156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="38BFE9A6A716AB6B7F4D79E68A85D4D3" ID-CLB-Dataset="28501" ID-DOI="http://dx.doi.org/10.3897/zookeys.826.26488" ID-GBIF-Dataset="9d62af83-e4f5-423b-8e12-7dd567f69921" ID-PMC="PMC6405737" ID-Pensoft-Pub="1313-2970-826-1" ID-PubMed="30858752" ID-ZooBank="B1550A3AE54744509A44BC4366D5E110" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-826-1" ModsDocOrigin="ZooKeys 826" ModsDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" checkinTime="1551446249571" checkinUser="pensoft" docAuthor="Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z." docDate="2019" docId="8E33AE68153C0B4E327366672B4333A0" docLanguage="en" docName="ZooKeys 826: 1-343" docOrigin="ZooKeys 826" docSource="http://dx.doi.org/10.3897/zookeys.826.26488" docTitle="Canthon (Glaphyrocanthon) ohausi Balthasar 1939, stat. n." docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="FFECFFBAFFFC2D25D00B9843FFD67749" masterDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" masterLastPageNumber="343" masterPageNumber="1" pageNumber="48" updateTime="1701379963039" updateUser="ExternalLinkService">
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<mods:title id="B0653886C1D3D56EF5A8F51D83331379">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="69544D916814EB036EAFE8F1CE82FDFB">Chamorro, William</mods:namePart>
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<mods:namePart id="B5B65EF1AD9FC48E4F6D33312E1024F4">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="902F60173CA203C79D8D5C8CCBACB624">senjo, Angelico</mods:namePart>
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<mods:namePart id="3E0B2B428AFE7B3462CD6F62BB1977CB">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="B0C2AEBF4463351C882EFE2E9A1B5E7E">text</mods:typeOfResource>
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<mods:date id="1EAA024FF9C5EFC2A15907547EF2E12E">2019</mods:date>
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<mods:detail id="710892DF33BD5747595CE32E4705A642" type="volume">
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<mods:number id="A3ADC6600EC4A8DD39EE3EE72C0DDDED">826</mods:number>
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<mods:classification id="76851DCE41A0108BF56D17A2D93FD75D">journal article</mods:classification>
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<mods:identifier id="F28C639B038702488AE1EBEFD76BC29B" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
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<treatment id="8E33AE68153C0B4E327366672B4333A0" ID-GBIF-Taxon="154127674" LSID="urn:lsid:plazi:treatment:8E33AE68153C0B4E327366672B4333A0" httpUri="http://treatment.plazi.org/id/8E33AE68153C0B4E327366672B4333A0" lastPageId="50" lastPageNumber="50" pageId="47" pageNumber="48">
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<subSubSection id="463037E963A771B2F42AD48E48479997" pageId="47" pageNumber="48" type="nomenclature">
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<paragraph id="FC5F95093146EC021C4E6E51D2BE7755" pageId="47" pageNumber="48">
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<taxonomicName id="4A09A1C1F8CC382B876639C05F3DF401" ID-CoL="QP9Q" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon (Glaphyrocanthon) ohausi" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="ohausi" subGenus="Glaphyrocanthon">Canthon (Glaphyrocanthon) ohausi Balthasar, 1939</taxonomicName>
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<taxonomicNameLabel id="433F2957A25973B4B29880525479F9D2" pageId="47" pageNumber="48">stat. n.</taxonomicNameLabel>
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Plates 13A, 56D
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</paragraph>
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</subSubSection>
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<subSubSection id="72BE9A42C86AAF5E598F7A04FB2D908F" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="reference_group">
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<paragraph id="229B55572261DF473A8DA63A8A5D162B" pageId="48" pageNumber="49">
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<taxonomicName id="2A264735FB935B49256301B49B4DE21E" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon angustatus" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="angustatus" subSpecies="ohausi">
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<pageBreakToken id="E5AA0B86A42CDD321370C65210659ACA" pageId="48" pageNumber="49" start="start">Canthon</pageBreakToken>
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angustatus ohausi
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</taxonomicName>
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Balthasar, 1939d: 216 (original description. Type locality: Ecuador).
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</paragraph>
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<paragraph id="4082761497418B9CD99E06054551FC4D" pageId="48" pageNumber="49">
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<taxonomicName id="03B513064B47BB2C1E617134B3525A3B" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon angustatus" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="angustatus" subSpecies="ohausi">Canthon angustatus ohausi</taxonomicName>
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:
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<bibRefCitation id="39A7CA621CD8A11DAEFEC4429464D631" author="Bezdek, A" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="262" pageNumber="263" pagination="349 - 378" title="Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic." volume="51" year="2011">Bezdek and Hajek 2011</bibRefCitation>
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: 361 (catalog of types NMPC);
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<bibRefCitation id="6B897E7E6AB188229A5881428F40E5F2" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 92 (cited for Ecuador).
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</paragraph>
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<paragraph id="B199A1D4B3CB5B50AC0CD9D9F5583643" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
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<taxonomicName id="47FADBF96CA9A31E28A919A1FAB90324" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon ohausi" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ohausi">Canthon ohausi</taxonomicName>
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<pageBreakToken id="5DD7C92FAE07D7CC1153FE044D639753" pageId="49" pageNumber="50" start="start">:</pageBreakToken>
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<bibRefCitation id="893AA4AEACD44B137A13BD27EFA33BCA" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 200 (list of species for Latin America);
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<bibRefCitation id="E21F6814ABB471C50FECF3EA7DB5B4C9" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
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: 314-315 (cited for Ecuador).
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</paragraph>
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</subSubSection>
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<subSubSection id="4E1A59EF67D944405062B5CD7EE29303" pageId="49" pageNumber="50" type="type specimens">
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<paragraph id="273F2D9B6670B15410CCD38782C816B5" pageId="49" pageNumber="50">Type specimens.</paragraph>
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<paragraph id="F6BB459293992CC5CA7332F8A072535D" pageId="49" pageNumber="50">
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<taxonomicName id="39BA7A40BCE41575677E46102F89EDA7" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon angustatus" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="angustatus" subSpecies="ohausi">Canthon angustatus ohausi</taxonomicName>
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Balthasar, 1939. The lectotype (♂) (here designated) and one paralectotype are deposited at the NMPC. Locality: Mera, examined.
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</paragraph>
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<paragraph id="39E2FE784ECFC58E69BE65FE712EF202" pageId="49" pageNumber="50">Lectotype (here designated) (♂): "Mera / Ecuador [p]", "Typus [p, red label, black margin]", "C. angustatus / ssp. m. / Typus! [p and hw] / Dr. V. Balthasar det. [p]", "LECTOTYPE ♂ / Canthon angus- / tatus ohausi / Balth. / des. F.Z.Vaz-de-Mello, 2013 [hw and p, red label, black margin]".</paragraph>
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<paragraph id="661737287FD65B165B0CE74251F2AD4C" pageId="49" pageNumber="50">Paralectotype (♀): "ECUADOR / Sabanilla / F. OhausS. [p]", "Senckenberg / Museum [p]", "Typus [p, red label, black margin]", "C. angustatus / ssp. Ohausi m. / Typus! [hw] / Dr. V. Balthasar det. [p]", "ssp. Ohausi / m. [hw, green label, black margin]", "PARALECTOTYPE / Canthon angusta- / tus ohausi ♀ / Balth. / des. F.Z.Vaz-de-Mello, 2013 [hw and p, yellow label, black margin]".</paragraph>
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</subSubSection>
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<subSubSection id="AC766523287B920B38286F4AE2BEA7C0" pageId="49" pageNumber="50" type="distribution">
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<paragraph id="2CECE8A05D6EE58FFB46C7937912D6F2" pageId="49" pageNumber="50">Distribution.</paragraph>
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<paragraph id="AE66467061BD9AB8ACD22F253748B846" pageId="49" pageNumber="50">Brazil, Colombia, Ecuador, Peru, and French Guiana.</paragraph>
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</subSubSection>
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<subSubSection id="79B477DC2293AD1190D5E90B39778BAB" pageId="49" pageNumber="50" type="records examined">
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<paragraph id="E0B130F7507B24D8B72A9BBEFE67FA92" pageId="49" pageNumber="50">Records examined.</paragraph>
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<paragraph id="2ECAA267A34AABE64B92A68C399EA010" pageId="49" pageNumber="50">
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NAPO:
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<normalizedToken id="3F076C7DBBEE4E43673EE9707B9EF41B" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="453269F182902CA9EC3BBD5845C8647B" originalValue="Nushiño">Nushino</normalizedToken>
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Gareno-Waponi, 370 m (1 specimen MUTPL). ORELLANA: Campo Palanda-Yuca Sur,
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<normalizedToken id="580F814D5015A6546C77542BA7EB335A" originalValue="Estación">Estacion</normalizedToken>
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Palanda 5, 320 m (1 specimen MUTPL);
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<normalizedToken id="44497E662BE72CBF0FC6619ECDC39E1A" originalValue="Estación">Estacion</normalizedToken>
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de Biodiversidad Tiputini, 215 m, Parque Nacional
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<normalizedToken id="EBBF674710A9170A2C0CBC16152989E7" originalValue="Yasuní">Yasuni</normalizedToken>
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(3 specimens MUTPL); Ines Arango road
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<normalizedToken id="8D39E09CCA45548FA280FAB3904D2F8C" originalValue="Tiwino-río">Tiwino-rio</normalizedToken>
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Shiripuno, 250 m (1 specimen MUTPL); road to Maxus km 45
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<normalizedToken id="9E6BDEA9B78E3F3545B9FA5BCA01A97B" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="7555A7A07C7DE2782594EEB07921590F" originalValue="Capirón">Capiron</normalizedToken>
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, 220 m, Parque Nacional
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<normalizedToken id="3DED6C79F1F84C1BDDEEE3FE100457FD" originalValue="Yasuní">Yasuni</normalizedToken>
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Tiputini (1 specimen MUTPL); Lago San Pedro, plataforma Copal, 310 m (1 specimen MUTPL). PASTAZA: Bosque Protector
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<normalizedToken id="F924FD38E036BB98452FD71C55066AF8" originalValue="Oglán">Oglan</normalizedToken>
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Alto, 945 m (2 specimens MUTPL; 1 specimen MGO-UC); Chuyayacu, Oleoducto km 25, 200 m (1 specimen MGO-UC); Mera (2 specimens NMPC); Pandanuque, 420 m (1 specimen MUTPL).
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<normalizedToken id="823E5DAB50ACC53F17ABC84371282A8F" originalValue="SUCUMBÍOS">SUCUMBIOS</normalizedToken>
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: Aucayacu
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<normalizedToken id="D7793F93849EBE0D7CD96DD621C05739" originalValue="Río">Rio</normalizedToken>
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El Eno, 16 km de Lago Agrio, 275 m (3 specimens CEMT; 36 specimens MGO-UC); Cascales Pozo Aguas Blancas, 385 m (2 specimens MUTPL); Cascales road to Lumbaqui Pozo Mascarey, 395 m (1 specimen MUTPL); Limoncocha, Reserva
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<normalizedToken id="2311A74EDD34053083596C1EFA408D98" originalValue="Biológica">Biologica</normalizedToken>
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(1 specimen MUTPL); Pacayacu Campo Libertador, Tapi, 265 m (1 specimen MUTPL). ZAMORA CHINCHIPE: Sabanilla [= El Tambo, ZAMORA CHINCHIPE] (1 specimen NMPC); Tundayme campamento Mirador, Enerentsa, 1030 m (1 specimen MUTPL).
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</paragraph>
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</subSubSection>
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<subSubSection id="FFD47B9F158A7650428DD1E57B7A273D" pageId="49" pageNumber="50" type="temporal data">
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<paragraph id="3A3632902485D3BC4013FFAE5C65F4BD" pageId="49" pageNumber="50">Temporal data.</paragraph>
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<paragraph id="2968D5CB61E4FE6BD51EC076181B9085" pageId="49" pageNumber="50">Collected in January, February, March, May, June, July, August, September, November, and December.</paragraph>
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</subSubSection>
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<subSubSection id="75C7EE17E3D9FCF17104F2ADC60444D3" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="remarks">
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<paragraph id="F6BA348060E6ABABA3A7D0C69B9FA754" pageId="49" pageNumber="50">Remarks.</paragraph>
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<paragraph id="9C2E21D2043742DC8D7E1351E6090187" pageId="49" pageNumber="50">Inhabits the lowland evergreen forests and the foothill evergreen forests of the Amazon region from 215-1300 m a.s.l. Collected with pitfall traps baited with carrion and human feces.</paragraph>
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<paragraph id="EC659C924789EE8C9A6B0C09A78DCD4C" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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Balthasar (1939) described
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<taxonomicName id="D3174A5E5FAC4D13E467DED40652BAAC" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon angustatus" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subSpecies" species="angustatus" subSpecies="ohausi">Canthon angustatus ohausi</taxonomicName>
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(type locality: Ecuador) as a variety of
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<taxonomicName id="6F9C1A20F0DD3CD2C3201D37AF8448E1" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon angustatus" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="angustatus">Canthon angustatus</taxonomicName>
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Harold, 1867 (type locality: Costa Rica), which made the former name available as a subspecific category (see
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<bibRefCitation id="2BF6E17CDAA2C211E6D86D8E7D56BCDA" author="Celi, J" journalOrPublisher="Madrid, Espana" pageId="265" pageNumber="266" title="Codigo Internacional de Nomenclatura Zoologica. Cuarta edicion adoptada por la Union Internacional de Ciencias Biologicas." year="1999">ICZN 1999</bibRefCitation>
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, Articles 45.6.4 and 46.6.4.1). However, upon examining the external and genital morphology of the type specimens of
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<taxonomicName id="FF2C29B0324A321D2A6CCCA812C309C2" lsidName="C. angustatus" pageId="49" pageNumber="50" rank="subSpecies" species="angustatus" subSpecies="ohausi">C. angustatus ohausi</taxonomicName>
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(lectotype ♂ here designated, deposited at the NMPC, Plate 56D) and
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<taxonomicName id="33CE7417E6D5318CD2A3A5D42CA5ABF2" lsidName="C. angustatus" pageId="49" pageNumber="50" rank="species" species="angustatus">C. angustatus</taxonomicName>
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(lectotype ♂ here designated, deposited at the MNHN, Plate 56C), specifically, the differences in the aedeagus and the shapes of the pronotal spots, we think that they belong to two distinct species. Therefore, maintain
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<pageBreakToken id="C9BDA794650EA0FEE5E1FC67CE33233C" pageId="50" pageNumber="51" start="start">ing</pageBreakToken>
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the subspecific name originally proposed by Balthasar, we elevate it to species level under the following new status:
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<taxonomicName id="70F82851A0A6F79AEA746884F46E73A2" class="Insecta" family="Scarabaeidae" genus="Canthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthon (Glaphyrocanthon) ohausi" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="ohausi" subGenus="Glaphyrocanthon">Canthon (Glaphyrocanthon) ohausi</taxonomicName>
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Balthasar, 1939 stat. n. The lectotype is here designated and illustrated (♂ Plate 56D) and originates from Mera, Ecuador (associated with Amazon forests).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |