212 lines
28 KiB
XML
212 lines
28 KiB
XML
<document id="2E9DB6E6217D7E7D03F2A10A4E871A76" ID-DOI="10.11646/zootaxa.3888.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10086415" ID-ZooBank="1B437AE1-CF28-4C1B-95B6-C31A295905A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730057475" checkinUser="felipe" docAuthor="Schwarzhans, Werner" docDate="2014" docId="463A8F36FF8EFFC7129798C9E51BFDC5" docLanguage="en" docName="zt03888p073.pdf.imf" docOrigin="Zootaxa 3888 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hymenocephalus grimaldii Weber 1913" docType="treatment" docVersion="2" lastPageNumber="64" masterDocId="BA03F74EFFB0FFF812009814E16BFFF8" masterDocTitle="Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species" masterLastPageNumber="73" masterPageNumber="1" pageNumber="63" updateTime="1699488604623" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="B50181C2371D71A86E9351352FF96F30">Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species</mods:title>
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<mods:name id="E68D131945BFE86BAE467EFE97634E12" type="personal">
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<mods:namePart id="CB4A363D17870F5E911ABCAE75E915B2">Schwarzhans, Werner</mods:namePart>
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<mods:title id="9A91978A0736EE32455D96EF71C0B555">Zootaxa</mods:title>
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<mods:part id="5D9D9F373A1C35D1A63E959BB1F45C52">
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<mods:date id="AE8CB16B53984E60EC0883718B4DDB20">2014</mods:date>
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<mods:number id="B0AF47285573DCF01E44A95052736CD6">2014-11-28</mods:number>
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<mods:number id="FDE8D46624B05CFC0589E71E9C0E0255">3888</mods:number>
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<taxonomicName id="099345A3FF8EFFC6129798C9E3EBFF0F" ID-CoL="3NJ8W" authority="Weber, 1913" authorityName="Weber" authorityYear="1913" box="[151,640,221,247]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="grimaldii">
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<emphasis id="FCE7E232FF8EFFC6129798C9E3EBFF0F" bold="true" box="[151,640,221,247]" pageId="62" pageNumber="63">
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<emphasis id="FCE7E232FF8EFFC6129798C9E0B4FF0F" bold="true" box="[151,479,221,247]" italics="true" pageId="62" pageNumber="63">Hymenocephalus grimaldii</emphasis>
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,
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TL,
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P.32344-006, 117+ mm TL,
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|
||
<emphasis id="FCE7E232FF8EFFC612979D77E049FA80" bold="true" box="[151,290,1379,1401]" pageId="62" pageNumber="63">FIGURE 34.</emphasis>
|
||
<taxonomicName id="099345A3FF8EFFC6132C9D77E32AFA81" authorityName="Weber" authorityYear="1913" box="[300,577,1379,1401]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="grimaldii">
|
||
<emphasis id="FCE7E232FF8EFFC6132C9D77E32AFA81" box="[300,577,1379,1401]" italics="true" pageId="62" pageNumber="63">Hymenocephalus grimaldii</emphasis>
|
||
</taxonomicName>
|
||
: A—WAM P.32344-006, lateral view of head, B–C—BSKU 098160, B—Lateral view of head, C—Dorsal view of head. D–H—Otoliths, D—Anterior view, E—Ventral view, F–H—Inner faces, D–E—WAM P.32344-006, F—BSKU 17010, G—BSKU 17011.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="86896DABFF8EFFC612C79DF6E3BBF8E3" pageId="62" pageNumber="63" type="diagnosis">
|
||
<paragraph id="CE2C3E20FF8EFFC612C79DF6E402F99F" blockId="62.[151,1437,1506,2035]" pageId="62" pageNumber="63">
|
||
<emphasis id="FCE7E232FF8EFFC612C79DF6E02AFA03" bold="true" box="[199,321,1506,1531]" pageId="62" pageNumber="63">Diagnosis.</emphasis>
|
||
Pelvic fin rays 8; pectoral fin rays 9–11, rarely 12; gill rakers 17–21. Barbel absent. Orbit moderately large, 34–38% HL, strongly oblique. Snout obtuse, short, barely protruding, 15–20% HL. Ventral striae extending to periproct. Preopercular supporter very long and narrow, 8–13% HL. Otolith very compressed with broad predorsal lobe, OL:OH = 0.75–0.8; colliculi fused; pseudocolliculum very long, TCL:PCL = 1.15–1.25.
|
||
</paragraph>
|
||
<paragraph id="CE2C3E20FF8EFFC612C79E66E3BBF8E3" blockId="62.[151,1437,1506,2035]" pageId="62" pageNumber="63">
|
||
<emphasis id="FCE7E232FF8EFFC612C79E66E009F973" bold="true" box="[199,354,1650,1675]" pageId="62" pageNumber="63">Comparison.</emphasis>
|
||
<taxonomicName id="099345A3FF8EFFC613689E66E3F3F973" authorityName="Weber" authorityYear="1913" box="[360,664,1650,1675]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="grimaldii">
|
||
<emphasis id="FCE7E232FF8EFFC613689E66E3F3F973" box="[360,664,1650,1675]" italics="true" pageId="62" pageNumber="63">Hymenocephalus grimaldii</emphasis>
|
||
</taxonomicName>
|
||
is readily recognized by the combination of low counts of pelvic and pectoral fin rays, absence of a barbel, strongly obliquely oriented orbit and very long and narrow preopercular supporter. The otolith morphology adds another set of easily recognizable characters such as the broad, but rather shallow predorsal lobe, the very compressed otolith with the longest pseudocolliculum observed in all
|
||
<taxonomicName id="099345A3FF8EFFC612979F16E0DBF8E3" box="[151,432,1794,1819]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="FCE7E232FF8EFFC612979F16E033F8E3" box="[151,344,1794,1819]" italics="true" pageId="62" pageNumber="63">Hymenocephalus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(TCL-PCL = 1.15–1.25).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="86896DABFF8EFFC712C79F32E4E8FE0D" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="description">
|
||
<paragraph id="CE2C3E20FF8EFFC712C79F32E588FF00" blockId="62.[151,1437,1506,2035]" lastBlockId="63.[151,1437,151,573]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
|
||
<emphasis id="FCE7E232FF8EFFC612C79F32E033F8C7" bold="true" box="[199,344,1830,1855]" pageId="62" pageNumber="63">Description.</emphasis>
|
||
Head morphology (n = 2) (
|
||
<figureCitation id="56A822A5FF8EFFC610AF9F32E254F8C7" box="[687,831,1830,1855]" captionStart="FIGURE 34" captionStartId="62.[151,250,1379,1401]" captionTargetBox="[164,1427,588,1347]" captionTargetId="figure-350@62.[156,1432,582,1354]" captionTargetPageId="62" captionText="FIGURE 34. Hymenocephalus grimaldii: A—WAM P.32344-006, lateral view of head, B–C—BSKU 098160, B—Lateral view of head, C—Dorsal view of head. D–H—Otoliths, D—Anterior view, E—Ventral view, F–H—Inner faces, D–E—WAM P.32344-006, F—BSKU 17010, G—BSKU 17011." figureDoi="http://doi.org/10.5281/zenodo.10086487" httpUri="https://zenodo.org/record/10086487/files/figure.png" pageId="62" pageNumber="63">Fig. 34A–C</figureCitation>
|
||
): Snout short, blunt, high, only slightly projecting, 15–20% HL, orbit diameter large, 34–38% HL, oriented at an angle of 70–80° inclination to long axis of fish, with the longest orbit diameter being 42–45% HL, interorbital width about 80% HW indicating the most advanced lateralization of the eyes. Barbel absent. Head canals well developed, infraorbital width 15–17% HL, supraorbital canal with 4 segments, width 15–18% HL, supratemporal canal above segment 3 of supraorbital canal, preopercular canal width 13–19% HL, postorbital-preopercular interspace 3–5% HL. Infranasal supporter small, thin; infraorbital supporter variable, extended below almost entire length of orbit (70–80% OD) in specimens from
|
||
<collectingCountry id="B6847EB0FF8FFFC7129798A8E06FFF2D" box="[151,260,188,213]" name="Indonesia" pageId="63" pageNumber="64">Indonesia</collectingCountry>
|
||
(
|
||
<figureCitation id="56A822A5FF8FFFC7131598A8E0EBFF2D" box="[277,384,188,213]" captionStart="FIGURE 34" captionStartId="62.[151,250,1379,1401]" captionTargetBox="[164,1427,588,1347]" captionTargetId="figure-350@62.[156,1432,582,1354]" captionTargetPageId="62" captionText="FIGURE 34. Hymenocephalus grimaldii: A—WAM P.32344-006, lateral view of head, B–C—BSKU 098160, B—Lateral view of head, C—Dorsal view of head. D–H—Otoliths, D—Anterior view, E—Ventral view, F–H—Inner faces, D–E—WAM P.32344-006, F—BSKU 17010, G—BSKU 17011." figureDoi="http://doi.org/10.5281/zenodo.10086487" httpUri="https://zenodo.org/record/10086487/files/figure.png" pageId="63" pageNumber="64">Fig. 34A</figureCitation>
|
||
), but below the rear part of the orbit only in the specimen off NW
|
||
<collectingCountry id="B6847EB0FF8FFFC7168A98A8E59AFF2D" box="[1162,1265,188,213]" name="Australia" pageId="63" pageNumber="64">Australia</collectingCountry>
|
||
(30–40% OD) (
|
||
<figureCitation id="56A822A5FF8FFFC7129F98CBE04CFF00" box="[159,295,223,248]" captionStart="FIGURE 34" captionStartId="62.[151,250,1379,1401]" captionTargetBox="[164,1427,588,1347]" captionTargetId="figure-350@62.[156,1432,582,1354]" captionTargetPageId="62" captionText="FIGURE 34. Hymenocephalus grimaldii: A—WAM P.32344-006, lateral view of head, B–C—BSKU 098160, B—Lateral view of head, C—Dorsal view of head. D–H—Otoliths, D—Anterior view, E—Ventral view, F–H—Inner faces, D–E—WAM P.32344-006, F—BSKU 17010, G—BSKU 17011." figureDoi="http://doi.org/10.5281/zenodo.10086487" httpUri="https://zenodo.org/record/10086487/files/figure.png" pageId="63" pageNumber="64">Fig. 34B–C</figureCitation>
|
||
); preopercular supporter very long and narrow, 8–13% HL, with straight rear margin.
|
||
</paragraph>
|
||
<paragraph id="CE2C3E20FF8FFFC712C79910E4E8FE0D" blockId="63.[151,1437,151,573]" pageId="63" pageNumber="64">
|
||
Otolith morphology (n = 8) (
|
||
<figureCitation id="56A822A5FF8FFFC710159910E3CFFEE5" box="[533,676,260,285]" captionStart="FIGURE 34" captionStartId="62.[151,250,1379,1401]" captionTargetBox="[164,1427,588,1347]" captionTargetId="figure-350@62.[156,1432,582,1354]" captionTargetPageId="62" captionText="FIGURE 34. Hymenocephalus grimaldii: A—WAM P.32344-006, lateral view of head, B–C—BSKU 098160, B—Lateral view of head, C—Dorsal view of head. D–H—Otoliths, D—Anterior view, E—Ventral view, F–H—Inner faces, D–E—WAM P.32344-006, F—BSKU 17010, G—BSKU 17011." figureDoi="http://doi.org/10.5281/zenodo.10086487" httpUri="https://zenodo.org/record/10086487/files/figure.png" pageId="63" pageNumber="64">Fig. 34D–H</figureCitation>
|
||
): Otolith large; OL:OH = 0.75–0.8; OH:OT = 3.2–3.5. Dorsal rim with moderately large but very broad, mostly smooth predorsal lobe, distally reaching close to the obtuse, broadly rounded posterior tip without indentation mark; ventral rim very deep, regularly curved, smooth, deepest anterior of the middle; anterior rim high, nearly vertical. Inner face slightly convex, with slightly supramedian sulcus. Colliculi completely fused, terminating far from anterior and posterior tips of otolith, sometimes dorsally reduced and then rather narrow; pseudocolliculum very long. TCL:PCL = 1.15–1.25. Dorsal depression indistinct; no ventral furrow, instead broad, indistinct depression at center of ventral field and parallel to ventral rim of otolith.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="86896DABFF8FFFC712C799EBE51BFDC5" pageId="63" pageNumber="64" type="distribution">
|
||
<paragraph id="CE2C3E20FF8FFFC712C799EBE51BFDC5" blockId="63.[151,1437,151,573]" pageId="63" pageNumber="64">
|
||
<emphasis id="FCE7E232FF8FFFC712C799EBE00BFDE0" bold="true" box="[199,352,511,536]" pageId="63" pageNumber="64">Distribution.</emphasis>
|
||
(
|
||
<figureCitation id="56A822A5FF8FFFC7136F99EBE0ABFDE0" box="[367,448,511,536]" captionStart="FIGURE 37" captionStartId="66.[151,250,976,998]" captionTargetBox="[158,1430,272,949]" captionTargetId="figure-236@66.[152,1435,265,954]" captionTargetPageId="66" captionText="FIGURE 37. Geographical distribution of Hymenocephalus aeger, H. megalops, H. striatissimus, H. torvus, H. grimaldii, H. neglectissimus, H. semipellucidus, H. sp. 1 and H." figureDoi="http://doi.org/10.5281/zenodo.10086493" httpUri="https://zenodo.org/record/10086493/files/figure.png" pageId="63" pageNumber="64">Fig. 37</figureCitation>
|
||
)
|
||
<taxonomicName id="099345A3FF8FFFC713CF99EBE395FDE0" authorityName="Weber" authorityYear="1913" box="[463,766,511,536]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="grimaldii">
|
||
<emphasis id="FCE7E232FF8FFFC713CF99EBE395FDE0" box="[463,766,511,536]" italics="true" pageId="63" pageNumber="64">Hymenocephalus grimaldii</emphasis>
|
||
</taxonomicName>
|
||
has been collected in the Timor and
|
||
<collectingRegion id="0C57F0C2FF8FFFC716989A14E5A1FDEF" box="[1176,1226,512,535]" country="Indonesia" name="Jawa Timur" pageId="63" pageNumber="64">Java</collectingRegion>
|
||
Seas of
|
||
<collectingCountry id="B6847EB0FF8FFFC7172899EBE4F3FDE0" box="[1320,1432,511,536]" name="Indonesia" pageId="63" pageNumber="64">Indonesia</collectingCountry>
|
||
, and
|
||
<specimenCount id="D895F5A9FF8FFFC712C69A30E00BFDC5" box="[198,352,548,573]" pageId="63" pageNumber="64" type="generic">one specimen</specimenCount>
|
||
has now been identified from off Northwest Cape, Western Australia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |