236 lines
47 KiB
XML
236 lines
47 KiB
XML
<document id="3FE4321021A6E04AD67D44AC64E9D3D8" ID-CLB-Dataset="29502" ID-DOI="10.11646/zootaxa.4484.1.1" ID-GBIF-Dataset="74b777c9-eae0-4770-8c86-dcbb10fb06b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437848" ID-ZooBank="D66524CF-9C6D-4DF4-8CA2-B2C9708CF5FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538125412450" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P. & Melnik, Viacheslav P." docDate="2018" docId="521587E45609553B09EEFF1F884FFE9B" docLanguage="en" docName="zootaxa.4484.1.1.pdf" docOrigin="Zootaxa 4484 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anyuta anastema Grischenko & Gordon & Melnik 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="61" masterDocId="AE2CFF9C563255070979FF888A4AFF87" masterDocTitle="Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining" masterLastPageNumber="91" masterPageNumber="1" pageNumber="60" updateTime="1698557051871" updateUser="ExternalLinkService">
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<mods:title id="91CDAE52FA9DD947DCB90C0BB2D7A7C1">Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining</mods:title>
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<mods:name id="FA37B5DB7A304861D9D8A8FA05CA86F2" type="personal">
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<mods:namePart id="B409C25BA226889C8859618BBD99B5CC">Grischenko, Andrei V.</mods:namePart>
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<mods:namePart id="84F4D99FA0E4E6BD47B97881C909B517">Gordon, Dennis P.</mods:namePart>
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<mods:name id="6AA50BFFBB96E0B77379768AB8B79D85" type="personal">
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<mods:roleTerm id="714FCF7C36BCAF98945239335974730F">Author</mods:roleTerm>
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<mods:namePart id="DDBAD75A49C75CFE24B8DD040EA321FD">Melnik, Viacheslav P.</mods:namePart>
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<mods:typeOfResource id="D2A9C29E23A976AFF4C448B4B09F723D">text</mods:typeOfResource>
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<mods:title id="418EDFC679F079F53F827A0483CB3980">Zootaxa</mods:title>
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<mods:part id="5E0898E31497137F216A46AEA627D315">
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<mods:date id="54244CE7716A3470CBB302880F7B75F3">2018</mods:date>
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<mods:number id="1F95A4C0C5AAB3EA051ECE069C6A30D1">2018-09-25</mods:number>
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<mods:number id="0796E8B9A92E4352909B24216BD9E151">4484</mods:number>
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<treatment id="521587E45609553B09EEFF1F884FFE9B" ID-DOI="http://doi.org/10.5281/zenodo.5989827" ID-GBIF-Taxon="148403736" ID-Zenodo-Dep="5989827" LSID="urn:lsid:plazi:treatment:521587E45609553B09EEFF1F884FFE9B" httpUri="http://treatment.plazi.org/id/521587E45609553B09EEFF1F884FFE9B" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
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<subSubSection id="92A665795609553C09EEFF1F8BE9FF54" pageId="59" pageNumber="60" type="nomenclature">
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<paragraph id="DA0336F25609553C09EEFF1F8BF9FF35" blockId="59.[151,435,151,211]" box="[151,435,151,178]" pageId="59" pageNumber="60">
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<heading id="814B819E5609553C09EEFF1F8BF9FF35" bold="true" box="[151,435,151,178]" fontSize="11" level="1" pageId="59" pageNumber="60" reason="1">
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<emphasis id="E8C8EAE05609553C09EEFF1F8BF9FF35" bold="true" box="[151,435,151,178]" pageId="59" pageNumber="60">
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<taxonomicName id="1DBC4D715609553C09EEFF1F8B20FF36" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[151,362,151,177]" class="Stenolaemata" family="Lichenoporidae" genus="Anyuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="59" pageNumber="60" phylum="Bryozoa" rank="species" species="anastema" status="sp. nov.">
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<emphasis id="E8C8EAE05609553C09EEFF1F8B20FF36" bold="true" box="[151,362,151,177]" italics="true" pageId="59" pageNumber="60">Anyuta anastema</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F3FB579B5609553C0809FF108BF9FF35" box="[368,435,152,178]" pageId="59" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="DA0336F25609553C09EEFF328BE9FF54" blockId="59.[151,435,151,211]" box="[151,419,186,211]" pageId="59" pageNumber="60">
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(
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<figureCitation id="42872A775609553C09E6FF328B54FF54" box="[159,286,186,211]" captionStart="FIGURE 2" captionStartId="10.[151,250,1774,1796]" captionTargetBox="[151,1436,392,1715]" captionTargetId="figure@10.[151,1436,392,1715]" captionTargetPageId="10" captionText="FIGURE 2. Colonies of some cyclostome bryozoans, in vivo, attached to polymetallic nodules. A–E, Pandanipora helix n. gen., n. sp.: A, specimen GLD4–09, Stn 190; B, specimen GLD4–12, Stn 262; C, specimen YMG4–07, Stn 143; D, specimen YMG4–13, Stn 295; E, specimen GLD 4–11, Stn 212. F, Tubuliporina sp. indet., specimen YMG18–01, Stn 7. G, H, Abyssoecia elevata n. gen., n. sp.: G, specimen GLD4–09, Stn 196; H, specimen GLD4–09, Stn 191. I, Discantenna metallica n. sp.: specimen GLD4–11, Stn 224. J, K, Frontohornera frontalis n. gen., n. sp.: J, specimen YMG4–07, Stn 124; K, specimen GLD4–11, Stn 210. L, Alyonushka hystricosa n. gen., n. sp.: specimen GLD4–09, Stn 199. M, Calyssopora volcano n. gen., n. sp.: specimen YMG18–01, Stn 33. N, O, Anyuta anastema n. gen., n. sp.: N, specimen GLD4–09, Stn 180; O, specimen YMG4–06, Stn 71. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437852/files/figure.png" pageId="59" pageNumber="60">Figs 2N, O</figureCitation>
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,
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<figureCitation id="42872A775609553C0853FF338B39FF54" box="[298,371,187,211]" captionStart-0="FIGURE 39" captionStart-1="FIGURE 40" captionStart-2="FIGURE 41" captionStartId-0="60.[151,250,1940,1962]" captionStartId-1="61.[151,250,1823,1845]" captionStartId-2="62.[151,250,1896,1918]" captionTargetBox-0="[166,1420,339,1927]" captionTargetBox-1="[222,1364,193,1809]" captionTargetBox-2="[161,1426,183,1886]" captionTargetId-0="figure@60.[166,1420,339,1927]" captionTargetId-1="figure@61.[222,1364,193,1809]" captionTargetId-2="figure@62.[161,1426,183,1886]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionTargetPageId-2="62" captionText-0="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." captionText-1="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." captionText-2="FIGURE 41. Anyuta anastema n. gen., n. sp. Progressive stages of development of young colonies. A–D, specimen YMG4–14, Stn 360, three-zooid stage; E–H, specimen YMG4–14, Stn 338, five–six-zooid stage; I–L, specimen YMG4–13, Stn 281, and M–P, specimen YMG4–06, Stn 71, showing later stages in which trabecular walls define spaces destined to become either autozooids or alveoli. Scale bars: 200 µm." httpUri-0="https://zenodo.org/record/1437927/files/figure.png" httpUri-1="https://zenodo.org/record/1437929/files/figure.png" httpUri-2="https://zenodo.org/record/1437931/files/figure.png" pageId="59" pageNumber="60">39–41</figureCitation>
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,
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<figureCitation id="42872A775609553C0807FF328BD1FF54" box="[382,411,186,211]" captionStart="FIGURE 53" captionStartId="78.[151,250,1823,1845]" captionTargetBox="[151,1436,210,1786]" captionTargetId="figure@78.[151,1436,210,1786]" captionTargetPageId="78" captionText="FIGURE 53. Anyuta anastema n. gen., n. sp. Micro-CT scans of paratype, NIWA 127725, as surface (C) and back-face isosurface renders showing colony interiors. A, view of colony from below; note absence of kenozooids from most of column; B, lateral abfrontal view of two small fascicles with entrance to dimorphic zooid indicated by arrowhead; note relative paucity of communication pores; C–E, exterior and interior views of colony base, showing lateral (C, D) and frontolateral (E) profiles with palisade of elongate-triangular kenozooidal chambers surrounding ancestrular protoecium (pr) and proximal part of its peristome (ap). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1437955/files/figure.png" pageId="59" pageNumber="60">53</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="92A665795609553C09EEFE8A88BCFE48" pageId="59" pageNumber="60" type="materials_examined">
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<paragraph id="DA0336F25609553C09EEFE8A88BCFE48" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
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<emphasis id="E8C8EAE05609553C09EEFE8A8BCCFE9C" bold="true" box="[151,390,258,283]" pageId="59" pageNumber="60">Material examined.</emphasis>
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<emphasis id="E8C8EAE05609553C08EBFE8A8BB3FE9C" box="[402,505,258,283]" italics="true" pageId="59" pageNumber="60">Holotype</emphasis>
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: ZIRAS 1/50718, colony detached from nodule, YMG R.V.
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<emphasis id="E8C8EAE05609553C0DB6FE8A8F07FE9C" box="[1231,1357,258,283]" italics="true" pageId="59" pageNumber="60">Gelendzhik</emphasis>
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cruise GLD4–09, Stn 180,
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<date id="AE0210325609553C08F9FEAE8811FEB8" box="[384,603,294,319]" pageId="59" pageNumber="60" value="2010-12-26">26 December 2010</date>
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, 13.03863° N, 133.39428° W,
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<quantity id="1D449B175609553C0AC5FEAF8E5CFEB9" box="[956,1046,295,319]" metricMagnitude="3" metricUnit="m" metricValue="4.919" pageId="59" pageNumber="60" unit="m" value="4919.0">4919 m</quantity>
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.
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<emphasis id="E8C8EAE05609553C0D5AFEAF8EC0FEB9" box="[1059,1162,295,318]" italics="true" pageId="59" pageNumber="60">Paratype</emphasis>
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: NIWA 127725, colony detached from nodule, YMG R.V.
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<emphasis id="E8C8EAE05609553C0B5AFEC288A5FEE4" box="[547,751,330,355]" italics="true" pageId="59" pageNumber="60">Yuzhmorgeologiya</emphasis>
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cruise YMG4–14, Stn 359,
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<date id="AE0210325609553C0D4EFEC38EBDFEE4" box="[1079,1271,330,355]" pageId="59" pageNumber="60" value="2016-01-19">19 January 2016</date>
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, 14.08687° N, 131.78558° W,
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<quantity id="1D449B175609553C0830FEE68BE9FE01" box="[329,419,366,391]" metricMagnitude="3" metricUnit="m" metricValue="5.122" pageId="59" pageNumber="60" unit="m" value="5122.0">5122 m</quantity>
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.
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<emphasis id="E8C8EAE05609553C08D6FEE688C6FE00" box="[431,652,366,391]" italics="true" pageId="59" pageNumber="60">Additional material</emphasis>
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: YMG18–01, Stn 29; YMG4–06, Stn 71; YMG4–07, Stns 119, 133; GLD4–08, Stn 144; GLD4–11, Stn 227; YMG4–13, Stns 281, 283; YMG4–14, Stns 338, 360. Total specimens examined 12, two (the largest) with dimorphic zooids.
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</paragraph>
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</subSubSection>
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<subSubSection id="92A665795609553C09BEFE52880DFD90" pageId="59" pageNumber="60" type="etymology">
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<paragraph id="DA0336F25609553C09BEFE52880DFD90" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
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<emphasis id="E8C8EAE05609553C09BEFE528B04FE74" bold="true" box="[199,334,474,499]" pageId="59" pageNumber="60">Etymology.</emphasis>
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Greek,
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<taxonomicName id="1DBC4D715609553C08D3FE54885CFE74" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[426,534,476,499]" class="Stenolaemata" family="Lichenoporidae" genus="Anyuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="59" pageNumber="60" phylum="Bryozoa" rank="species" species="anastema">
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<emphasis id="E8C8EAE05609553C08D3FE54885CFE74" box="[426,534,476,499]" italics="true" pageId="59" pageNumber="60">anastema</emphasis>
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</taxonomicName>
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, height, tallness, alluding to the elevated form of the colony with the branches lifted up, used as a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="92A665795609553C09BEFDAA8E30F830" pageId="59" pageNumber="60" type="description">
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<paragraph id="DA0336F25609553C09BEFDAA8B59FD68" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
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<emphasis id="E8C8EAE05609553C09BEFDAA8B13FDBC" bold="true" box="[199,345,546,571]" pageId="59" pageNumber="60">Description.</emphasis>
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Colony erect, pedunculate and subcalyciform, with outwardly flaring capitulum of short branches (
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<figureCitation id="42872A775609553C09E6FDCE8B61FDD8" box="[159,299,582,607]" captionStart="FIGURE 39" captionStartId="60.[151,250,1940,1962]" captionTargetBox="[166,1420,339,1927]" captionTargetId="figure@60.[166,1420,339,1927]" captionTargetPageId="60" captionText="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437927/files/figure.png" pageId="59" pageNumber="60">Fig. 39A, B</figureCitation>
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). Stalk (column) cylindrical, tapering proximally before expanding to form circular disk (
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<figureCitation id="42872A775609553C0C49FDCE8AE5FD04" captionStart="FIGURE 39" captionStartId="60.[151,250,1940,1962]" captionTargetBox="[166,1420,339,1927]" captionTargetId="figure@60.[166,1420,339,1927]" captionTargetPageId="60" captionText="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437927/files/figure.png" pageId="59" pageNumber="60">Fig. 39C, D</figureCitation>
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). Branches of
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<typeStatus id="050788505609553C082CFDE38BCFFD04" box="[341,389,619,643]" pageId="59" pageNumber="60">type</typeStatus>
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colony disposed more or less in two clusters, each comprising two fascicles of zooids with associated structures. Length of outwardly inclined branches shorter than stalk. All surfaces interior-walled, with variably imbricating platy microstructure or the plate-like crystalline ‘tablets’ commonly irregular and less angular (
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<figureCitation id="42872A775609553C09E6FD5E8B4DFD68" box="[159,263,726,751]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">Fig. 40C</figureCitation>
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).
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</paragraph>
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<paragraph id="DA0336F25609553C09BEFD72886DFC6C" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
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Autozooidal tubes numbering
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each of the fascicles, the tubes indicated by their entire, round peristomial rims (
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<figureCitation id="42872A775609553C09A2FC968B3AFCB1" box="[219,368,798,823]" captionStart="FIGURE 39" captionStartId="60.[151,250,1940,1962]" captionTargetBox="[166,1420,339,1927]" captionTargetId="figure@60.[166,1420,339,1927]" captionTargetPageId="60" captionText="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437927/files/figure.png" pageId="59" pageNumber="60">Figs 39C, D</figureCitation>
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,
|
||
<figureCitation id="42872A775609553C0806FC978BA1FCB1" box="[383,491,799,823]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">40D–G, I</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775609553C0882FC978826FCB0" box="[507,620,799,823]" captionStart="FIGURE 41" captionStartId="62.[151,250,1896,1918]" captionTargetBox="[161,1426,183,1886]" captionTargetId="figure@62.[161,1426,183,1886]" captionTargetPageId="62" captionText="FIGURE 41. Anyuta anastema n. gen., n. sp. Progressive stages of development of young colonies. A–D, specimen YMG4–14, Stn 360, three-zooid stage; E–H, specimen YMG4–14, Stn 338, five–six-zooid stage; I–L, specimen YMG4–13, Stn 281, and M–P, specimen YMG4–06, Stn 71, showing later stages in which trabecular walls define spaces destined to become either autozooids or alveoli. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437931/files/figure.png" pageId="59" pageNumber="60">41A, E, I</figureCitation>
|
||
). Exterior skeletal surface of peristomial walls minutely irregular and uneven (
|
||
<figureCitation id="42872A775609553C0983FCCA8BC2FCDC" box="[250,392,834,859]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">Fig. 40B, G</figureCitation>
|
||
), lightly textured, with variable expression of partial longitudinal and/or transverse ridges, striae and convexities. Pores simple, open, of variable shape, sparse. Peristomial rim even or slightly uneven, smooth or minutely granular. Skeletal ultrastructure of interior surface of peristomes similar to that of exterior but smoother and crystallites less discrete; no spinules or pustules. CT scans reveal that interzooidal communication pores are very sparse (
|
||
<figureCitation id="42872A775609553C08EBFC5A8853FC6C" box="[402,537,978,1003]" captionStart="FIGURE 53" captionStartId="78.[151,250,1823,1845]" captionTargetBox="[151,1436,210,1786]" captionTargetId="figure@78.[151,1436,210,1786]" captionTargetPageId="78" captionText="FIGURE 53. Anyuta anastema n. gen., n. sp. Micro-CT scans of paratype, NIWA 127725, as surface (C) and back-face isosurface renders showing colony interiors. A, view of colony from below; note absence of kenozooids from most of column; B, lateral abfrontal view of two small fascicles with entrance to dimorphic zooid indicated by arrowhead; note relative paucity of communication pores; C–E, exterior and interior views of colony base, showing lateral (C, D) and frontolateral (E) profiles with palisade of elongate-triangular kenozooidal chambers surrounding ancestrular protoecium (pr) and proximal part of its peristome (ap). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1437955/files/figure.png" pageId="59" pageNumber="60">Fig. 53A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F25609553C09BEFC7F8E2DFB18" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
|
||
<materialsCitation id="6AD43CAF5609553C09BEFC7F8E2DFB18" ID-GBIF-Occurrence="1913869016" collectingDate="1941-01" location="Center" pageId="59" pageNumber="60" specimenCount="1" stateProvince="Center">
|
||
<collectingRegion id="1878F8105609553C09BEFC7F8B58FB88" box="[199,274,1015,1039]" country="France" name="Centre" pageId="59" pageNumber="60">Center</collectingRegion>
|
||
of sides of capitulum with alveolar/kenozooidal chambers (
|
||
<figureCitation id="42872A775609553C0AACFC7E8E0CFB88" box="[981,1094,1014,1039]" captionStart="FIGURE 39" captionStartId="60.[151,250,1940,1962]" captionTargetBox="[166,1420,339,1927]" captionTargetId="figure@60.[166,1420,339,1927]" captionTargetPageId="60" captionText="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437927/files/figure.png" pageId="59" pageNumber="60">Figs 39A</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775609553C0D2FFC7F8EC2FB88" box="[1110,1160,1015,1039]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">40A</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775609553C0DE0FC7F8EA6FB89" box="[1177,1260,1015,1039]" captionStart="FIGURE 41" captionStartId="62.[151,250,1896,1918]" captionTargetBox="[161,1426,183,1886]" captionTargetId="figure@62.[161,1426,183,1886]" captionTargetPageId="62" captionText="FIGURE 41. Anyuta anastema n. gen., n. sp. Progressive stages of development of young colonies. A–D, specimen YMG4–14, Stn 360, three-zooid stage; E–H, specimen YMG4–14, Stn 338, five–six-zooid stage; I–L, specimen YMG4–13, Stn 281, and M–P, specimen YMG4–06, Stn 71, showing later stages in which trabecular walls define spaces destined to become either autozooids or alveoli. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437931/files/figure.png" pageId="59" pageNumber="60">
|
||
<date id="AE0210325609553C0DE0FC7F8E8AFB88" box="[1177,1216,1015,1039]" pageId="59" pageNumber="60" value="1941-01">
|
||
<collectingDate id="BE46E9DA5609553C0DE0FC7F8E8AFB88" box="[1177,1216,1015,1039]" pageId="59" pageNumber="60" value="1941-01">41I</collectingDate>
|
||
</date>
|
||
, M
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775609553C0D85FC7E8F7BFB88" box="[1276,1329,1014,1039]" captionStart="FIGURE 53" captionStartId="78.[151,250,1823,1845]" captionTargetBox="[151,1436,210,1786]" captionTargetId="figure@78.[151,1436,210,1786]" captionTargetPageId="78" captionText="FIGURE 53. Anyuta anastema n. gen., n. sp. Micro-CT scans of paratype, NIWA 127725, as surface (C) and back-face isosurface renders showing colony interiors. A, view of colony from below; note absence of kenozooids from most of column; B, lateral abfrontal view of two small fascicles with entrance to dimorphic zooid indicated by arrowhead; note relative paucity of communication pores; C–E, exterior and interior views of colony base, showing lateral (C, D) and frontolateral (E) profiles with palisade of elongate-triangular kenozooidal chambers surrounding ancestrular protoecium (pr) and proximal part of its peristome (ap). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1437955/files/figure.png" pageId="59" pageNumber="60">53A</figureCitation>
|
||
). Alveoli forming as low walls, typically arcuate, on upper surfaces. As these walls grow, the extrazooidal cavities they enclose become deeper, cupuliform, and kenozooid-like, filling in spaces between autozooidal peristomes, sometimes in stepwise series. Some alveoli very elongate, extending up frontal longitudinal faces of autozooidal peristomes (
|
||
<figureCitation id="42872A775609553C0866FB0E8BC1FB18" box="[287,395,1158,1183]" captionStart="FIGURE 39" captionStartId="60.[151,250,1940,1962]" captionTargetBox="[166,1420,339,1927]" captionTargetId="figure@60.[166,1420,339,1927]" captionTargetPageId="60" captionText="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437927/files/figure.png" pageId="59" pageNumber="60">Figs 39A</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775609553C08EFFB0F8B82FB18" box="[406,456,1159,1183]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">40A</figureCitation>
|
||
) and abfrontally as flattened alveoli/kenozooids (
|
||
<figureCitation id="42872A775609553C0A8BFB0E8E11FB18" box="[1010,1115,1158,1183]" captionStart="FIGURE 53" captionStartId="78.[151,250,1823,1845]" captionTargetBox="[151,1436,210,1786]" captionTargetId="figure@78.[151,1436,210,1786]" captionTargetPageId="78" captionText="FIGURE 53. Anyuta anastema n. gen., n. sp. Micro-CT scans of paratype, NIWA 127725, as surface (C) and back-face isosurface renders showing colony interiors. A, view of colony from below; note absence of kenozooids from most of column; B, lateral abfrontal view of two small fascicles with entrance to dimorphic zooid indicated by arrowhead; note relative paucity of communication pores; C–E, exterior and interior views of colony base, showing lateral (C, D) and frontolateral (E) profiles with palisade of elongate-triangular kenozooidal chambers surrounding ancestrular protoecium (pr) and proximal part of its peristome (ap). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1437955/files/figure.png" pageId="59" pageNumber="60">Fig. 53A</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="DA0336F25609553C09BEFB228967FAD4" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
|
||
Stalk with subparallel longitudinal ridges separating furrows in which are simple communication pores in generally linear series; ridges flare outward to circular base, giving appearance of pleated skirt (
|
||
<figureCitation id="42872A775609553C0DBAFB468F69FB60" box="[1219,1315,1230,1255]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">Fig. 40J</figureCitation>
|
||
). CT scans reveal
|
||
<emphasis id="E8C8EAE05609553C099DFB7C8ABAFA8C" box="[228,240,1268,1291]" italics="true" pageId="59" pageNumber="60">c</emphasis>
|
||
. 18 highly elongate-triangular kenozooidal chambers in basal part of stalk; these wider basally, acicular distally, surrounding ancestrular protoecium in palisade arrangement (
|
||
<figureCitation id="42872A775609553C0A93FA9E8ECEFAA8" box="[1002,1156,1302,1327]" captionStart="FIGURE 53" captionStartId="78.[151,250,1823,1845]" captionTargetBox="[151,1436,210,1786]" captionTargetId="figure@78.[151,1436,210,1786]" captionTargetPageId="78" captionText="FIGURE 53. Anyuta anastema n. gen., n. sp. Micro-CT scans of paratype, NIWA 127725, as surface (C) and back-face isosurface renders showing colony interiors. A, view of colony from below; note absence of kenozooids from most of column; B, lateral abfrontal view of two small fascicles with entrance to dimorphic zooid indicated by arrowhead; note relative paucity of communication pores; C–E, exterior and interior views of colony base, showing lateral (C, D) and frontolateral (E) profiles with palisade of elongate-triangular kenozooidal chambers surrounding ancestrular protoecium (pr) and proximal part of its peristome (ap). Scale bars: 100 µm." httpUri="https://zenodo.org/record/1437955/files/figure.png" pageId="59" pageNumber="60">Fig. 53D, E</figureCitation>
|
||
); these lacking porous communications with neighbors or exterior colony surface.
|
||
</paragraph>
|
||
<paragraph id="DA0336F25609553C09BEFAD78E47F980" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
|
||
In autozooidal clusters of mature colonies are smaller-diameter tubes with their openings encircled by subhorizontal subcircular flanges (
|
||
<figureCitation id="42872A775609553C0B5BFA0A8890FA1D" box="[546,730,1410,1435]" captionStart="FIGURE 39" captionStartId="60.[151,250,1940,1962]" captionTargetBox="[166,1420,339,1927]" captionTargetId="figure@60.[166,1420,339,1927]" captionTargetPageId="60" captionText="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437927/files/figure.png" pageId="59" pageNumber="60">Figs 39A, B, D</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775609553C0B91FA0B8937FA1C" box="[744,893,1411,1435]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">40A, B, D–I</figureCitation>
|
||
), interpreted as gonozooids. Peristomial rim of smaller tubes somewhat inclined over opening (inferred ooeciopore), slightly reducing its diameter (
|
||
<figureCitation id="42872A775609553C0C5DFA2E8FDAFA38" box="[1316,1424,1446,1471]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">Fig. 40K</figureCitation>
|
||
). Encircling flange white, suggestive of different
|
||
<typeStatus id="050788505609553C0BC0FA4388A3FA64" box="[697,745,1483,1507]" pageId="59" pageNumber="60">type</typeStatus>
|
||
of carbonate from rest of colony; a ring of tiny perforations around inner edge of flange against side of inferred ooeciostome (
|
||
<figureCitation id="42872A775609553C0A01FA668E4AF980" box="[888,1024,1518,1543]" captionStart="FIGURE 40" captionStartId="61.[151,250,1823,1845]" captionTargetBox="[222,1364,193,1809]" captionTargetId="figure@61.[222,1364,193,1809]" captionTargetPageId="61" captionText="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">Fig. 40K, L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F25609553C09BEF99A895BF884" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
|
||
Ancestrular zooid not seen in isolation; earliest growth stage encountered comprising three zooids (
|
||
<figureCitation id="42872A775609553C0C16F99A8AA2F9C8" captionStart="FIGURE 41" captionStartId="62.[151,250,1896,1918]" captionTargetBox="[161,1426,183,1886]" captionTargetId="figure@62.[161,1426,183,1886]" captionTargetPageId="62" captionText="FIGURE 41. Anyuta anastema n. gen., n. sp. Progressive stages of development of young colonies. A–D, specimen YMG4–14, Stn 360, three-zooid stage; E–H, specimen YMG4–14, Stn 338, five–six-zooid stage; I–L, specimen YMG4–13, Stn 281, and M–P, specimen YMG4–06, Stn 71, showing later stages in which trabecular walls define spaces destined to become either autozooids or alveoli. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437931/files/figure.png" pageId="59" pageNumber="60">Fig. 41A–D</figureCitation>
|
||
)—erect ancestrular peristome flanked laterofrontally by pair of daughter zooids budded from its base and concealing it frontally, its abfrontal surface coarsely ridged, with sparse pores. Peristomial tubes of ancestrular zooid and two daughters elongating and increasing in height and wall thickness. Developing colony becoming vasiform (
|
||
<figureCitation id="42872A775609553C0872F92A8B86F93C" box="[267,460,1698,1723]" captionStart="FIGURE 41" captionStartId="62.[151,250,1896,1918]" captionTargetBox="[161,1426,183,1886]" captionTargetId="figure@62.[161,1426,183,1886]" captionTargetPageId="62" captionText="FIGURE 41. Anyuta anastema n. gen., n. sp. Progressive stages of development of young colonies. A–D, specimen YMG4–14, Stn 360, three-zooid stage; E–H, specimen YMG4–14, Stn 338, five–six-zooid stage; I–L, specimen YMG4–13, Stn 281, and M–P, specimen YMG4–06, Stn 71, showing later stages in which trabecular walls define spaces destined to become either autozooids or alveoli. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437931/files/figure.png" pageId="59" pageNumber="60">Fig. 41J–L, N–P</figureCitation>
|
||
); budding of additional zooids from distal abfrontal parts of autozooids, with ~3 walls forming sides of each bud (
|
||
<figureCitation id="42872A775609553C08ABF94E8810F958" box="[466,602,1734,1759]" captionStart="FIGURE 41" captionStartId="62.[151,250,1896,1918]" captionTargetBox="[161,1426,183,1886]" captionTargetId="figure@62.[161,1426,183,1886]" captionTargetPageId="62" captionText="FIGURE 41. Anyuta anastema n. gen., n. sp. Progressive stages of development of young colonies. A–D, specimen YMG4–14, Stn 360, three-zooid stage; E–H, specimen YMG4–14, Stn 338, five–six-zooid stage; I–L, specimen YMG4–13, Stn 281, and M–P, specimen YMG4–06, Stn 71, showing later stages in which trabecular walls define spaces destined to become either autozooids or alveoli. Scale bars: 200 µm." httpUri="https://zenodo.org/record/1437931/files/figure.png" pageId="59" pageNumber="60">
|
||
Fig.
|
||
<date id="AE0210325609553C0B71F94F887AF958" box="[520,560,1735,1759]" pageId="59" pageNumber="60" value="1941-01">41I</date>
|
||
, M
|
||
</figureCitation>
|
||
). Developing colony stem typically with longitudinal ridges and alveolar furrows, these less obvious with secondary calcification.
|
||
</paragraph>
|
||
<paragraph id="DA0336F25609553C09BEF8868E30F830" blockId="59.[151,1437,258,2011]" pageId="59" pageNumber="60">
|
||
<emphasis id="E8C8EAE05609553C09BEF8868B8DF8A0" bold="true" box="[199,455,1806,1831]" pageId="59" pageNumber="60">Measurements (mm).</emphasis>
|
||
<typeStatus id="050788505609553C08A9F8868871F8A0" box="[464,571,1806,1831]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
, ZIRAS 1/50718 (
|
||
<figureCitation id="42872A775609553C0A69F88689DCF8A0" box="[784,918,1806,1831]" captionStart-0="FIGURE 39" captionStart-1="FIGURE 40" captionStartId-0="60.[151,250,1940,1962]" captionStartId-1="61.[151,250,1823,1845]" captionTargetBox-0="[166,1420,339,1927]" captionTargetBox-1="[222,1364,193,1809]" captionTargetId-0="figure@60.[166,1420,339,1927]" captionTargetId-1="figure@61.[222,1364,193,1809]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURE 39. Anyuta anastema n. gen., n. sp. A–D, holotype, ZIRAS 1/50718, in apical and lateral profiles; note the clusters of fascicles, at least three of which have dimorphic orifices with terminal flanges (dimorphic orifices arrowed). Scale bars: 500 µm." captionText-1="FIGURE 40. Anyuta anastema n. gen., n. sp. Holotype, ZIRAS 1/50718. A, part of fascicle with autozooidal peristomes and two dimorphic peristomes with flanges; note large shallow alveoli on sides of peristomes; B, two peristomes, left one dimorphic; C, skeletal microstructure of part of peristome in L; D–G, subfascicles with both autozooidal and flanged dimorphic peristomes; H, close-up of squat flanged dimorphic peristome in E; note small alveoli on flanks of peristome; I, close-up of lower dimorphic peristome in F, showing it to be derived from an autozooidal peristome by its partial closure; J, base of column; K, L, close-ups of perforated flanges of dimorphic peristomes. Scale bars: A, B, D, F, 250 µm; G–J, 100 µm; K, L, 50 µm; C, 25 µm." httpUri-0="https://zenodo.org/record/1437927/files/figure.png" httpUri-1="https://zenodo.org/record/1437929/files/figure.png" pageId="59" pageNumber="60">Figs 39–40</figureCitation>
|
||
): Colony height 4.41; capitulum 3.28 × 2.80; base 1.03 × 0.95; stalk height 2.78; narrowest diameter 0.40 × 0.34; ZL 0.873–1.460 (1.147 ± 0.204); PeL 0.330–0.565 (0.448 ± 0.076); PeD 0.231–0.248 (0.241 ± 0.006); ApL 0.210–0.243 (0.228 ± 0.009); ApW 0.193–0.218 (0.202 ± 0.008). Dimorphic zooids (
|
||
<emphasis id="E8C8EAE05609553C0BABF8F488AAF814" box="[722,736,1916,1939]" italics="true" pageId="59" pageNumber="60">n</emphasis>
|
||
= 4): GZH 0.152–0.211 (0.181 ± 0.024); GZD 0.251–0.325 (0.271 ± 0.036); GZFD 0.181–0.199 (0.188 ± 0.007); OpD 0.091–0.101 (0.094 ± 0.007).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A665795609553B09BEF84A88CAFF52" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="discussion">
|
||
<paragraph id="DA0336F25609553B09BEF84A88CAFF52" blockId="59.[151,1437,258,2011]" lastBlockId="60.[151,1436,151,285]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
|
||
<emphasis id="E8C8EAE05609553C09BEF84A8B71F85C" bold="true" box="[199,315,1986,2011]" pageId="59" pageNumber="60">Remarks.</emphasis>
|
||
<taxonomicName id="1DBC4D715609553C0838F84B8848F85C" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[321,514,1987,2011]" class="Stenolaemata" family="Lichenoporidae" genus="Anyuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="59" pageNumber="60" phylum="Bryozoa" rank="species" species="anastema" status="gen. nov.">
|
||
<emphasis id="E8C8EAE05609553C0838F84B8848F85C" box="[321,514,1987,2011]" italics="true" pageId="59" pageNumber="60">Anyuta anastema</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3FB579B5609553C0B71F84A88E8F85C" box="[520,674,1986,2011]" pageId="59" pageNumber="60" rank="species">
|
||
<emphasis id="E8C8EAE05609553C0B71F84A881CF85C" bold="true" box="[520,598,1986,2011]" pageId="59" pageNumber="60">n. gen.</emphasis>
|
||
,
|
||
<emphasis id="E8C8EAE05609553C0B1DF84A88E8F85C" bold="true" box="[612,674,1986,2011]" pageId="59" pageNumber="60">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is represented in the Russian-sector collection by twelve specimens. The capitulum of the smaller of the two largest colonies is skewed and less symmetrical than in the
|
||
<typeStatus id="05078850560E553B0D91FF1F8F00FF37" box="[1256,1354,151,176]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
|
||
colony and the alveoli are more open and elongate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A66579560E553B09BEFF57884FFE9B" pageId="60" pageNumber="61" type="distribution">
|
||
<paragraph id="DA0336F2560E553B09BEFF57884FFE9B" blockId="60.[151,1436,151,285]" pageId="60" pageNumber="61">
|
||
<emphasis id="E8C8EAE0560E553B09BEFF578B2AFF7F" bold="true" box="[199,352,223,248]" pageId="60" pageNumber="61">Distribution.</emphasis>
|
||
Recorded from 12 stations within coordinates 12.55602–
|
||
<geoCoordinate id="BF885035560E553B0A8FFF578EC3FF70" box="[1014,1161,223,248]" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="1" value="14.32132">14.32132° N</geoCoordinate>
|
||
, 129.11345–
|
||
<geoCoordinate id="BF885035560E553B0C65FF578AF8FE9B" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="1" value="-134.5119">134.51190° W</geoCoordinate>
|
||
, at depth range
|
||
<quantity id="1D449B17560E553B081BFE8C884AFE9B" box="[354,512,260,285]" metricMagnitude="3" metricUnit="m" metricValue="4.9515" metricValueMax="5.212999999999999" metricValueMin="4.6899999999999995" pageId="60" pageNumber="61" unit="m" value="4951.5" valueMax="5213.0" valueMin="4690.0">4690–5213 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |