253 lines
50 KiB
XML
253 lines
50 KiB
XML
<document id="FCD209F3166B62ADB81A7E55B2A5B6DB" ID-CLB-Dataset="29502" ID-DOI="10.11646/zootaxa.4484.1.1" ID-GBIF-Dataset="74b777c9-eae0-4770-8c86-dcbb10fb06b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437848" ID-ZooBank="D66524CF-9C6D-4DF4-8CA2-B2C9708CF5FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538125412450" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P. & Melnik, Viacheslav P." docDate="2018" docId="521587E4561E552A09EEFC16884FFC23" docLanguage="en" docName="zootaxa.4484.1.1.pdf" docOrigin="Zootaxa 4484 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Calyssopora volcano Grischenko & Gordon & Melnik 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="AE2CFF9C563255070979FF888A4AFF87" masterDocTitle="Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining" masterLastPageNumber="91" masterPageNumber="1" pageNumber="45" updateTime="1698557051871" updateUser="ExternalLinkService">
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<mods:title id="FCF4CB5A979522F6155F2B264A1E099D">Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining</mods:title>
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<mods:name id="F75013B54BFA258742DF11B85F2E3398" type="personal">
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<mods:roleTerm id="B435EC75A3E48E642F33515874CDCE98">Author</mods:roleTerm>
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<mods:namePart id="3026A3F95C69625D9AB93FFAADA8058F">Grischenko, Andrei V.</mods:namePart>
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<mods:namePart id="8F3B426B70831D6D6FA83180EF03A304">Gordon, Dennis P.</mods:namePart>
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<mods:name id="9C20712D2A2B7C12620114F88F6EA704" type="personal">
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<mods:namePart id="E46CB9CD34B106D2C24ED0FD986B3DF2">Melnik, Viacheslav P.</mods:namePart>
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<mods:title id="BC1BE3FFED1F5CA22C94CF5B8F05F2A5">Zootaxa</mods:title>
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<mods:part id="2228A7D861D6B36885B70AF909FA0CEA">
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<mods:date id="92FB604199A5567C3DCB3621543EA416">2018</mods:date>
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<mods:number id="9D42A2C508D3F350C92CAE7117F425DA">2018-09-25</mods:number>
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<mods:number id="06DBFCA659668D7F72657D5A2C36EB59">4484</mods:number>
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<mods:identifier id="291B3BAEF2EADE3A8D6AC2AE28A07F7F" type="DOI">10.11646/zootaxa.4484.1.1</mods:identifier>
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<treatment id="521587E4561E552A09EEFC16884FFC23" ID-DOI="http://doi.org/10.5281/zenodo.5989819" ID-GBIF-Taxon="148403738" ID-Zenodo-Dep="5989819" LSID="urn:lsid:plazi:treatment:521587E4561E552A09EEFC16884FFC23" httpUri="http://treatment.plazi.org/id/521587E4561E552A09EEFC16884FFC23" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
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<subSubSection id="92A66579561E552B09EEFC168B16FC5D" pageId="44" pageNumber="45" type="nomenclature">
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<paragraph id="DA0336F2561E552B09EEFC168B90FC3F" blockId="44.[151,474,926,986]" box="[151,474,926,952]" pageId="44" pageNumber="45">
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<heading id="814B819E561E552B09EEFC168B90FC3F" bold="true" box="[151,474,926,952]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
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<emphasis id="E8C8EAE0561E552B09EEFC168B90FC3F" bold="true" box="[151,474,926,952]" pageId="44" pageNumber="45">
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<taxonomicName id="1DBC4D71561E552B09EEFC168BDAFC3F" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[151,400,926,952]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="44" pageNumber="45" phylum="Bryozoa" rank="species" species="volcano" status="sp. nov.">
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<emphasis id="E8C8EAE0561E552B09EEFC168BDAFC3F" bold="true" box="[151,400,926,952]" italics="true" pageId="44" pageNumber="45">Calyssopora volcano</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F3FB579B561E552B08E1FC168B90FC3F" box="[408,474,926,952]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="DA0336F2561E552B09EEFC498B16FC5D" blockId="44.[151,474,926,986]" box="[151,348,961,986]" pageId="44" pageNumber="45">
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(
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<figureCitation id="42872A77561E552B09E6FC498B4BFC5D" box="[159,257,961,986]" captionStart="FIGURE 2" captionStartId="10.[151,250,1774,1796]" captionTargetBox="[151,1436,392,1715]" captionTargetId="figure@10.[151,1436,392,1715]" captionTargetPageId="10" captionText="FIGURE 2. Colonies of some cyclostome bryozoans, in vivo, attached to polymetallic nodules. A–E, Pandanipora helix n. gen., n. sp.: A, specimen GLD4–09, Stn 190; B, specimen GLD4–12, Stn 262; C, specimen YMG4–07, Stn 143; D, specimen YMG4–13, Stn 295; E, specimen GLD 4–11, Stn 212. F, Tubuliporina sp. indet., specimen YMG18–01, Stn 7. G, H, Abyssoecia elevata n. gen., n. sp.: G, specimen GLD4–09, Stn 196; H, specimen GLD4–09, Stn 191. I, Discantenna metallica n. sp.: specimen GLD4–11, Stn 224. J, K, Frontohornera frontalis n. gen., n. sp.: J, specimen YMG4–07, Stn 124; K, specimen GLD4–11, Stn 210. L, Alyonushka hystricosa n. gen., n. sp.: specimen GLD4–09, Stn 199. M, Calyssopora volcano n. gen., n. sp.: specimen YMG18–01, Stn 33. N, O, Anyuta anastema n. gen., n. sp.: N, specimen GLD4–09, Stn 180; O, specimen YMG4–06, Stn 71. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437852/files/figure.png" pageId="44" pageNumber="45">Figs 2M</figureCitation>
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,
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<figureCitation id="42872A77561E552B0875FC4A8B19FC5D" box="[268,339,962,986]" captionStart-0="FIGURE 28" captionStart-1="FIGURE 29" captionStart-2="FIGURE 30" captionStartId-0="46.[151,250,1932,1954]" captionStartId-1="47.[151,250,1837,1859]" captionStartId-2="48.[151,250,1846,1868]" captionTargetBox-0="[174,1418,193,1917]" captionTargetBox-1="[194,1399,193,1821]" captionTargetBox-2="[183,1407,185,1834]" captionTargetId-0="figure@46.[174,1418,193,1917]" captionTargetId-1="figure@47.[194,1399,193,1821]" captionTargetId-2="figure@48.[183,1407,185,1834]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionTargetPageId-2="48" captionText-0="FIGURE 28. Calyssopora volcano n. gen., n. sp. A, B, holotype, ZIRAS 1/50714, with a single gonozooid; C, D, paratype 1, ZIRAS 2/50715, with two ooeciostomes and therefore two gonozooids presumed; E, F, paratype 2, ZIRAS 3/50716, with possibly newly completed gonozooid, having large alveolate spaces at left-hand periphery. Scale bars: 250 µm." captionText-1="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." captionText-2="FIGURE 30. Calyssopora volcano n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. A, B, specimen YMG4–13, Stn 291; C, D, specimen YMG4–07, Stn 139; E, F, specimen YMG4–14, Stn 358; G, H, specimen YMG4–04, Stn 57; I, J, specimen YMG4–13, Stn 283; K, L, specimen YMG4–13, Stn 300; M, N, specimen GLD4–09, Stn 198; O, P, specimen YMG4–13, Stn 274. A, B, four-zooid stage; C–F, four–five-zooid stages; G–J, five–six-zooid stages; K–P, later submature stages with calyciform colony center adequately structured with trabeculae and alveoli to support gonozooid development. Scale bars: 200 µm." httpUri-0="https://zenodo.org/record/1437905/files/figure.png" httpUri-1="https://zenodo.org/record/1437907/files/figure.png" httpUri-2="https://zenodo.org/record/1437909/files/figure.png" pageId="44" pageNumber="45">28–30</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="92A66579561E552B09EEFB8189B3FA99" pageId="44" pageNumber="45" type="materials_examined">
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<paragraph id="DA0336F2561E552B09EEFB8189B3FA99" blockId="44.[151,1437,1033,2030]" pageId="44" pageNumber="45">
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<emphasis id="E8C8EAE0561E552B09EEFB818BCBFBA5" bold="true" box="[151,385,1033,1058]" pageId="44" pageNumber="45">Material examined.</emphasis>
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<emphasis id="E8C8EAE0561E552B08F1FB818BA4FBA5" box="[392,494,1033,1058]" italics="true" pageId="44" pageNumber="45">Holotype</emphasis>
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: ZIRAS 1/50714, colony attached to nodule particle, YMG R.V.
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<emphasis id="E8C8EAE0561E552B0DB4FB818FD3FBA5" box="[1229,1433,1033,1058]" italics="true" pageId="44" pageNumber="45">Yuzhmorgeologiya</emphasis>
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cruise YMG4–14, Stn 357,
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<date id="AE021032561E552B08A8FBA688DAFBC1" box="[465,656,1069,1094]" pageId="44" pageNumber="45" value="2016-01-18">18 January 2016</date>
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, 14.03237° N, 131.74563° W,
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<quantity id="1D449B17561E552B0A97FBA58E0DFBC2" box="[1006,1095,1069,1094]" metricMagnitude="3" metricUnit="m" metricValue="5.073" pageId="44" pageNumber="45" unit="m" value="5073.0">5073 m</quantity>
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.
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<emphasis id="E8C8EAE0561E552B0D2AFBA68EF0FBC2" box="[1107,1210,1070,1093]" italics="true" pageId="44" pageNumber="45">Paratype</emphasis>
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1: ZIRAS 2/50715, colony attached to nodule particle, YMG R.V.
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<emphasis id="E8C8EAE0561E552B0BD3FBD98962FBED" box="[682,808,1105,1130]" italics="true" pageId="44" pageNumber="45">Gelendzhik</emphasis>
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cruise GLD4–12, Stn 251,
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<date id="AE021032561E552B0D1AFBD98EB2FBED" box="[1123,1272,1105,1130]" pageId="44" pageNumber="45" value="2013-04-06">6 April 2013</date>
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, 12.80823° N, 132.69682° W,
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<quantity id="1D449B17561E552B0824FBFE8B8BFB0A" box="[349,449,1142,1166]" metricMagnitude="3" metricUnit="m" metricValue="4.837" pageId="44" pageNumber="45" unit="m" value="4837.0">4837 m</quantity>
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.
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<emphasis id="E8C8EAE0561E552B08AEFBFE8874FB0A" box="[471,574,1142,1165]" italics="true" pageId="44" pageNumber="45">Paratype</emphasis>
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2: ZIRAS 3/50716, colony attached to nodule particle, YMG R.V.
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<emphasis id="E8C8EAE0561E552B09EEFB118B2FFB35" box="[151,357,1177,1202]" italics="true" pageId="44" pageNumber="45">Yuzhmorgeologiya</emphasis>
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cruise YMG4–14, Stn 325,
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<date id="AE021032561E552B0BB5FB1289FFFB35" box="[716,949,1177,1202]" pageId="44" pageNumber="45" value="2015-12-20">20 December 2015</date>
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, 13.58832° N, 131.68838° W,
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<quantity id="1D449B17561E552B0C41FB118FD2FB36" box="[1336,1432,1177,1202]" metricMagnitude="3" metricUnit="m" metricValue="5.031" pageId="44" pageNumber="45" unit="m" value="5031.0">5031 m</quantity>
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.
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<emphasis id="E8C8EAE0561E552B09EEFB358B3CFB51" box="[151,374,1213,1238]" italics="true" pageId="44" pageNumber="45">Additional material</emphasis>
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: YMG18–01, Stns 7, 15, 24, 33; YMG4–04, Stn 57; YMG4–06, Stns 70, 92, 99, 101, 103; YMG4–07, Stn 139; GLD4–09, Stns 173, 178, 185, 198; GLD4–11, Stn 216; GLD4–12, Stn 271; YMG4–13, Stns 274, 283, 291, 300; YMG4–14, Stns 348, 358. Total specimens examined 27.
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</paragraph>
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</subSubSection>
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<subSubSection id="92A66579561E552B09BEFAA188DDFAE1" pageId="44" pageNumber="45" type="etymology">
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<paragraph id="DA0336F2561E552B09BEFAA188DDFAE1" blockId="44.[151,1437,1033,2030]" pageId="44" pageNumber="45">
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<emphasis id="E8C8EAE0561E552B09BEFAA18B04FAC5" bold="true" box="[199,334,1321,1346]" pageId="44" pageNumber="45">Etymology.</emphasis>
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Anglicized spelling of the Italian
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<emphasis id="E8C8EAE0561E552B0BABFAA18961FAC5" box="[722,811,1321,1346]" italics="true" pageId="44" pageNumber="45">vulcano</emphasis>
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, alluding to the volcano-like gonozooid with its craterlike ooeciopore; used as a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="92A66579561E552A09BEFAF98BE2FE73" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="description">
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<paragraph id="DA0336F2561E552B09BEFAF98914F949" blockId="44.[151,1437,1033,2030]" pageId="44" pageNumber="45">
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<emphasis id="E8C8EAE0561E552B09BEFAF98B13FA0D" bold="true" box="[199,345,1393,1418]" pageId="44" pageNumber="45">Description.</emphasis>
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Colony erect, thickly pedunculate, with autozooidal peristomes radiating outwards around periphery of capitulum (
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<figureCitation id="42872A77561E552B08D4FA1D8849FA29" box="[429,515,1429,1454]" captionStart="FIGURE 28" captionStartId="46.[151,250,1932,1954]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure@46.[174,1418,193,1917]" captionTargetPageId="46" captionText="FIGURE 28. Calyssopora volcano n. gen., n. sp. A, B, holotype, ZIRAS 1/50714, with a single gonozooid; C, D, paratype 1, ZIRAS 2/50715, with two ooeciostomes and therefore two gonozooids presumed; E, F, paratype 2, ZIRAS 3/50716, with possibly newly completed gonozooid, having large alveolate spaces at left-hand periphery. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437905/files/figure.png" pageId="44" pageNumber="45">Fig. 28</figureCitation>
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), up to maximum of 22 peristomes in single irregular whorl. Skeletal surface of column with weakly defined ridges and furrows, granular-tubercular (
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<figureCitation id="42872A77561E552B0ABEFA318E1FFA55" box="[967,1109,1465,1490]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29A–C</figureCitation>
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), with sparse pores; surface ultrastructure with irregularly arranged crystallites lacking obvious preferred growth direction (
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<figureCitation id="42872A77561E552B0C67FA558FDAFA71" box="[1310,1424,1501,1526]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29H</figureCitation>
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). Capitulum of submature colony infundibuliform (
|
||
<figureCitation id="42872A77561E552B0BB0F989897AF99D" box="[713,816,1537,1562]" captionStart="FIGURE 30" captionStartId="48.[151,250,1846,1868]" captionTargetBox="[183,1407,185,1834]" captionTargetId="figure@48.[183,1407,185,1834]" captionTargetPageId="48" captionText="FIGURE 30. Calyssopora volcano n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. A, B, specimen YMG4–13, Stn 291; C, D, specimen YMG4–07, Stn 139; E, F, specimen YMG4–14, Stn 358; G, H, specimen YMG4–04, Stn 57; I, J, specimen YMG4–13, Stn 283; K, L, specimen YMG4–13, Stn 300; M, N, specimen GLD4–09, Stn 198; O, P, specimen YMG4–13, Stn 274. A, B, four-zooid stage; C–F, four–five-zooid stages; G–J, five–six-zooid stages; K–P, later submature stages with calyciform colony center adequately structured with trabeculae and alveoli to support gonozooid development. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437909/files/figure.png" pageId="44" pageNumber="45">Fig. 30L</figureCitation>
|
||
), bottom of funnel comprising deep alveolar chambers delimited by trabeculae (
|
||
<figureCitation id="42872A77561E552B08C6F9AD8864F9B9" box="[447,558,1573,1598]" captionStart="FIGURE 30" captionStartId="48.[151,250,1846,1868]" captionTargetBox="[183,1407,185,1834]" captionTargetId="figure@48.[183,1407,185,1834]" captionTargetPageId="48" captionText="FIGURE 30. Calyssopora volcano n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. A, B, specimen YMG4–13, Stn 291; C, D, specimen YMG4–07, Stn 139; E, F, specimen YMG4–14, Stn 358; G, H, specimen YMG4–04, Stn 57; I, J, specimen YMG4–13, Stn 283; K, L, specimen YMG4–13, Stn 300; M, N, specimen GLD4–09, Stn 198; O, P, specimen YMG4–13, Stn 274. A, B, four-zooid stage; C–F, four–five-zooid stages; G–J, five–six-zooid stages; K–P, later submature stages with calyciform colony center adequately structured with trabeculae and alveoli to support gonozooid development. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437909/files/figure.png" pageId="44" pageNumber="45">Fig. 30O</figureCitation>
|
||
). Additional thin walls cross gaps between adjacent peristomes at colony periphery where additional autozooidal chambers develop to fill vacancies in whorl as colony expands (
|
||
<figureCitation id="42872A77561E552B0C4DF9C18FD2F9E5" box="[1332,1432,1609,1634]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Figs 29J</figureCitation>
|
||
,
|
||
<figureCitation id="42872A77561E552B09EEF9E68B40F901" box="[151,266,1646,1670]" captionStart="FIGURE 30" captionStartId="48.[151,250,1846,1868]" captionTargetBox="[183,1407,185,1834]" captionTargetId="figure@48.[183,1407,185,1834]" captionTargetPageId="48" captionText="FIGURE 30. Calyssopora volcano n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. A, B, specimen YMG4–13, Stn 291; C, D, specimen YMG4–07, Stn 139; E, F, specimen YMG4–14, Stn 358; G, H, specimen YMG4–04, Stn 57; I, J, specimen YMG4–13, Stn 283; K, L, specimen YMG4–13, Stn 300; M, N, specimen GLD4–09, Stn 198; O, P, specimen YMG4–13, Stn 274. A, B, four-zooid stage; C–F, four–five-zooid stages; G–J, five–six-zooid stages; K–P, later submature stages with calyciform colony center adequately structured with trabeculae and alveoli to support gonozooid development. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437909/files/figure.png" pageId="44" pageNumber="45">30A, I, O</figureCitation>
|
||
); some of these openings seem to represent adjacent extrazooidal spaces, their openings narrowed in mature colony (
|
||
<figureCitation id="42872A77561E552B0830F9198BFAF92D" box="[329,432,1681,1706]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29B</figureCitation>
|
||
). Increasing calcification reduces openings in calyx center to expanding porous platform that serves as irregular open ‘floor’ to gonozooid(s) (
|
||
<figureCitation id="42872A77561E552B0B90F93D8918F949" box="[745,850,1717,1742]" captionStart="FIGURE 30" captionStartId="48.[151,250,1846,1868]" captionTargetBox="[183,1407,185,1834]" captionTargetId="figure@48.[183,1407,185,1834]" captionTargetPageId="48" captionText="FIGURE 30. Calyssopora volcano n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. A, B, specimen YMG4–13, Stn 291; C, D, specimen YMG4–07, Stn 139; E, F, specimen YMG4–14, Stn 358; G, H, specimen YMG4–04, Stn 57; I, J, specimen YMG4–13, Stn 283; K, L, specimen YMG4–13, Stn 300; M, N, specimen GLD4–09, Stn 198; O, P, specimen YMG4–13, Stn 274. A, B, four-zooid stage; C–F, four–five-zooid stages; G–J, five–six-zooid stages; K–P, later submature stages with calyciform colony center adequately structured with trabeculae and alveoli to support gonozooid development. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437909/files/figure.png" pageId="44" pageNumber="45">Fig. 30O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F2561E552B09BEF9518F3FF8D9" blockId="44.[151,1437,1033,2030]" pageId="44" pageNumber="45">
|
||
Autozooidal peristomes in single subregular series, skeletal surface like that of column, with sparse, simple pores (
|
||
<figureCitation id="42872A77561E552B099EF9758B1AF891" box="[231,336,1789,1814]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29G</figureCitation>
|
||
) having irregular outline, tending to be in single longitudinal series on lower side (
|
||
<figureCitation id="42872A77561E552B0C5EF9758FDAF891" box="[1319,1424,1789,1814]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29L</figureCitation>
|
||
). Developing apertures slightly irregular, subrounded (
|
||
<figureCitation id="42872A77561E552B0B90F8A9893BF8BD" box="[745,881,1825,1850]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29A, K</figureCitation>
|
||
) or having bilateral symmetry, opening obliquely outwards and downwards (
|
||
<figureCitation id="42872A77561E552B08BDF8CD8860F8D9" box="[452,554,1861,1886]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29L</figureCitation>
|
||
). No spinules or prickles on inner surface of peristome tubes (
|
||
<figureCitation id="42872A77561E552B0D99F8CD8F22F8D9" box="[1248,1384,1861,1886]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29K, L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F2561E552A09BEF8E188E9FF53" blockId="44.[151,1437,1033,2030]" lastBlockId="45.[151,1436,151,933]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
|
||
Gonozooid large and capacious (
|
||
<figureCitation id="42872A77561E552B0B44F8E18882F805" box="[573,712,1897,1922]" captionStart="FIGURE 28" captionStartId="46.[151,250,1932,1954]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure@46.[174,1418,193,1917]" captionTargetPageId="46" captionText="FIGURE 28. Calyssopora volcano n. gen., n. sp. A, B, holotype, ZIRAS 1/50714, with a single gonozooid; C, D, paratype 1, ZIRAS 2/50715, with two ooeciostomes and therefore two gonozooids presumed; E, F, paratype 2, ZIRAS 3/50716, with possibly newly completed gonozooid, having large alveolate spaces at left-hand periphery. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437905/files/figure.png" pageId="44" pageNumber="45">Fig. 28A, B</figureCitation>
|
||
), 1–2 per mature colony (
|
||
<figureCitation id="42872A77561E552B0A8BF8E18E35F805" box="[1010,1151,1897,1922]" captionStart="FIGURE 28" captionStartId="46.[151,250,1932,1954]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure@46.[174,1418,193,1917]" captionTargetPageId="46" captionText="FIGURE 28. Calyssopora volcano n. gen., n. sp. A, B, holotype, ZIRAS 1/50714, with a single gonozooid; C, D, paratype 1, ZIRAS 2/50715, with two ooeciostomes and therefore two gonozooids presumed; E, F, paratype 2, ZIRAS 3/50716, with possibly newly completed gonozooid, having large alveolate spaces at left-hand periphery. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437905/files/figure.png" pageId="44" pageNumber="45">Fig. 28C, D</figureCitation>
|
||
). Floor of gonozooid not discrete and continuous. If solo, gonozooid in center of calyx or to one side, chamber roof sloping upwards to ooeciostome (
|
||
<figureCitation id="42872A77561E552B084AF8398BF4F84D" box="[307,446,1969,1994]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29A–D</figureCitation>
|
||
), resembling volcanic cone in profile, with lateral ridges and furrows and porous cancelli that are mostly on periphery of gonozooid (
|
||
<figureCitation id="42872A77561E552B0BF2F85D88BFF869" box="[651,757,2005,2030]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="44" pageNumber="45">Fig. 29D</figureCitation>
|
||
); ooeciopore variably elevated, subcircular to suboval (
|
||
<figureCitation id="42872A77561E552A0C16F85D8B51FF37" captionStart="FIGURE 28" captionStartId="46.[151,250,1932,1954]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure@46.[174,1418,193,1917]" captionTargetPageId="46" captionText="FIGURE 28. Calyssopora volcano n. gen., n. sp. A, B, holotype, ZIRAS 1/50714, with a single gonozooid; C, D, paratype 1, ZIRAS 2/50715, with two ooeciostomes and therefore two gonozooids presumed; E, F, paratype 2, ZIRAS 3/50716, with possibly newly completed gonozooid, having large alveolate spaces at left-hand periphery. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437905/files/figure.png" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">Fig. 28A, C, E</figureCitation>
|
||
). Surface microstructure of gonozooid like that of column and zooidal peristomes, with weak imbrication of irregular crystallites (
|
||
<figureCitation id="42872A77561F552A0B54FF3488DCFF53" box="[557,662,188,213]" captionStart="FIGURE 29" captionStartId="47.[151,250,1837,1859]" captionTargetBox="[194,1399,193,1821]" captionTargetId="figure@47.[194,1399,193,1821]" captionTargetPageId="47" captionText="FIGURE 29. Calyssopora volcano n. gen., n. sp. A, D, L, holotype, ZIRAS 1/50714; B, E, G–I, K, paratype 1, ZIRAS 2/ 50715; C, F, J, paratype 2, ZIRAS 3/50716. A–C, oblique profiles of fertile colonies respectively shown in Fig. 28A, B, Fig. 28B, C and Fig. 28E, F; D–F, peristome of gonozooids respectively shown in A–C; G, calyx periphery showing short autozooidal peristomes, shallow alveoli and pores; H, skeletal microstructure of slope of ooeciostome; I, peristomes in oblique profile; J, abfrontal side of peristomes; K, two autozooidal apertures with large alveolus between; L, oblique bilaterally symmetrical autozooidal apertures. Scale bars: A–C, J, 250 µm; D–G, I, K, L, 150 µm; H, 25 µm." httpUri="https://zenodo.org/record/1437907/files/figure.png" pageId="45" pageNumber="46">Fig. 29H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F2561F552A09BEFF578993FEC7" blockId="45.[151,1436,151,933]" pageId="45" pageNumber="46">
|
||
Ancestrula not seen in isolation, but evidently erect judging from smallest (four-zooid) colony stages seen (
|
||
<figureCitation id="42872A77561F552A0C16FF578ABBFE9B" captionStart="FIGURE 30" captionStartId="48.[151,250,1846,1868]" captionTargetBox="[183,1407,185,1834]" captionTargetId="figure@48.[183,1407,185,1834]" captionTargetPageId="48" captionText="FIGURE 30. Calyssopora volcano n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. A, B, specimen YMG4–13, Stn 291; C, D, specimen YMG4–07, Stn 139; E, F, specimen YMG4–14, Stn 358; G, H, specimen YMG4–04, Stn 57; I, J, specimen YMG4–13, Stn 283; K, L, specimen YMG4–13, Stn 300; M, N, specimen GLD4–09, Stn 198; O, P, specimen YMG4–13, Stn 274. A, B, four-zooid stage; C–F, four–five-zooid stages; G–J, five–six-zooid stages; K–P, later submature stages with calyciform colony center adequately structured with trabeculae and alveoli to support gonozooid development. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437909/files/figure.png" pageId="45" pageNumber="46">Fig. 30A, B</figureCitation>
|
||
). Trabeculae form in center of developing colony to support colony elevation and expansion, with extrazooidal alveolar spaces between trabeculae (
|
||
<figureCitation id="42872A77561F552A0BB9FEAF8981FEC7" box="[704,971,295,320]" captionStart="FIGURE 30" captionStartId="48.[151,250,1846,1868]" captionTargetBox="[183,1407,185,1834]" captionTargetId="figure@48.[183,1407,185,1834]" captionTargetPageId="48" captionText="FIGURE 30. Calyssopora volcano n. gen., n. sp. Progressive stages of development of ancestrular and young colonies. A, B, specimen YMG4–13, Stn 291; C, D, specimen YMG4–07, Stn 139; E, F, specimen YMG4–14, Stn 358; G, H, specimen YMG4–04, Stn 57; I, J, specimen YMG4–13, Stn 283; K, L, specimen YMG4–13, Stn 300; M, N, specimen GLD4–09, Stn 198; O, P, specimen YMG4–13, Stn 274. A, B, four-zooid stage; C–F, four–five-zooid stages; G–J, five–six-zooid stages; K–P, later submature stages with calyciform colony center adequately structured with trabeculae and alveoli to support gonozooid development. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437909/files/figure.png" pageId="45" pageNumber="46">Fig. 30A, C, I, K, M, O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F2561F552A09BEFEC48BE2FE73" blockId="45.[151,1436,151,933]" pageId="45" pageNumber="46">
|
||
<emphasis id="E8C8EAE0561F552A09BEFEC48B8CFEE2" bold="true" box="[199,454,332,357]" pageId="45" pageNumber="46">Measurements (mm).</emphasis>
|
||
<typeStatus id="05078850561F552A08B6FEC48871FEE2" box="[463,571,332,357]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
|
||
, ZIRAS 1/50714 (
|
||
<figureCitation id="42872A77561F552A0A74FEC489D2FEE3" box="[781,920,332,357]" captionStart="FIGURE 28" captionStartId="46.[151,250,1932,1954]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure@46.[174,1418,193,1917]" captionTargetPageId="46" captionText="FIGURE 28. Calyssopora volcano n. gen., n. sp. A, B, holotype, ZIRAS 1/50714, with a single gonozooid; C, D, paratype 1, ZIRAS 2/50715, with two ooeciostomes and therefore two gonozooids presumed; E, F, paratype 2, ZIRAS 3/50716, with possibly newly completed gonozooid, having large alveolate spaces at left-hand periphery. Scale bars: 250 µm." httpUri="https://zenodo.org/record/1437905/files/figure.png" pageId="45" pageNumber="46">Fig. 28A, B</figureCitation>
|
||
): Colony height 1.78; capitulum 2.07 × 1.98; base at substratum 0.85 × 0.93 with narrowest cross-section 0.60 × 0.64; ZL 0.591–0.980 (0.809 ± 0.122); PeL 0.372–0.597 (0.492 ± 0.069); PeD 0.137–0.177 (0.155 ± 0.013); ApL 0.152–0.185 (0.172 ± 0.011); ApW 0.128–0.153 (0.138 ± 0.009). Gonozooid (
|
||
<emphasis id="E8C8EAE0561F552A0B04FE3188C1FE57" box="[637,651,441,464]" italics="true" pageId="45" pageNumber="46">n</emphasis>
|
||
= 1): GZL 0.644; GZW 0.817; OsL 0.155; OsW 0.127; OsH 0.138; OpL 0.133; OpW 0.119.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A66579561F552A09BEFE778F19FD02" pageId="45" pageNumber="46" type="discussion">
|
||
<paragraph id="DA0336F2561F552A09BEFE778F19FD02" blockId="45.[151,1436,151,933]" pageId="45" pageNumber="46">
|
||
<emphasis id="E8C8EAE0561F552A09BEFE778B71FD9F" bold="true" box="[199,315,511,536]" pageId="45" pageNumber="46">Remarks.</emphasis>
|
||
<taxonomicName id="1DBC4D71561F552A083BFE77887AFD9F" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[322,560,511,536]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="46" phylum="Bryozoa" rank="species" species="volcano" status="sp. nov.">
|
||
<emphasis id="E8C8EAE0561F552A083BFE77887AFD9F" box="[322,560,511,536]" italics="true" pageId="45" pageNumber="46">Calyssopora volcano</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8C8EAE0561F552A0B4EFE77883CFD9F" bold="true" box="[567,630,511,536]" pageId="45" pageNumber="46">
|
||
<taxonomicNameLabel id="F3FB579B561F552A0B4EFE77883CFD9F" box="[567,630,511,536]" pageId="45" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="1DBC4D71561F552A0A75FE778E5CFD9F" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[780,1046,511,536]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="46" phylum="Bryozoa" rank="species" species="vasiformis" status="sp. nov.">
|
||
<emphasis id="E8C8EAE0561F552A0A75FE778E5CFD9F" box="[780,1046,511,536]" italics="true" pageId="45" pageNumber="46">Calyssopora vasiformis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8C8EAE0561F552A0D64FE778E17FD9F" bold="true" box="[1053,1117,511,536]" pageId="45" pageNumber="46">
|
||
<taxonomicNameLabel id="F3FB579B561F552A0D64FE778E17FD9F" box="[1053,1117,511,536]" pageId="45" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(below) in overall form and the number of peristomes, and from
|
||
<taxonomicName id="1DBC4D71561F552A0B4FFDAC891FFDBA" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[566,853,548,573]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="46" phylum="Bryozoa" rank="species" species="clarionensis" status="sp. nov.">
|
||
<emphasis id="E8C8EAE0561F552A0B4FFDAC891FFDBA" box="[566,853,548,573]" italics="true" pageId="45" pageNumber="46">Calyssopora clarionensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8C8EAE0561F552A0A24FDAC89D7FDBA" bold="true" box="[861,925,548,573]" pageId="45" pageNumber="46">
|
||
<taxonomicNameLabel id="F3FB579B561F552A0A24FDAC89D7FDBA" box="[861,925,548,573]" pageId="45" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, in having volcaniform gonozooids in which the frontal surface rises to a frontally facing ooeciostome with a crater-like ooeciopore. The gonozooid in
|
||
<taxonomicName id="1DBC4D71561F552A0CFDFDC08B6AFD02" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="46" phylum="Bryozoa" rank="species" species="clarionensis">
|
||
<emphasis id="E8C8EAE0561F552A0CFDFDC08B6AFD02" italics="true" pageId="45" pageNumber="46">A. clarionensis</emphasis>
|
||
</taxonomicName>
|
||
is rather more bulging than volcaniform, and the ooeciostome is strongly curved and hood-like.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A66579561F552A09BEFD07896CFCDA" pageId="45" pageNumber="46" type="reference_group">
|
||
<paragraph id="DA0336F2561F552A09BEFD07896CFCDA" blockId="45.[151,1436,151,933]" pageId="45" pageNumber="46">
|
||
<taxonomicName id="1DBC4D71561F552A09BEFD07895EFD2F" authority="Gordon & Taylor, 2010" authorityName="Gordon & Taylor" authorityYear="2010" box="[199,788,655,680]" class="Stenolaemata" family="Lichenoporidae" genus="Dartevellopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="46" phylum="Bryozoa" rank="species" species="granulosa">
|
||
<emphasis id="E8C8EAE0561F552A09BEFD078BB8FD2F" box="[199,498,655,680]" italics="true" pageId="45" pageNumber="46">Dartevellopora granulosa</emphasis>
|
||
<bibRefCitation id="BE2D4B03561F552A0884FD07895EFD2F" box="[509,788,655,680]" pageId="45" pageNumber="46" refId="ref34174" refString="Gordon, D. P. & Taylor, P. D. (2010) New seamount- and ridge-associated cyclostome Bryozoa from New Zealand. Zootaxa, 2533, 43 - 68." type="journal article">Gordon & Taylor, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
was only doubtfully included in
|
||
<taxonomicName id="1DBC4D71561F552A0DDFFD078F19FD2F" ID-CoL="84PGT" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[1190,1363,655,680]" class="Stenolaemata" family="Lichenoporidae" genus="Dartevellopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="46" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="E8C8EAE0561F552A0DDFFD078F19FD2F" box="[1190,1363,655,680]" italics="true" pageId="45" pageNumber="46">Dartevellopora</emphasis>
|
||
</taxonomicName>
|
||
when described. In some respects, it resembles the species of
|
||
<taxonomicName id="1DBC4D71561F552A0A2BFD3C8995FD4A" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[850,991,692,717]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="64" phylum="Bryozoa" rank="genus" status="gen. nov.">
|
||
<emphasis id="E8C8EAE0561F552A0A2BFD3C8995FD4A" box="[850,991,692,717]" italics="true" pageId="45" pageNumber="46">Calyssopora</emphasis>
|
||
</taxonomicName>
|
||
, but differs in that there is a clear differentiation between a smoothly spreading skirt-like base and pustulose erect part of the column. The center of its capitulum is less calyciform than in
|
||
<taxonomicName id="1DBC4D71561F552A0B2FFD7488AEFC92" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[598,740,764,789]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="64" phylum="Bryozoa" rank="genus" status="gen. nov.">
|
||
<emphasis id="E8C8EAE0561F552A0B2FFD7488AEFC92" box="[598,740,764,789]" italics="true" pageId="45" pageNumber="46">Calyssopora</emphasis>
|
||
</taxonomicName>
|
||
, comprising a subhorizontal platform with evenly distributed circular pores, unlike the open alveolate and trabecular mesh of
|
||
<taxonomicName id="1DBC4D71561F552A0A18FC9789A4FCBF" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[865,1006,799,824]" class="Stenolaemata" family="Lichenoporidae" genus="Calyssopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="64" phylum="Bryozoa" rank="genus" status="gen. nov.">
|
||
<emphasis id="E8C8EAE0561F552A0A18FC9789A4FCBF" box="[865,1006,799,824]" italics="true" pageId="45" pageNumber="46">Calyssopora</emphasis>
|
||
</taxonomicName>
|
||
. Until fertile colonies of
|
||
<taxonomicName id="1DBC4D71561F552A0C71FCA98FD6FCBF" authorityName="Gordon & Taylor" authorityYear="2010" box="[1288,1436,799,824]" class="Stenolaemata" family="Lichenoporidae" genus="Dartevellopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="45" pageNumber="46" phylum="Bryozoa" rank="species" species="granulosa">
|
||
<emphasis id="E8C8EAE0561F552A0C71FCA98FD6FCBF" box="[1288,1436,799,824]" italics="true" pageId="45" pageNumber="46">D. granulosa</emphasis>
|
||
</taxonomicName>
|
||
are found, its generic attribution must remain problematic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A66579561F552A09BEFCEF884FFC23" pageId="45" pageNumber="46" type="distribution">
|
||
<paragraph id="DA0336F2561F552A09BEFCEF884FFC23" blockId="45.[151,1436,151,933]" pageId="45" pageNumber="46">
|
||
<emphasis id="E8C8EAE0561F552A09BEFCEF8B2AFC07" bold="true" box="[199,352,871,896]" pageId="45" pageNumber="46">Distribution.</emphasis>
|
||
Recorded from 26 stations within coordinates 12.51953–
|
||
<geoCoordinate id="BF885035561F552A0A8DFCEF8EC2FCF8" box="[1012,1160,871,896]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="1" value="14.1649">14.16490° N</geoCoordinate>
|
||
, 129.04471–
|
||
<geoCoordinate id="BF885035561F552A0C62FCEF8AF8FC23" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="1" value="-134.60008">134.60008° W</geoCoordinate>
|
||
, at depth range
|
||
<quantity id="1D449B17561F552A081BFC04884AFC23" box="[354,512,908,933]" metricMagnitude="3" metricUnit="m" metricValue="4.9105" metricValueMax="5.148999999999999" metricValueMin="4.672" pageId="45" pageNumber="46" unit="m" value="4910.5" valueMax="5149.0" valueMin="4672.0">4672–5149 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |