248 lines
26 KiB
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248 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.646.11397" ID-GBIF-Dataset="0ce110f8-a8ad-44d9-9c71-7c26094f31bd" ID-PMC="PMC5299438" ID-Pensoft-Pub="1313-2970-646-25" ID-PubMed="28228674" ID-ZBK="82CC88F888B049D490AF1F9D02B1B444" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-646-25" ModsDocOrigin="ZooKeys 646" ModsDocTitle="Towards a revision of the genus Periclimenes: resurrection of Ancylocaris Schenkel, 1902, and designation of three new genera (Crustacea, Decapoda, Palaemonidae)" checkinTime="1484692217103" checkinUser="pensoft" docAuthor="Ďuris, Zdenek & Horka, Ivona" docDate="2017" docId="48728D5A973BBC5195B3CC7B9B63F4A3" docLanguage="en" docName="ZooKeys 646: 25-44" docOrigin="ZooKeys 646" docSource="http://dx.doi.org/10.3897/zookeys.646.11397" docTitle="Actinimenes Ďuris & Horka, 2017, gen. n." docType="treatment" docUuid="A1D1A9D6-406C-4EB3-B750-494A81EEAF9A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="30" masterDocId="FFD4FFD1FFEBFF92FFBCFFD2FFC6FFE8" masterDocTitle="Towards a revision of the genus Periclimenes: resurrection of Ancylocaris Schenkel, 1902, and designation of three new genera (Crustacea, Decapoda, Palaemonidae)" masterLastPageNumber="44" masterPageNumber="25" pageNumber="28" updateTime="1668163901988" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Towards a revision of the genus Periclimenes: resurrection of Ancylocaris Schenkel, 1902, and designation of three new genera (Crustacea, Decapoda, Palaemonidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ďuris, Zdenek</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Horka, Ivona</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>646</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>25</mods:start>
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<mods:end>44</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.646.11397</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.646.11397</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-646-25</mods:identifier>
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<mods:identifier type="ZooBank">82CC88F888B049D490AF1F9D02B1B444</mods:identifier>
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<treatment ID-GBIF-Taxon="127900302" LSID="urn:lsid:zoobank.org:act:A1D1A9D6-406C-4EB3-B750-494A81EEAF9A" httpUri="http://treatment.plazi.org/id/48728D5A973BBC5195B3CC7B9B63F4A3" lastPageId="5" lastPageNumber="30" pageId="3" pageNumber="28">
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<subSubSection pageId="3" pageNumber="28" type="nomenclature">
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<paragraph pageId="3" pageNumber="28">
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<taxonomicName LSID="http://zoobank.org/A1D1A9D6-406C-4EB3-B750-494A81EEAF9A" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="28">gen. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="28" type="type species">
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<paragraph pageId="3" pageNumber="28">Type species.</paragraph>
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<paragraph pageId="3" pageNumber="28">
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Periclimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Periclimenes ornatus" order="Decapoda" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornatus">Periclimenes ornatus</taxonomicName>
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Bruce, 1969, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="28" type="included species">
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<paragraph pageId="3" pageNumber="28">Included species.</paragraph>
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<paragraph pageId="3" pageNumber="28">
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes inornatus" order="Decapoda" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="inornatus">Actinimenes inornatus</taxonomicName>
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(Kemp, 1922), comb. n. (Fig. 3C);
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes ornatus" order="Decapoda" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornatus">Actinimenes ornatus</taxonomicName>
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(Bruce, 1969), comb. n. (Fig. 3D); and
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes ornatellus" order="Decapoda" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornatellus">Actinimenes ornatellus</taxonomicName>
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(Bruce, 1979), comb. n.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28" type="diagnosis">
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<paragraph pageId="3" pageNumber="28">Diagnosis.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
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Carapace smooth; rostrum well developed, compressed, dorsal and ventral margins convex, with 7-10 dorsal teeth (posterior tooth behind orbits) and 0-2 ventral teeth, lateral carinae and orbit feebly developed, epigastric and supraorbital spines absent, inferior orbital angle usually produced, rounded, antennal tooth marginal, hepatic tooth close to level of latter. Pleon smooth, third segment not posteriorly produced, pleura rounded. Telson with two pairs of moderately large dorsal marginal spines situated on anterior and posterior thirds of telson length; three pairs of posterior spines, lateral spines smaller than dorsal spines. Eyes with globular cornea, small additional pigment spot dorsally on corneal margin. Antennule well developed. Antennal basicerite armed with lateral tooth; scaphocerite well developed, moderately broad, with distolateral tooth small, not reaching distal end of lamella. Mandible without palp, molar process robust, incisor process as usual for the family. Maxillula with bilobed palp, laciniae as usual for the family. Maxilla with simple palp, basal endite slender, deeply bilobed, coxal endite obsolete, scaphognathite moderately broad. First maxilliped with simple palp, basal endite fused with coxal endite, exopod with large caridean lobe,
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<normalizedToken originalValue="flagellum">flagellum</normalizedToken>
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slender with several plumose distal setae, epipod feebly bilobed. Second maxilliped with normal endopod, propodus not produced distomesially, exopod similar to
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<normalizedToken originalValue="first">first</normalizedToken>
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maxilliped, without accessory lobe, coxa with oval epipod without podobranch. Third maxilliped with slender endopod, ischiomerus fused to basis, exopod as in second maxilliped, coxa with oval lateral plate, arthrobranch rudimentary or lacking. Fourth thoracic sternite with broad transverse ridge subdivided by median incision. First pereiopods moderately slender, chela with
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<normalizedToken originalValue="fingers">fingers</normalizedToken>
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subequal to palm, deeply subspatulate with entire cutting edges, coxa with setose distoventral lobe.
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<pageBreakToken pageId="4" pageNumber="29" start="start">Second</pageBreakToken>
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pereiopods well developed, smooth, similar and equal,
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<normalizedToken originalValue="fingers">fingers</normalizedToken>
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with several small recurved teeth on proximal half, palm subcylindrical, longer than fingers, carpo-propodal articulation terminal, carpus much shorter than palm, merus unarmed, coxa without distoventral lobe. Ambulatory pereiopods moderately slender, propodus without ventral spines, dactyli with stout unarmed corpus, unguis elongate, curved. Endopod of male first pleopod simple, elliptic, feebly spinulose medioproximally, with several setulose setae distolaterally. Male second pleopod with appendix masculina slender, with several simple terminal and lateral setae. Uropods normal, exopod with small distolateral tooth and normal movable spine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="29" type="etymology">
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<paragraph pageId="4" pageNumber="29">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="29">
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From
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<taxonomicName lsidName="Palaemonoidea" order="Actiniaria" pageId="4" pageNumber="29" rank="order">Actiniaria</taxonomicName>
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, the order of
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<taxonomicName class="Anthozoa" lsidName="Palaemonoidea" pageId="4" pageNumber="29" rank="class">Anthozoa</taxonomicName>
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which comprises the host sea anemones for the genus, and
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Periclimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Periclimenes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Periclimenes</taxonomicName>
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to which genus the species previously belonged; gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="29" type="figures">
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<paragraph pageId="4" pageNumber="29">Figures</paragraph>
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<paragraph pageId="4" pageNumber="29">
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(selected).
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<bibRefCitation author="Kemp, S" journalOrPublisher="Records of the Indian Museum" pageId="14" pageNumber="39" pagination="113 - 288" title="Notes on CrustaceaDecapoda in the Indian Museum, XV. Pontoniinae." url="http://www.biodiversitylibrary.org/part/146043/summary" volume="24" year="1922">Kemp (1922</bibRefCitation>
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: figs 43-46),
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<bibRefCitation author="Bruce, AJ" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="12" pageNumber="37" pagination="89 - 153" title="A report on some pontoniid shrimps collected from the Seychelle Islands by the F. R. V. Manihine, 1972, with a review of the Seychelles pontoniid shrimp fauna." url="https://doi.org/10.1111/j.1096-3642.1976.tb01012.x" volume="59" year="1976">Bruce (1976</bibRefCitation>
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: 10-11;
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<bibRefCitation author="Bruce, AJ" journalOrPublisher="Crustaceana" pageId="12" pageNumber="37" pagination="209 - 230" title="A report on a small collection of pontoniine shrimps from Eniwetok Atoll." url="https://www.jstor.org/stable/25027504?seq=1page_scan_tab_contents" volume="5" year="1979">1979</bibRefCitation>
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: figs 3B, 4-7, Pl. 1: fig.
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<normalizedToken originalValue="B–E">B-E</normalizedToken>
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;
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<bibRefCitation author="Bruce, AJ" editor="Morton, BS" journalOrPublisher="Hong Kong University Press, Hong Kong" pageId="12" pageNumber="37" pagination="233 - 284" title="The pontoniine shrimp fauna of Hong Kong." url="http://www.reefbase.org/resource_center/publication/pub_20467.aspx" volumeTitle="Proceedings of the 1 st International Marine Biology Workshop: The marine flora and fauna of Hong Kong and southern China, Hong Kong, 1980" year="1982">1982</bibRefCitation>
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: figs 11-12),
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<bibRefCitation author="Fransen, CHJM" journalOrPublisher="Netherlands Journal of Sea Research" pageId="14" pageNumber="39" pagination="131 - 147" title="Notes on caridean shrimps collected during the Snellius-II expedition. I. Associates of Anthozoa." url="https://doi.org/10.1016/0077-7579(89)90008-2" volume="23" year="1989">Fransen (1989</bibRefCitation>
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: figs 2-3).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="30" pageId="4" pageNumber="29" type="systematic position">
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<paragraph pageId="4" pageNumber="29">Systematic position.</paragraph>
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<paragraph pageId="4" pageNumber="29">
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Based on recent molecular studies (
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<bibRefCitation author="Gan, ZB" journalOrPublisher="Journal of Zoological Systematics and Evolutionary Research" pageId="14" pageNumber="39" pagination="282 - 290" title="Phylogeny of Indo-West Pacific pontoniine shrimps (Crustacea: Decapoda: Caridea) based on multilocus analysis." url="https://doi.org/10.1111/jzs.12108" volume="53" year="2015">Gan et al. 2015</bibRefCitation>
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,
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<bibRefCitation author="Horka, I" journalOrPublisher="Scientific Reports" pageId="14" pageNumber="39" title="Multiple host switching events shape the evolution of symbiotic palaemonid shrimps (Crustacea: Decapoda)." url="https://doi.org/10.1038/srep26486" volume="6" year="2016">
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<normalizedToken originalValue="Horká">Horka</normalizedToken>
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et al. 2016
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</bibRefCitation>
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), species of the genus
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
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gen. n. show a close phylogenetic relationship to two groups of taxa,
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Zenopontonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zenopontonia" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Zenopontonia</taxonomicName>
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Bruce, 1975 and some other echinoderm-associated taxa on the one hand, and to the
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Periclimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Periclimenes diversipes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="diversipes">Periclimenes diversipes</taxonomicName>
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species group on the other hand. While the latter are also cnidarian associates, they are distinctly smaller species with more slender ambulatory pereiopods and dactyls, but mainly with very distinctive second pereiopods with both or at least one of the chelae of a specific subspatulate shape, with fingers generally longer than palm. Further, the species of the latter group share a similar shape of rostrum, and position of the carapacial teeth, subspatulate first pereiopod chela, short carpus of second pereiopods, simple ambulatory dactyli with elongate sharp unguis, and the shape and spinulation of the male pleopods (
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<bibRefCitation author="Bruce, AJ" journalOrPublisher="The Beagle, Records of the Northern Territory Museum of Arts and Sciences" pageId="12" pageNumber="37" pagination="149 - 156" title="Periclimenesgonioporae sp. nov. (Crustacea: Decapoda: Palaemonidae), a new coelenterate-associated shrimp." url="https://search.informit.com.au/documentSummary;dn=942059688801559;res=IELHSS" volume="6" year="1989">Bruce 1989</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="4" pageNumber="29">
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Zenopontonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zenopontonia" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Zenopontonia</taxonomicName>
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as well as other related echinoderm-associated taxa, such as
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Periclimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Periclimenes colemani" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="colemani">Periclimenes colemani</taxonomicName>
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Bruce, 1975, and
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Lipkemenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lipkemenes lanipes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="lanipes">Lipkemenes lanipes</taxonomicName>
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(Kemp, 1922), are generally also very similar to
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
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by the position of antennal and hepatic teeth, an incised transverse ridge on the fourth thoracic sternite, the deeply subspatulate chelae of the first pereiopod, the shape of the chela of the second pereiopod and the very short carpus, and by the shape and spinulation of the male pleopods (
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<bibRefCitation author="Bruce, AJ" journalOrPublisher="The Beagle, Records of the Northern Territory Museum of Arts and Sciences" pageId="12" pageNumber="37" pagination="149 - 156" title="Periclimenesgonioporae sp. nov. (Crustacea: Decapoda: Palaemonidae), a new coelenterate-associated shrimp." url="https://search.informit.com.au/documentSummary;dn=942059688801559;res=IELHSS" volume="6" year="1989">Bruce 1989</bibRefCitation>
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). These species however differ from
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
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spp. by a more down curved rostrum, more posteriorly situated and smaller dorsal telson spines, fine pectination on the fingers of the first pereiopod, and the ambulatory dactyli having small distoventral tooth on the corpus, sometimes also fully reduced (
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<bibRefCitation author="Marin, I" editor="Britayev, TA" journalOrPublisher="KMK Scientific Press, Moscow" pageId="14" pageNumber="39" pagination="345 - 405" title="New records and associations of pontoniine shrimps (Crustacea: Decapoda: Caridea: Palaemonidae: Pontoniinae) from the Nhatrang Bay, Vietnam; with taxonomic remarks on some species from the Indo-West Pacific region." volumeTitle="Benthic fauna of the Bay of Nhatrang, Southern Vietnam" year="2012">Marin 2012</bibRefCitation>
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). Some mollusc- or ascidian-associated genera, e.g.
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Anchistus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchistus" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Anchistus</taxonomicName>
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Borradaile, 1898,
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Paranchistus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paranchistus" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Paranchistus</taxonomicName>
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Holthuis, 1952, or
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Dasella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasella" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Dasella</taxonomicName>
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Lebour, 1945, show some phylogenetic relationship to these echinoderm associated taxa (
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<bibRefCitation author="Gan, ZB" journalOrPublisher="Journal of Zoological Systematics and Evolutionary Research" pageId="14" pageNumber="39" pagination="282 - 290" title="Phylogeny of Indo-West Pacific pontoniine shrimps (Crustacea: Decapoda: Caridea) based on multilocus analysis." url="https://doi.org/10.1111/jzs.12108" volume="53" year="2015">Gan et al. 2015</bibRefCitation>
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,
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<bibRefCitation author="Horka, I" journalOrPublisher="Scientific Reports" pageId="14" pageNumber="39" title="Multiple host switching events shape the evolution of symbiotic palaemonid shrimps (Crustacea: Decapoda)." url="https://doi.org/10.1038/srep26486" volume="6" year="2016">
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<normalizedToken originalValue="Horká">Horka</normalizedToken>
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||
et al. 2016
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</bibRefCitation>
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), and thus more remotely also to
|
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
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gen. n. All species of these genera differ however from
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
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gen. n. by specialized biunguiculate ambulatory dactyli.
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</paragraph>
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<paragraph lastPageId="5" lastPageNumber="30" pageId="4" pageNumber="29">
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The three species of
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
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gen. n. were previously thought to be part of the '
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<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Periclimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Periclimenes brevicarpalis" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="brevicarpalis">Periclimenes brevicarpalis</taxonomicName>
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<normalizedToken originalValue="group’">group'</normalizedToken>
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(
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<bibRefCitation author="Bruce, AJ" journalOrPublisher="Zoologische Verhandelingen" pageId="13" pageNumber="38" pagination="1 - 44" title="Observations upon some pontoniine shrimps from Aqaba, Jordan." url="http://www.repository.naturalis.nl/document/148926" volume="205" year="1983">Bruce and Svoboda 1983</bibRefCitation>
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,
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<bibRefCitation author="Bruce, AJ" journalOrPublisher="Zootaxa" pageId="13" pageNumber="38" pagination="45 - 54" title="Periclimenesnevillei sp. nov. (Crustacea: Decapoda: Pontoniinae) from Vanuatu." url="http://www.mapress.com/j/zt/article/view/8665" volume="2510" year="2010">Bruce 2010</bibRefCitation>
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), although
|
||
<bibRefCitation author="Fransen, CHJM" journalOrPublisher="Netherlands Journal of Sea Research" pageId="14" pageNumber="39" pagination="131 - 147" title="Notes on caridean shrimps collected during the Snellius-II expedition. I. Associates of Anthozoa." url="https://doi.org/10.1016/0077-7579(89)90008-2" volume="23" year="1989">
|
||
<pageBreakToken pageId="5" pageNumber="30" start="start">Fransen</pageBreakToken>
|
||
(1989)
|
||
</bibRefCitation>
|
||
regarded them as comprising a '
|
||
<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Periclimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Periclimenes inornatus" order="Decapoda" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="inornatus">Periclimenes inornatus</taxonomicName>
|
||
<normalizedToken originalValue="group’">group'</normalizedToken>
|
||
of identical composition of the new genus (see also Remarks for
|
||
<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Ancylocaris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancylocaris brevicarpalis" order="Decapoda" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="brevicarpalis">Ancylocaris brevicarpalis</taxonomicName>
|
||
comb. n., above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="30" type="distribution">
|
||
<paragraph pageId="5" pageNumber="30">Distribution.</paragraph>
|
||
<paragraph pageId="5" pageNumber="30">
|
||
Widely distributed in the Indo-West
|
||
<normalizedToken originalValue="Pacific">Pacific</normalizedToken>
|
||
from the Red Sea and Kenya to Japan, Marshall Islands, and Fiji.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="30" type="ecology">
|
||
<paragraph pageId="5" pageNumber="30">Ecology.</paragraph>
|
||
<paragraph pageId="5" pageNumber="30">
|
||
The species of the present genus are all obligate associates of sea anemones (
|
||
<taxonomicName lsidName="Palaemonoidea" pageId="5" pageNumber="30" phylum="Cnidaria" rank="phylum">Cnidaria</taxonomicName>
|
||
:
|
||
<taxonomicName lsidName="Palaemonoidea" order="Actiniaria" pageId="5" pageNumber="30" rank="order">Actiniaria</taxonomicName>
|
||
) (see
|
||
<bibRefCitation author="Fransen, CHJM" journalOrPublisher="Netherlands Journal of Sea Research" pageId="14" pageNumber="39" pagination="131 - 147" title="Notes on caridean shrimps collected during the Snellius-II expedition. I. Associates of Anthozoa." url="https://doi.org/10.1016/0077-7579(89)90008-2" volume="23" year="1989">Fransen 1989</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mueller, H-G" journalOrPublisher="H. - G. Mueller, Wetzlar" pageId="14" pageNumber="39" title="Catalogue of Indo-West Pacific pontoniine shrimps" url="http://www.vliz.be/imisdocs/publications/17544.pdf" year="1993">
|
||
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
|
||
1993
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="30" type="key">
|
||
<paragraph pageId="5" pageNumber="30">
|
||
Key to species identification of
|
||
<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes" order="Decapoda" pageId="5" pageNumber="30" phylum="Arthropoda" rank="genus">Actinimenes</taxonomicName>
|
||
gen. n.
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="30">
|
||
<table pageId="5" pageNumber="30">
|
||
<tr pageId="5" pageNumber="30">
|
||
<td colspan="1" pageId="5" pageNumber="30" rowspan="1">
|
||
<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes ornatus" order="Decapoda" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="ornatus">Actinimenes ornatus</taxonomicName>
|
||
(Bruce, 1969), comb. n.
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="30">
|
||
<td colspan="1" pageId="5" pageNumber="30" rowspan="1">2</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="30">
|
||
<td colspan="1" pageId="5" pageNumber="30" rowspan="1">
|
||
<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes inornatus" order="Decapoda" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="inornatus">Actinimenes inornatus</taxonomicName>
|
||
(Kemp, 1922), comb. n.
|
||
</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="30">
|
||
<td colspan="1" pageId="5" pageNumber="30" rowspan="1">
|
||
<taxonomicName class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinimenes ornatellus" order="Decapoda" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="ornatellus">Actinimenes ornatellus</taxonomicName>
|
||
(Bruce, 1979), comb. n.
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |