonomicName id="26DD4D05020D8F4EFE81F919FE2ED389" authorityName="Simon" authorityYear="1877" box="[371,466,1768,1789]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D3A9EA94020D8F4EFE81F919FE2ED389" box="[371,466,1768,1789]" italics="true" pageId="12" pageNumber="11">Chaerilus</emphasis>
|
||
</taxonomicName>
|
||
, collection localities.
|
||
<emphasis id="D3A9EA94020D8F4EFD5EF919FCECD389" bold="true" box="[684,784,1768,1789]" pageId="12" pageNumber="11">Figure 43</emphasis>
|
||
.
|
||
<taxonomicName id="26DD4D05020D8F4EFCECF919FC7BD389" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[798,903,1768,1789]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
|
||
<emphasis id="D3A9EA94020D8F4EFCECF919FC7BD389" box="[798,903,1768,1789]" italics="true" pageId="12" pageNumber="11">C. majkusi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF020D8F4EFC7DF919FC3BD389" box="[911,967,1768,1789]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="D3A9EA94020D8F4EFC7DF919FC5AD389" bold="true" box="[911,934,1768,1789]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA94020D8F4EFC46F919FC3DD389" bold="true" box="[948,961,1768,1789]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, type locality.
|
||
<emphasis id="D3A9EA94020D8F4EFBACF919FB3ED389" bold="true" box="[1118,1218,1768,1789]" pageId="12" pageNumber="11">Figure 44</emphasis>
|
||
.
|
||
<taxonomicName id="26DD4D05020D8F4EFB22F919FAA5D389" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[1232,1369,1768,1789]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA94020D8F4EFB22F919FAA5D389" box="[1232,1369,1768,1789]" italics="true" pageId="12" pageNumber="11">C. neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF020D8F4EFA93F919FA65D389" box="[1377,1433,1768,1789]" pageId="12" pageNumber="11" rank="species">
|
||
<emphasis id="D3A9EA94020D8F4EFA93F919FA84D389" bold="true" box="[1377,1400,1768,1789]" pageId="12" pageNumber="11">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA94020D8F4EFA74F919FA6FD389" bold="true" box="[1414,1427,1768,1789]" pageId="12" pageNumber="11">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, type locality.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B5A2660E020C8F4FFF32F984FAE0D3F8" box="[192,1308,1653,1676]" pageId="13" pageNumber="12" startId="13.[192,278,1653,1676]" targetBox="[208,1429,213,1619]" targetPageId="13">
|
||
<paragraph id="E1623686020C8F4FFF32F984FAE0D3F8" blockId="13.[192,1308,1653,1676]" box="[192,1308,1653,1676]" pageId="13" pageNumber="12">
|
||
<emphasis id="D3A9EA94020C8F4FFF32F984FE94D3F8" bold="true" box="[192,360,1653,1676]" pageId="13" pageNumber="12">Figures 45–46:</emphasis>
|
||
<taxonomicName id="26DD4D05020C8F4FFE9CF986FDC1D3F8" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[366,573,1655,1676]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA94020C8F4FFE9CF986FDC1D3F8" box="[366,573,1655,1676]" italics="true" pageId="13" pageNumber="12">Chaerilus neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF020C8F4FFDB1F986FD84D3F8" box="[579,632,1655,1676]" pageId="13" pageNumber="12" rank="species">
|
||
<emphasis id="D3A9EA94020C8F4FFDB1F986FDA6D3F8" bold="true" box="[579,602,1655,1676]" pageId="13" pageNumber="12">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA94020C8F4FFD94F986FD8FD3F8" bold="true" box="[614,627,1655,1676]" pageId="13" pageNumber="12">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype male, dorsal (45) and ventral (46) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E1623686020C8F4FFF32F940FD4DD393" blockId="13.[192,792,1713,1767]" pageId="13" pageNumber="12">carinae, second to fourth segments with 8 carinae. All metasomal segments very sparsely granulated.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9C7650D020C8F56FF32F8FCFD41D3CE" lastPageId="20" lastPageNumber="19" pageId="13" pageNumber="12" type="description">
|
||
<paragraph id="E1623686020C8F4FFF32F8FCFC5CD250" blockId="13.[192,793,1805,1890]" lastBlockId="13.[840,1440,1713,1890]" pageId="13" pageNumber="12">
|
||
DESCRIPTION. Total length of male
|
||
<typeStatus id="3E668824020C8F4FFDCDF8FCFD61D250" box="[575,669,1805,1828]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
<quantity id="26259B63020C8F4FFD56F8FCFCEDD250" box="[676,785,1805,1828]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" pageId="13" pageNumber="12" unit="mm" value="23.25">23.25 mm</quantity>
|
||
, female unknown. Two well developed pairs of lateral eyes and one pair of median eyes. The chelicerae are finely granulated, yellow and strongly reticulate, anteriorly black. For the position and distribution of trichobothria, see
|
||
<figureCitation id="79E62A03020C8F4FFBF2F91EFB76D272" box="[1024,1162,1775,1798]" captionStart="Figures 52–66" captionStartId="15.[192,278,1734,1757]" captionTargetBox="[204,1435,217,1702]" captionTargetId="figure-6@15.[192,1439,212,1704]" captionTargetPageId="15" captionText="Figures 52–66: Chaerilus neradorum sp. n., holotype male, pedipalp chela, dorsal (52), externodorsal (53), and ventrointernal (54) views. Pedipalp patella, dorsal (55), external (56) and ventral (57) views. Pedipalp femur and trochanter, dorsal (58) and ventral (59) views. Movable (60) and fixed (61) fingers. Left legs I–IV, retrolateral/ventral aspects (62–66). The trichobothrial pattern is indicated in Figures 52–58." pageId="13" pageNumber="12">Figs. 52–58</figureCitation>
|
||
. For measurements, see
|
||
<tableCitation id="AC5F033D020C8F4FFCBAF8FCFC5CD250" box="[840,928,1805,1828]" captionStart="Table 1" captionStartId="17.[192,257,818,841]" captionText="Table 1: Comparative measurements of Chaerilus majkusi sp. n. and Chaerilus neradorum sp. n. types. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="13" pageNumber="12">Table 1.</tableCitation>
|
||
</paragraph>
|
||
<paragraph id="E1623686020C8F4FFCBAF8DDFC12D216" blockId="13.[840,1440,1713,1890]" pageId="13" pageNumber="12">
|
||
COLORATION (
|
||
<figureCitation id="79E62A03020C8F4FFC1FF8DDFB88D237" box="[1005,1140,1836,1859]" captionStart="Figures 45–46" captionStartId="13.[192,278,1653,1676]" captionTargetBox="[208,1429,213,1619]" captionTargetId="figure-124@13.[192,1439,192,1623]" captionTargetPageId="13" captionText="Figures 45–46: Chaerilus neradorum sp. n., holotype male, dorsal (45) and ventral (46) views. Scale bar: 10 mm." pageId="13" pageNumber="12">Figs. 45–46</figureCitation>
|
||
). The color is orange to brown, spotted.
|
||
</paragraph>
|
||
<caption id="B5A2660E020F8F4CFF32F90BFB71D25E" pageId="14" pageNumber="13" startId="14.[192,278,1786,1809]" targetBox="[191,1440,191,1757]" targetPageId="14">
|
||
<paragraph id="E1623686020F8F4CFF32F90BFB71D25E" blockId="14.[192,1440,1786,1834]" pageId="14" pageNumber="13">
|
||
<emphasis id="D3A9EA94020F8F4CFF32F90BFE94D265" bold="true" box="[192,360,1786,1809]" pageId="14" pageNumber="13">Figures 47–51:</emphasis>
|
||
<taxonomicName id="26DD4D05020F8F4CFE9DF90DFDC3D265" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[367,575,1788,1809]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA94020F8F4CFE9DF90DFDC3D265" box="[367,575,1788,1809]" italics="true" pageId="14" pageNumber="13">Chaerilus neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF020F8F4CFDB7F90DFD87D265" box="[581,635,1788,1809]" pageId="14" pageNumber="13" rank="species">
|
||
<emphasis id="D3A9EA94020F8F4CFDB7F90DFDA0D265" bold="true" box="[581,604,1788,1809]" pageId="14" pageNumber="13">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA94020F8F4CFD9BF90DFD8AD265" bold="true" box="[617,630,1788,1809]" pageId="14" pageNumber="13">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype male, carapace and tergites I–III (47), sternopectinal region and sternites (48), metasoma and telson, lateral (49), ventral (50), and dorsal (51) views. Scale bar: 5 mm (49–51).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B5A2660E020E8F4DFF32F937FDDBD253" pageId="15" pageNumber="14" startId="15.[192,278,1734,1757]" targetBox="[204,1435,217,1702]" targetPageId="15">
|
||
<paragraph id="E1623686020E8F4DFF32F937FDDBD253" blockId="15.[192,1441,1734,1831]" pageId="15" pageNumber="14">
|
||
<emphasis id="D3A9EA94020E8F4DFF32F937FE95D3A9" bold="true" box="[192,361,1734,1757]" pageId="15" pageNumber="14">Figures 52–66:</emphasis>
|
||
<taxonomicName id="26DD4D05020E8F4DFE82F936FDBDD3A8" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[368,577,1735,1756]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA94020E8F4DFE82F936FDBDD3A8" box="[368,577,1735,1756]" italics="true" pageId="15" pageNumber="14">Chaerilus neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF020E8F4DFDB5F936FD81D3A8" box="[583,637,1735,1756]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="D3A9EA94020E8F4DFDB5F936FDA2D3A8" bold="true" box="[583,606,1735,1756]" pageId="15" pageNumber="14">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA94020E8F4DFD99F936FD84D3A8" bold="true" box="[619,632,1735,1756]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype male, pedipalp chela, dorsal (52), externodorsal (53), and ventrointernal (5 4) views. Pedipalp patella, dorsal (55), external (56) and ventral (57) views. Pedipalp femur and trochanter, dorsal (58) and ventral (59) views. Movable (60) and fixed (61) fingers. Left legs I–IV, retrolateral/ventral aspects (62–66). The trichobothrial pattern is indicated in Figures 52–58.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B5A2660E02118F52FF32F949FEF1D393" pageId="16" pageNumber="15" startId="16.[192,278,1720,1743]" targetBox="[191,1440,191,1690]" targetPageId="16">
|
||
<paragraph id="E162368602118F52FF32F949FEF1D393" blockId="16.[192,1441,1720,1767]" pageId="16" pageNumber="15">
|
||
<emphasis id="D3A9EA9402118F52FF32F949FE97D3BB" bold="true" box="[192,363,1720,1743]" pageId="16" pageNumber="15">Figures 67–68:</emphasis>
|
||
<taxonomicName id="26DD4D0502118F52FE86F94BFDBAD3BB" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[372,582,1722,1743]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402118F52FE86F94BFDBAD3BB" box="[372,582,1722,1743]" italics="true" pageId="16" pageNumber="15">Chaerilus neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02118F52FDBDF94BFD7BD3BB" box="[591,647,1722,1743]" pageId="16" pageNumber="15" rank="species">
|
||
<emphasis id="D3A9EA9402118F52FDBDF94BFD9AD3BB" bold="true" box="[591,614,1722,1743]" pageId="16" pageNumber="15">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402118F52FD87F94BFD7ED3BB" bold="true" box="[629,642,1722,1743]" pageId="16" pageNumber="15">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype (67) and
|
||
<taxonomicName id="26DD4D0502118F52FC7AF94BFB1ED3BB" authority="Kovarik, 2012" authorityName="Kovarik" authorityYear="2012" box="[904,1250,1722,1743]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="cimrmani">
|
||
<emphasis id="D3A9EA9402118F52FC7AF94BFBB0D3BB" box="[904,1100,1722,1743]" italics="true" pageId="16" pageNumber="15">Chaerilus cimrmani</emphasis>
|
||
<bibRefCitation id="854C4B7702118F52FBA7F94BFB1ED3BB" author="KOVARIK" box="[1109,1250,1722,1743]" pageId="16" pageNumber="15" pagination="1 - 14" refId="ref7828" refString="KOVARIK, F. 2012. Five New Species of Chaerilus Simon, 1877 from China, Indonesia, Malaysia, Philippines, Thailand, and Vietnam (Scorpiones: Chaerilidae). Euscorpius, 149: 1 - 14." type="journal article" year="2012">Kovařík, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (68) in vivo habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E162368602118F53FF32F8FCFE1BD6B8" blockId="16.[192,792,1805,1889]" lastBlockId="17.[192,793,918,1861]" lastPageId="17" lastPageNumber="16" pageId="16" pageNumber="15">
|
||
CARAPACE AND MESOSOMA (
|
||
<figureCitation id="79E62A0302118F52FDF4F8FCFD77D250" box="[518,651,1805,1828]" captionStart="Figures 47–51" captionStartId="14.[192,278,1786,1809]" captionTargetBox="[191,1440,191,1757]" captionTargetId="figure-76@14.[192,1439,192,1756]" captionTargetPageId="14" captionText="Figures 47–51: Chaerilus neradorum sp. n., holotype male, carapace and tergites I–III (47), sternopectinal region and sternites (48), metasoma and telson, lateral (49), ventral (50), and dorsal (51) views. Scale bar: 5 mm (49–51)." pageId="16" pageNumber="15">Figs. 47–48</figureCitation>
|
||
). The entire carapace is covered by large granules which do not form carinae. The anterior margin of the carapace is almost straight to weakly concave. The mesosomal tergites are irregularly, sparsely granulated. All sternites are smooth without carinae and granulation. Sternite V with smooth patch weakly indicated. Pectinal teeth number
|
||
<quantity id="26259B6302108F53FEEAFC44FEBDD6B8" box="[280,321,949,972]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="17" pageNumber="16" unit="in" value="6.0">6 in</quantity>
|
||
male
|
||
<typeStatus id="3E66882402108F53FE70FC44FE1FD6B8" box="[386,483,949,972]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E162368602108F53FCE6FF39FAF6D664" pageId="17" pageNumber="16">
|
||
<table id="93DDC426021070BDFEF6FF39FAA7D664" box="[260,1371,200,784]" gridcols="5" gridrows="19" pageId="17" pageNumber="16">
|
||
<tr id="5FED34C4021070BDFEF6FF39FAA7D5AB" box="[260,1371,200,223]" gridrow="0" pageId="17" pageNumber="16" rowspan-0="1" rowspan-1="1">
|
||
<th id="1C3C5DB8021070BDFD70FF39FBA0D5AB" box="[642,1116,200,223]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="17" pageNumber="16">
|
||
<taxonomicName id="26DD4D0502108F53FCE6FF39FC7BD5AB" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[788,903,200,223]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FCE6FF39FC7BD5AB" box="[788,903,200,223]" italics="true" pageId="17" pageNumber="16">C. majkusi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D3A9EA9402108F53FC7FFF39FC35D5AB" bold="true" box="[909,969,200,223]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="C89A57EF02108F53FC7FFF39FC35D5AB" box="[909,969,200,223]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="1C3C5DB8021070BDFB8FFF39FAA7D5AB" box="[1149,1371,200,223]" gridcol="4" gridrow="0" pageId="17" pageNumber="16">
|
||
<taxonomicName id="26DD4D0502108F53FB72FF39FAE9D5AB" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[1152,1301,200,223]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FB72FF39FAE9D5AB" box="[1152,1301,200,223]" italics="true" pageId="17" pageNumber="16">C. neradorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D3A9EA9402108F53FAEEFF39FAA4D5AB" bold="true" box="[1308,1368,200,223]" pageId="17" pageNumber="16">
|
||
<taxonomicNameLabel id="C89A57EF02108F53FAEEFF39FAA4D5AB" box="[1308,1368,200,223]" pageId="17" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FF16FAA7D58A" box="[260,1371,231,254]" gridrow="1" pageId="17" pageNumber="16" rowspan-1="1">
|
||
<th id="1C3C5DB8021070BDFEF6FF16FE19D58A" box="[260,485,231,254]" gridcol="0" gridrow="1" pageId="17" pageNumber="16">
|
||
<emphasis id="D3A9EA9402108F53FEE5FF16FE19D58A" bold="true" box="[279,485,231,254]" pageId="17" pageNumber="16">Dimensions (MM)</emphasis>
|
||
</th>
|
||
<td id="1C3C5DB8021070BDFD70FF16FC9CD58A" box="[642,864,231,254]" gridcol="2" gridrow="1" pageId="17" pageNumber="16">
|
||
♂
|
||
<emphasis id="D3A9EA9402108F53FD5CFF16FCECD58A" bold="true" box="[686,784,231,254]" pageId="17" pageNumber="16">holotype</emphasis>
|
||
</td>
|
||
<td id="1C3C5DB8021070BDFC8CFF16FBA0D58A" box="[894,1116,231,254]" gridcol="3" gridrow="1" pageId="17" pageNumber="16">
|
||
<emphasis id="D3A9EA9402108F53FC38FF16FBB7D58A" bold="true" box="[970,1099,231,254]" pageId="17" pageNumber="16">♀ paratype</emphasis>
|
||
</td>
|
||
<td id="1C3C5DB8021070BDFB8FFF16FAA7D58A" box="[1149,1371,231,254]" gridcol="4" gridrow="1" pageId="17" pageNumber="16">
|
||
♂
|
||
<emphasis id="D3A9EA9402108F53FB3AFF16FAD6D58A" bold="true" box="[1224,1322,231,254]" pageId="17" pageNumber="16">holotype</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FEFBFAA7D455" box="[260,1371,266,289]" gridrow="2" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FEFBFE19D455" box="[260,485,266,289]" gridcol="0" gridrow="2" pageId="17" pageNumber="16">Carapace</th>
|
||
<td id="1C3C5DB8021070BDFDF3FEFBFD9BD455" box="[513,615,266,289]" gridcol="1" gridrow="2" pageId="17" pageNumber="16">L / W</td>
|
||
<td id="1C3C5DB8021070BDFD70FEFBFC9CD455" box="[642,864,266,289]" gridcol="2" gridrow="2" pageId="17" pageNumber="16">3.925 / 4.175</td>
|
||
<td id="1C3C5DB8021070BDFC8CFEFBFBA0D455" box="[894,1116,266,289]" gridcol="3" gridrow="2" pageId="17" pageNumber="16">3.825 / 4.175</td>
|
||
<td id="1C3C5DB8021070BDFB8FFEFBFAA7D455" box="[1149,1371,266,289]" gridcol="4" gridrow="2" pageId="17" pageNumber="16">3.850 / 3.550</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FED9FAA7D44B" box="[260,1371,296,319]" gridrow="3" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FED9FE19D44B" box="[260,485,296,319]" gridcol="0" gridrow="3" pageId="17" pageNumber="16">Mesosoma</th>
|
||
<td id="1C3C5DB8021070BDFDF3FED9FD9BD44B" box="[513,615,296,319]" gridcol="1" gridrow="3" pageId="17" pageNumber="16">L</td>
|
||
<td id="1C3C5DB8021070BDFD70FED9FC9CD44B" box="[642,864,296,319]" gridcol="2" gridrow="3" pageId="17" pageNumber="16">5.950</td>
|
||
<td id="1C3C5DB8021070BDFC8CFED9FBA0D44B" box="[894,1116,296,319]" gridcol="3" gridrow="3" pageId="17" pageNumber="16">10.300</td>
|
||
<td id="1C3C5DB8021070BDFB8FFED9FAA7D44B" box="[1149,1371,296,319]" gridcol="4" gridrow="3" pageId="17" pageNumber="16">5.300</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FEB6FAA7D42A" box="[260,1371,327,350]" gridrow="4" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FEB6FE19D42A" box="[260,485,327,350]" gridcol="0" gridrow="4" pageId="17" pageNumber="16">Tergite VII</th>
|
||
<td id="1C3C5DB8021070BDFDF3FEB6FD9BD42A" box="[513,615,327,350]" gridcol="1" gridrow="4" pageId="17" pageNumber="16">L / W</td>
|
||
<td id="1C3C5DB8021070BDFD70FEB6FC9CD42A" box="[642,864,327,350]" gridcol="2" gridrow="4" pageId="17" pageNumber="16">1.375 / 3.525</td>
|
||
<td id="1C3C5DB8021070BDFC8CFEB6FBA0D42A" box="[894,1116,327,350]" gridcol="3" gridrow="4" pageId="17" pageNumber="16">1.950 / 3.950</td>
|
||
<td id="1C3C5DB8021070BDFB8FFEB6FAA7D42A" box="[1149,1371,327,350]" gridcol="4" gridrow="4" pageId="17" pageNumber="16">2.075 / 3.200</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FE97FAA7D409" box="[260,1371,358,381]" gridrow="5" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FE97FE19D409" box="[260,485,358,381]" gridcol="0" gridrow="5" pageId="17" pageNumber="16">Metasoma & telson</th>
|
||
<td id="1C3C5DB8021070BDFDF3FE97FD9BD409" box="[513,615,358,381]" gridcol="1" gridrow="5" pageId="17" pageNumber="16">L</td>
|
||
<td id="1C3C5DB8021070BDFD70FE97FC9CD409" box="[642,864,358,381]" gridcol="2" gridrow="5" pageId="17" pageNumber="16">17.025</td>
|
||
<td id="1C3C5DB8021070BDFC8CFE97FBA0D409" box="[894,1116,358,381]" gridcol="3" gridrow="5" pageId="17" pageNumber="16">14.625</td>
|
||
<td id="1C3C5DB8021070BDFB8FFE97FAA7D409" box="[1149,1371,358,381]" gridcol="4" gridrow="5" pageId="17" pageNumber="16">14.100</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FE75FAA7D4EF" box="[260,1371,388,411]" gridrow="6" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FE75FE19D4EF" box="[260,485,388,411]" gridcol="0" gridrow="6" pageId="17" pageNumber="16">Segment I</th>
|
||
<td id="1C3C5DB8021070BDFDF3FE75FD9BD4EF" box="[513,615,388,411]" gridcol="1" gridrow="6" pageId="17" pageNumber="16">L / W / D</td>
|
||
<td id="1C3C5DB8021070BDFD70FE75FC9CD4EF" box="[642,864,388,411]" gridcol="2" gridrow="6" pageId="17" pageNumber="16">1.675 / 2.425 / 1.938</td>
|
||
<td id="1C3C5DB8021070BDFC8CFE75FBA0D4EF" box="[894,1116,388,411]" gridcol="3" gridrow="6" pageId="17" pageNumber="16">1.325 / 2.258 / 1.875</td>
|
||
<td id="1C3C5DB8021070BDFB8FFE75FAA7D4EF" box="[1149,1371,388,411]" gridcol="4" gridrow="6" pageId="17" pageNumber="16">1.350 / 2.250 / 1.650</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FE52FAA7D4CE" box="[260,1371,419,442]" gridrow="7" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FE52FE19D4CE" box="[260,485,419,442]" gridcol="0" gridrow="7" pageId="17" pageNumber="16">Segment II</th>
|
||
<td id="1C3C5DB8021070BDFDF3FE52FD9BD4CE" box="[513,615,419,442]" gridcol="1" gridrow="7" pageId="17" pageNumber="16">L / W / D</td>
|
||
<td id="1C3C5DB8021070BDFD70FE52FC9CD4CE" box="[642,864,419,442]" gridcol="2" gridrow="7" pageId="17" pageNumber="16">1.800 / 2.013 / 1.838</td>
|
||
<td id="1C3C5DB8021070BDFC8CFE52FBA0D4CE" box="[894,1116,419,442]" gridcol="3" gridrow="7" pageId="17" pageNumber="16">1.550 / 1.950 / 1.700</td>
|
||
<td id="1C3C5DB8021070BDFB8FFE52FAA7D4CE" box="[1149,1371,419,442]" gridcol="4" gridrow="7" pageId="17" pageNumber="16">1.675 / 1.900 / 1.475</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FE33FAA7D4AD" box="[260,1371,450,473]" gridrow="8" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FE33FE19D4AD" box="[260,485,450,473]" gridcol="0" gridrow="8" pageId="17" pageNumber="16">Segment III</th>
|
||
<td id="1C3C5DB8021070BDFDF3FE33FD9BD4AD" box="[513,615,450,473]" gridcol="1" gridrow="8" pageId="17" pageNumber="16">L / W / D</td>
|
||
<td id="1C3C5DB8021070BDFD70FE33FC9CD4AD" box="[642,864,450,473]" gridcol="2" gridrow="8" pageId="17" pageNumber="16">2.000 / 1.950 / 1.700</td>
|
||
<td id="1C3C5DB8021070BDFC8CFE33FBA0D4AD" box="[894,1116,450,473]" gridcol="3" gridrow="8" pageId="17" pageNumber="16">1.700 / 1.825 / 1.600</td>
|
||
<td id="1C3C5DB8021070BDFB8FFE33FAA7D4AD" box="[1149,1371,450,473]" gridcol="4" gridrow="8" pageId="17" pageNumber="16">1.750 / 1.775 / 1.450</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FE11FAA7D483" box="[260,1371,480,503]" gridrow="9" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FE11FE19D483" box="[260,485,480,503]" gridcol="0" gridrow="9" pageId="17" pageNumber="16">Segment IV</th>
|
||
<td id="1C3C5DB8021070BDFDF3FE11FD9BD483" box="[513,615,480,503]" gridcol="1" gridrow="9" pageId="17" pageNumber="16">L / W / D</td>
|
||
<td id="1C3C5DB8021070BDFD70FE11FC9CD483" box="[642,864,480,503]" gridcol="2" gridrow="9" pageId="17" pageNumber="16">2.325 / 1.850 / 1.675</td>
|
||
<td id="1C3C5DB8021070BDFC8CFE11FBA0D483" box="[894,1116,480,503]" gridcol="3" gridrow="9" pageId="17" pageNumber="16">1.950 / 1.725 / 1.550</td>
|
||
<td id="1C3C5DB8021070BDFB8FFE11FAA7D483" box="[1149,1371,480,503]" gridcol="4" gridrow="9" pageId="17" pageNumber="16">1.975 / 1.650 / 1.400</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FE0EFAA7D762" box="[260,1371,511,534]" gridrow="10" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FE0EFE19D762" box="[260,485,511,534]" gridcol="0" gridrow="10" pageId="17" pageNumber="16">Segment V</th>
|
||
<td id="1C3C5DB8021070BDFDF3FE0EFD9BD762" box="[513,615,511,534]" gridcol="1" gridrow="10" pageId="17" pageNumber="16">L / W / D</td>
|
||
<td id="1C3C5DB8021070BDFD70FE0EFC9CD762" box="[642,864,511,534]" gridcol="2" gridrow="10" pageId="17" pageNumber="16">4.325 / 1.850 / 1.750</td>
|
||
<td id="1C3C5DB8021070BDFC8CFE0EFBA0D762" box="[894,1116,511,534]" gridcol="3" gridrow="10" pageId="17" pageNumber="16">3.725 / 1.775 / 1.575</td>
|
||
<td id="1C3C5DB8021070BDFB8FFE0EFAA7D762" box="[1149,1371,511,534]" gridcol="4" gridrow="10" pageId="17" pageNumber="16">3.425 / 1.550 / 1.366</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FDEFFAA7D741" box="[260,1371,542,565]" gridrow="11" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FDEFFE19D741" box="[260,485,542,565]" gridcol="0" gridrow="11" pageId="17" pageNumber="16">Telson</th>
|
||
<td id="1C3C5DB8021070BDFDF3FDEFFD9BD741" box="[513,615,542,565]" gridcol="1" gridrow="11" pageId="17" pageNumber="16">L / W / D</td>
|
||
<td id="1C3C5DB8021070BDFD70FDEFFC9CD741" box="[642,864,542,565]" gridcol="2" gridrow="11" pageId="17" pageNumber="16">4.900 / 1.775 / 1.600</td>
|
||
<td id="1C3C5DB8021070BDFC8CFDEFFBA0D741" box="[894,1116,542,565]" gridcol="3" gridrow="11" pageId="17" pageNumber="16">4.375 / 1.750 / 1.525</td>
|
||
<td id="1C3C5DB8021070BDFB8FFDEFFAA7D741" box="[1149,1371,542,565]" gridcol="4" gridrow="11" pageId="17" pageNumber="16">3.925 / 1.625 / 1.463</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FDCDFAA7D727" box="[260,1371,572,595]" gridrow="12" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FDCDFE19D727" box="[260,485,572,595]" gridcol="0" gridrow="12" pageId="17" pageNumber="16">Pedipalp</th>
|
||
<td id="1C3C5DB8021070BDFDF3FDCDFD9BD727" box="[513,615,572,595]" gridcol="1" gridrow="12" pageId="17" pageNumber="16">L</td>
|
||
<td id="1C3C5DB8021070BDFD70FDCDFC9CD727" box="[642,864,572,595]" gridcol="2" gridrow="12" pageId="17" pageNumber="16">12.825</td>
|
||
<td id="1C3C5DB8021070BDFC8CFDCDFBA0D727" box="[894,1116,572,595]" gridcol="3" gridrow="12" pageId="17" pageNumber="16">13.325</td>
|
||
<td id="1C3C5DB8021070BDFB8FFDCDFAA7D727" box="[1149,1371,572,595]" gridcol="4" gridrow="12" pageId="17" pageNumber="16">11.375</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FDAAFAA7D706" box="[260,1371,603,626]" gridrow="13" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FDAAFE19D706" box="[260,485,603,626]" gridcol="0" gridrow="13" pageId="17" pageNumber="16">Femur</th>
|
||
<td id="1C3C5DB8021070BDFDF3FDAAFD9BD706" box="[513,615,603,626]" gridcol="1" gridrow="13" pageId="17" pageNumber="16">L / W</td>
|
||
<td id="1C3C5DB8021070BDFD70FDAAFC9CD706" box="[642,864,603,626]" gridcol="2" gridrow="13" pageId="17" pageNumber="16">3.025 / 1.375</td>
|
||
<td id="1C3C5DB8021070BDFC8CFDAAFBA0D706" box="[894,1116,603,626]" gridcol="3" gridrow="13" pageId="17" pageNumber="16">3.075 / 1.500</td>
|
||
<td id="1C3C5DB8021070BDFB8FFDAAFAA7D706" box="[1149,1371,603,626]" gridcol="4" gridrow="13" pageId="17" pageNumber="16">2.525 / 1.325</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FD8BFAA7D7E5" box="[260,1371,634,657]" gridrow="14" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FD8BFE19D7E5" box="[260,485,634,657]" gridcol="0" gridrow="14" pageId="17" pageNumber="16">Patella</th>
|
||
<td id="1C3C5DB8021070BDFDF3FD8BFD9BD7E5" box="[513,615,634,657]" gridcol="1" gridrow="14" pageId="17" pageNumber="16">L / W</td>
|
||
<td id="1C3C5DB8021070BDFD70FD8BFC9CD7E5" box="[642,864,634,657]" gridcol="2" gridrow="14" pageId="17" pageNumber="16">3.250 / 1.500</td>
|
||
<td id="1C3C5DB8021070BDFC8CFD8BFBA0D7E5" box="[894,1116,634,657]" gridcol="3" gridrow="14" pageId="17" pageNumber="16">3.300 / 1.675</td>
|
||
<td id="1C3C5DB8021070BDFB8FFD8BFAA7D7E5" box="[1149,1371,634,657]" gridcol="4" gridrow="14" pageId="17" pageNumber="16">2.900 / 1.600</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FD69FAA7D7DB" box="[260,1371,664,687]" gridrow="15" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FD69FE19D7DB" box="[260,485,664,687]" gridcol="0" gridrow="15" pageId="17" pageNumber="16">Chela</th>
|
||
<td id="1C3C5DB8021070BDFDF3FD69FD9BD7DB" box="[513,615,664,687]" gridcol="1" gridrow="15" pageId="17" pageNumber="16">L</td>
|
||
<td id="1C3C5DB8021070BDFD70FD69FC9CD7DB" box="[642,864,664,687]" gridcol="2" gridrow="15" pageId="17" pageNumber="16">6.550</td>
|
||
<td id="1C3C5DB8021070BDFC8CFD69FBA0D7DB" box="[894,1116,664,687]" gridcol="3" gridrow="15" pageId="17" pageNumber="16">6.950</td>
|
||
<td id="1C3C5DB8021070BDFB8FFD69FAA7D7DB" box="[1149,1371,664,687]" gridcol="4" gridrow="15" pageId="17" pageNumber="16">5.950</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FD46FAA7D7BA" box="[260,1371,695,718]" gridrow="16" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FD46FE19D7BA" box="[260,485,695,718]" gridcol="0" gridrow="16" pageId="17" pageNumber="16">Manus</th>
|
||
<td id="1C3C5DB8021070BDFDF3FD46FD9BD7BA" box="[513,615,695,718]" gridcol="1" gridrow="16" pageId="17" pageNumber="16">W / D</td>
|
||
<td id="1C3C5DB8021070BDFD70FD46FC9CD7BA" box="[642,864,695,718]" gridcol="2" gridrow="16" pageId="17" pageNumber="16">3.500 / 2.700</td>
|
||
<td id="1C3C5DB8021070BDFC8CFD46FBA0D7BA" box="[894,1116,695,718]" gridcol="3" gridrow="16" pageId="17" pageNumber="16">3.050 / 2.500</td>
|
||
<td id="1C3C5DB8021070BDFB8FFD46FAA7D7BA" box="[1149,1371,695,718]" gridcol="4" gridrow="16" pageId="17" pageNumber="16">2.475 / 2.125</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FD27FAA7D799" box="[260,1371,726,749]" gridrow="17" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FD27FE19D799" box="[260,485,726,749]" gridcol="0" gridrow="17" pageId="17" pageNumber="16">Movable finger</th>
|
||
<td id="1C3C5DB8021070BDFDF3FD27FD9BD799" box="[513,615,726,749]" gridcol="1" gridrow="17" pageId="17" pageNumber="16">L</td>
|
||
<td id="1C3C5DB8021070BDFD70FD27FC9CD799" box="[642,864,726,749]" gridcol="2" gridrow="17" pageId="17" pageNumber="16">3.425</td>
|
||
<td id="1C3C5DB8021070BDFC8CFD27FBA0D799" box="[894,1116,726,749]" gridcol="3" gridrow="17" pageId="17" pageNumber="16">3.925</td>
|
||
<td id="1C3C5DB8021070BDFB8FFD27FAA7D799" box="[1149,1371,726,749]" gridcol="4" gridrow="17" pageId="17" pageNumber="16">3.175</td>
|
||
</tr>
|
||
<tr id="5FED34C4021070BDFEF6FD08FAA7D664" box="[260,1371,761,784]" gridrow="18" pageId="17" pageNumber="16">
|
||
<th id="1C3C5DB8021070BDFEF6FD08FE19D664" box="[260,485,761,784]" gridcol="0" gridrow="18" pageId="17" pageNumber="16">Total</th>
|
||
<td id="1C3C5DB8021070BDFDF3FD08FD9BD664" box="[513,615,761,784]" gridcol="1" gridrow="18" pageId="17" pageNumber="16">L</td>
|
||
<td id="1C3C5DB8021070BDFD70FD08FC9CD664" box="[642,864,761,784]" gridcol="2" gridrow="18" pageId="17" pageNumber="16">26.90</td>
|
||
<td id="1C3C5DB8021070BDFC8CFD08FBA0D664" box="[894,1116,761,784]" gridcol="3" gridrow="18" pageId="17" pageNumber="16">28.75</td>
|
||
<td id="1C3C5DB8021070BDFB8FFD08FAA7D664" box="[1149,1371,761,784]" gridcol="4" gridrow="18" pageId="17" pageNumber="16">23.25</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="B5A2660E02108F53FF32FCC3FC88D615" pageId="17" pageNumber="16" startId="17.[192,257,818,841]">
|
||
<paragraph id="E162368602108F53FF32FCC3FC88D615" blockId="17.[192,1440,818,865]" pageId="17" pageNumber="16">
|
||
<emphasis id="D3A9EA9402108F53FF32FCC3FEDCD63D" bold="true" box="[192,288,818,841]" pageId="17" pageNumber="16">Table 1:</emphasis>
|
||
Comparative measurements of
|
||
<taxonomicName id="26DD4D0502108F53FDAAFCC2FCF5D63C" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[600,777,819,840]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FDAAFCC2FCF5D63C" box="[600,777,819,840]" italics="true" pageId="17" pageNumber="16">Chaerilus majkusi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02108F53FCE2FCC2FCBAD63C" box="[784,838,819,840]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="D3A9EA9402108F53FCE2FCC2FCD9D63C" bold="true" box="[784,805,819,840]" pageId="17" pageNumber="16">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402108F53FCC1FCC2FCBCD63C" bold="true" box="[819,832,819,840]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="26DD4D0502108F53FC8BFCC2FBB6D63C" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[889,1098,819,840]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FC8BFCC2FBB6D63C" box="[889,1098,819,840]" italics="true" pageId="17" pageNumber="16">Chaerilus neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02108F53FBBDFCC2FB7BD63C" box="[1103,1159,819,840]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="D3A9EA9402108F53FBBDFCC2FB9AD63C" bold="true" box="[1103,1126,819,840]" pageId="17" pageNumber="16">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402108F53FB81FCC2FB7CD63C" bold="true" box="[1139,1152,819,840]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
types. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E162368602108F53FF32FC22FE92D069" blockId="17.[192,793,918,1861]" pageId="17" pageNumber="16">
|
||
METASOMA AND TELSON (
|
||
<figureCitation id="79E62A0302108F53FE2EFC22FDA2D69E" box="[476,606,979,1002]" captionStart="Figures 47–51" captionStartId="14.[192,278,1786,1809]" captionTargetBox="[191,1440,191,1757]" captionTargetId="figure-76@14.[192,1439,192,1756]" captionTargetPageId="14" captionText="Figures 47–51: Chaerilus neradorum sp. n., holotype male, carapace and tergites I–III (47), sternopectinal region and sternites (48), metasoma and telson, lateral (49), ventral (50), and dorsal (51) views. Scale bar: 5 mm (49–51)." pageId="17" pageNumber="16">Figs. 49–51</figureCitation>
|
||
). The first metasomal segment bears 10 carinae, the second to fourth bear eight carinae, and the fifth segment bears seven carinae of which one ventral carina posteriorly branches in a "Y" configuration. All carinae are composed of large, sparse granules. The spaces between carinae are irregularly very sparsely granulated on all surfaces, less so on the dorsal surface. Granules on the dorsal surface may indicated a pair of carinae. All segments are sparsely hirsute. The telson is elongate, smooth and sparsely hirsute.
|
||
</paragraph>
|
||
<paragraph id="E162368602108F53FF32FAD4FE53D324" blockId="17.[192,793,918,1861]" pageId="17" pageNumber="16">
|
||
PEDIPALPS (
|
||
<figureCitation id="79E62A0302108F53FEB7FAD4FE36D048" box="[325,458,1317,1340]" captionStart="Figures 52–66" captionStartId="15.[192,278,1734,1757]" captionTargetBox="[204,1435,217,1702]" captionTargetId="figure-6@15.[192,1439,212,1704]" captionTargetPageId="15" captionText="Figures 52–66: Chaerilus neradorum sp. n., holotype male, pedipalp chela, dorsal (52), externodorsal (53), and ventrointernal (54) views. Pedipalp patella, dorsal (55), external (56) and ventral (57) views. Pedipalp femur and trochanter, dorsal (58) and ventral (59) views. Movable (60) and fixed (61) fingers. Left legs I–IV, retrolateral/ventral aspects (62–66). The trichobothrial pattern is indicated in Figures 52–58." pageId="17" pageNumber="16">Figs. 52–61</figureCitation>
|
||
). The pedipalp chela is wide and swollen in the male. The movable and fixed fingers of pedipalp bear 11 imbricated rows of granules. The chela has eight partly granulated carinae. The patella has five only weakly indicated carinae and the femur has four or five partly granulated carinae. The spaces between carinae on the femur are covered by unevenly spaced granules. The chela is finely granulated and the patella is smooth except for several solitary granules on their internal surfaces.
|
||
</paragraph>
|
||
<paragraph id="E162368602108F53FF32F9A6FE4DD231" blockId="17.[192,793,918,1861]" pageId="17" pageNumber="16">
|
||
LEGS (
|
||
<figureCitation id="79E62A0302108F53FEFAF9A6FE74D31A" box="[264,392,1623,1646]" captionStart="Figures 52–66" captionStartId="15.[192,278,1734,1757]" captionTargetBox="[204,1435,217,1702]" captionTargetId="figure-6@15.[192,1439,212,1704]" captionTargetPageId="15" captionText="Figures 52–66: Chaerilus neradorum sp. n., holotype male, pedipalp chela, dorsal (52), externodorsal (53), and ventrointernal (54) views. Pedipalp patella, dorsal (55), external (56) and ventral (57) views. Pedipalp femur and trochanter, dorsal (58) and ventral (59) views. Movable (60) and fixed (61) fingers. Left legs I–IV, retrolateral/ventral aspects (62–66). The trichobothrial pattern is indicated in Figures 52–58." pageId="17" pageNumber="16">Figs. 62–66</figureCitation>
|
||
). The legs are sparsely hirsute, without bristlecombs and carinae. The femora and patellae have several granules dorsally, other surfaces are smooth. The patellae have several median spines dorsally. The tarsomeres bear two rows of spiniform setae and 2 – 4 outer spiniform setae. Spiniform setae formula is 5–6/6–7: 6–7/7–8: 7–8/8–9: 7–8/7–9 (omitting outer spiniform setae).
|
||
</paragraph>
|
||
<paragraph id="E162368602108F53FCBAFC67FB67D00D" blockId="17.[840,1441,918,1401]" pageId="17" pageNumber="16">
|
||
KARYOTYPE (
|
||
<figureCitation id="79E62A0302108F53FC2DFC67FB81D6D9" box="[991,1149,918,941]" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="17" pageNumber="16">Figs. 117–118</figureCitation>
|
||
). We analyzed the karyotype of the male
|
||
<typeStatus id="3E66882402108F53FBF5FC44FB94D6B8" box="[1031,1128,949,972]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
. The diploid complement of this species is 108 chromosomes in the majority of mitotic metaphases (
|
||
<figureCitation id="79E62A0302108F53FBDFFC03FB70D17D" box="[1069,1164,1010,1033]" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="17" pageNumber="16">Fig. 117</figureCitation>
|
||
). However, the chromosomes are small and the correct number is not easy to count in some observed cells as a consequence of an early segregation of some chromatids during mitotic metaphase. This effect is evident especially in segregation of chromosomes during late metaphase I (
|
||
<figureCitation id="79E62A0302108F53FA84FB7AFC8AD1B5" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="17" pageNumber="16">Fig. 118</figureCitation>
|
||
). It is the reason we cannot exclude even 106 as the correct diploid number of chromosomes in this species. We are not able to specify morphology of the chromosomes at this moment. However, it is evident that they gradually decrease in length from 2.63% to 1.07% of the haploid set. We did not observe chiasmata during postpachytene and metaphase I.
|
||
</paragraph>
|
||
<paragraph id="E162368602108F53FCBAFA6EFAFED324" blockId="17.[840,1441,1439,1861]" pageId="17" pageNumber="16">
|
||
AFFINITIES.
|
||
<taxonomicName id="26DD4D0502108F53FC3FFA6EFB4AD0C2" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[973,1206,1439,1462]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FC3FFA6EFB4AD0C2" box="[973,1206,1439,1462]" italics="true" pageId="17" pageNumber="16">Chaerilus neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02108F53FB4DFA6EFB00D0C2" box="[1215,1276,1439,1462]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="D3A9EA9402108F53FB4DFA6EFB2AD0C2" bold="true" box="[1215,1238,1439,1462]" pageId="17" pageNumber="16">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402108F53FB14FA6EFB09D0C2" bold="true" box="[1254,1269,1439,1462]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is reliably distinguished from all other
|
||
<taxonomicName id="26DD4D0502108F53FB8CFA4FFAB8D0A1" box="[1150,1348,1470,1493]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="D3A9EA9402108F53FB8CFA4FFB1BD0A1" box="[1150,1255,1470,1493]" italics="true" pageId="17" pageNumber="16">Chaerilus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the following unique combination characters: total length of male
|
||
<typeStatus id="3E66882402108F53FC50FA0AFBFCD366" box="[930,1024,1531,1554]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
<quantity id="26259B6302108F53FBFAFA0AFB89D366" box="[1032,1141,1531,1554]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" pageId="17" pageNumber="16" unit="mm" value="23.25">23.25 mm</quantity>
|
||
; movable and fixed fingers of pedipalp with 11 cutting edges; chela length/width ratio in male 2.4; fingers straight in male.
|
||
</paragraph>
|
||
<paragraph id="E162368602108F56FC8AF9A6FD41D3CE" blockId="17.[840,1441,1439,1861]" lastBlockId="20.[192,792,1669,1722]" lastPageId="20" lastPageNumber="19" pageId="17" pageNumber="16">
|
||
Type locality of
|
||
<taxonomicName id="26DD4D0502108F53FBC0F9A6FB37D31A" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[1074,1227,1623,1646]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FBC0F9A6FB37D31A" box="[1074,1227,1623,1646]" italics="true" pageId="17" pageNumber="16">C. neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02108F53FB20F9A6FAEDD31A" box="[1234,1297,1623,1646]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="D3A9EA9402108F53FB20F9A6FB17D31A" bold="true" box="[1234,1259,1623,1646]" pageId="17" pageNumber="16">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402108F53FB09F9A6FAF6D31A" bold="true" box="[1275,1290,1623,1646]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is inside the area of distribution of
|
||
<taxonomicName id="26DD4D0502108F53FBAFF987FA63D3F9" authority="Kovarik, 2012" authorityName="Kovarik" authorityYear="2012" box="[1117,1439,1654,1677]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="cimrmani">
|
||
<emphasis id="D3A9EA9402108F53FBAFF987FB10D3F9" box="[1117,1260,1654,1677]" italics="true" pageId="17" pageNumber="16">C. cimrmani</emphasis>
|
||
<bibRefCitation id="854C4B7702108F53FB0EF987FA63D3F9" author="KOVARIK" box="[1276,1439,1654,1677]" pageId="17" pageNumber="16" pagination="1 - 14" refId="ref7828" refString="KOVARIK, F. 2012. Five New Species of Chaerilus Simon, 1877 from China, Indonesia, Malaysia, Philippines, Thailand, and Vietnam (Scorpiones: Chaerilidae). Euscorpius, 149: 1 - 14." type="journal article" year="2012">Kovařík, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
which can be distinguished from
|
||
<taxonomicName id="26DD4D0502108F53FB4DF964FAA5D3D8" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[1215,1369,1685,1708]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FB4DF964FAA5D3D8" box="[1215,1369,1685,1708]" italics="true" pageId="17" pageNumber="16">C. neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02108F53FA92F964FA5CD3D8" box="[1376,1440,1685,1708]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="D3A9EA9402108F53FA92F964FA85D3D8" bold="true" box="[1376,1401,1685,1708]" pageId="17" pageNumber="16">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402108F53FA79F964FA66D3D8" bold="true" box="[1419,1434,1685,1708]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by larger size (total length of male
|
||
<typeStatus id="3E66882402108F53FB3EF942FAD6D3BE" box="[1228,1322,1715,1738]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
<quantity id="26259B6302108F53FAC0F942FA5DD3BE" box="[1330,1441,1715,1738]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" pageId="17" pageNumber="16" unit="mm" value="23.25">23.25 mm</quantity>
|
||
for
|
||
<taxonomicName id="26DD4D0502108F53FC9CF923FBF9D39D" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[878,1029,1746,1769]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402108F53FC9CF923FBF9D39D" box="[878,1029,1746,1769]" italics="true" pageId="17" pageNumber="16">C. neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02108F53FBF9F923FBB4D39D" box="[1035,1096,1746,1769]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="D3A9EA9402108F53FBF9F923FBD8D39D" bold="true" box="[1035,1060,1746,1769]" pageId="17" pageNumber="16">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402108F53FBC0F923FBBDD39D" bold="true" box="[1074,1089,1746,1769]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
vs.
|
||
<quantity id="26259B6302108F53FB86F923FB1AD39D" box="[1140,1254,1746,1769]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.2" metricValueMin="3.1" pageId="17" pageNumber="16" unit="mm" value="36.5" valueMax="42.0" valueMin="31.0">31–42 mm</quantity>
|
||
of
|
||
<taxonomicName id="26DD4D0502108F53FAF9F923FA6DD39D" authorityName="Kovarik" authorityYear="2012" box="[1291,1425,1746,1769]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="cimrmani">
|
||
<emphasis id="D3A9EA9402108F53FAF9F923FA6DD39D" box="[1291,1425,1746,1769]" italics="true" pageId="17" pageNumber="16">C. cimrmani</emphasis>
|
||
</taxonomicName>
|
||
). Other differences are evident in comparing
|
||
<figureCitation id="79E62A0302108F53FAB1F900FA5CD27C" box="[1347,1440,1777,1800]" captionStart="Figures 52–66" captionStartId="15.[192,278,1734,1757]" captionTargetBox="[204,1435,217,1702]" captionTargetId="figure-6@15.[192,1439,212,1704]" captionTargetPageId="15" captionText="Figures 52–66: Chaerilus neradorum sp. n., holotype male, pedipalp chela, dorsal (52), externodorsal (53), and ventrointernal (54) views. Pedipalp patella, dorsal (55), external (56) and ventral (57) views. Pedipalp femur and trochanter, dorsal (58) and ventral (59) views. Movable (60) and fixed (61) fingers. Left legs I–IV, retrolateral/ventral aspects (62–66). The trichobothrial pattern is indicated in Figures 52–58." pageId="17" pageNumber="16">Figs. 52</figureCitation>
|
||
versus 105, which shows that the male of
|
||
<taxonomicName id="26DD4D0502108F53FAEAF8FEFA5CD252" authorityName="Kovarik" authorityYear="2012" box="[1304,1440,1807,1830]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="cimrmani">
|
||
<emphasis id="D3A9EA9402108F53FAEAF8FEFA5CD252" box="[1304,1440,1807,1830]" italics="true" pageId="17" pageNumber="16">C. cimrmani</emphasis>
|
||
</taxonomicName>
|
||
has wider, more swollen chelae, shorter chela fingers, and stronger granulation of carapace and metasomal segments, than the male of
|
||
<taxonomicName id="26DD4D0502158F56FE16F952FD85D3CE" authority="Kovařík & Lowe & Šťáhlavský, 2018" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2018" box="[484,633,1699,1722]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="neradorum" status="sp. nov.">
|
||
<emphasis id="D3A9EA9402158F56FE16F952FD85D3CE" box="[484,633,1699,1722]" italics="true" pageId="20" pageNumber="19">C. neradorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C89A57EF02158F56FD72F952FD41D3CE" box="[640,701,1699,1722]" pageId="20" pageNumber="19" rank="species">
|
||
<emphasis id="D3A9EA9402158F56FD72F952FD65D3CE" bold="true" box="[640,665,1699,1722]" pageId="20" pageNumber="19">sp</emphasis>
|
||
.
|
||
<emphasis id="D3A9EA9402158F56FD55F952FD4AD3CE" bold="true" box="[679,694,1699,1722]" pageId="20" pageNumber="19">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |