193 lines
33 KiB
XML
193 lines
33 KiB
XML
<document id="9BB6C4D40C61C571D93E69051A187162" ID-CLB-Dataset="54577" ID-DOI="10.6620/ZS.2022.61-13" ID-GBIF-Dataset="854b8390-f85f-4b0a-8c7a-2bfc1da39225" ID-PMC="PMC9579960" ID-PubMed="36330021" ID-Zenodo-Dep="8074520" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687184145743" checkinUser="valdenar" docAuthor="Ng, Peter K L & Grinang, Jongkar" docDate="2022" docId="CF2E433D3C39FFA8FE83FEE3379CFDD9" docLanguage="en" docName="ZoolStud.61.13.1-35.pdf" docOrigin="Zoological studies 61 (13)" docSource="http://dx.doi.org/10.6620/ZS.2022.61-13" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Borneosa niah Ng & Grinang 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="12" masterDocId="33173B453C32FFA3FFA0FF9E341AFFCF" masterDocTitle="Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species." masterLastPageNumber="35" masterPageNumber="1" pageNumber="12" updateTime="1687519204558" updateUser="ExternalLinkService">
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<mods:title id="68CF763B218B48A8EDD1F4E4A2F20749">Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species.</mods:title>
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<mods:namePart id="332725A45CFA0902CD446CEC18C85476">Ng, Peter K L</mods:namePart>
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<mods:roleTerm id="096079576F256C02A4D6CE7479546043">Author</mods:roleTerm>
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<mods:namePart id="8EF9E315EAA004285510A21DC91C20EA">Grinang, Jongkar</mods:namePart>
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<mods:title id="A3708934CC7AAF6AD4CC33E219AC439A">Zoological studies</mods:title>
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<mods:part id="E9C06CA550E0DE13970626E023C80251">
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<mods:date id="243022F2D0951E4FA6A6943B20D11395">2022</mods:date>
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<mods:number id="E056D2405A41ACC0A4D835985297006A">2022-05-05</mods:number>
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<mods:number id="B4B928F167B7236E26A50E10DAE313EE">61</mods:number>
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<mods:number id="1BE0B679B08DA8490F49C9949DA5034F">13</mods:number>
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<mods:url id="74603AE2D2D3BEDFA0E00ACD303E05C0">http://dx.doi.org/10.6620/ZS.2022.61-13</mods:url>
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<mods:identifier id="51C38900AB3610F43108B9D1740108FB" type="DOI">10.6620/ZS.2022.61-13</mods:identifier>
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<treatment id="CF2E433D3C39FFA8FE83FEE3379CFDD9" ID-DOI="http://doi.org/10.5281/zenodo.8055966" ID-Zenodo-Dep="8055966" LSID="urn:lsid:plazi:treatment:CF2E433D3C39FFA8FE83FEE3379CFDD9" httpUri="http://treatment.plazi.org/id/CF2E433D3C39FFA8FE83FEE3379CFDD9" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection id="0F9DA1A03C39FFA8FE83FEE3365CFE59" box="[291,582,381,406]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="4738F22B3C39FFA8FE83FEE3365CFE59" blockId="11.[162,710,381,490]" box="[291,582,381,406]" pageId="11" pageNumber="12">
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<heading id="1C7045473C39FFA8FE83FEE3365CFE59" bold="true" box="[291,582,381,406]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="12" reason="2">
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<emphasis id="75F32E393C39FFA8FE83FEE3365CFE59" bold="true" box="[291,582,381,406]" pageId="11" pageNumber="12">
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<taxonomicName id="808789A83C39FFA8FE83FEE335C6FE59" ID-CoL="9933R" authority="Ng & Grinang, 2022" authorityName="Ng & Grinang" authorityYear="2022" box="[291,476,381,406]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="niah" status="sp. nov.">
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<emphasis id="75F32E393C39FFA8FE83FEE335C6FE59" bold="true" box="[291,476,381,406]" italics="true" pageId="11" pageNumber="12">Borneosa niah</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6EC093423C39FFA8FE43FEE3365CFE59" box="[483,582,381,406]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0F9DA1A03C39FFA8FEECFE03379CFDD9" pageId="11" pageNumber="12" type="description">
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<paragraph id="4738F22B3C39FFA8FEECFE033612FE25" blockId="11.[162,710,381,490]" pageId="11" pageNumber="12">
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(
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<figureCitation id="DFBCEEAE3C39FFA8FEF4FE033585FE7A" box="[340,415,413,437]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Figs. 8</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C39FFA8FE0BFE0335A0FE7A" box="[427,442,413,437]" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C39FFA8FE66FE03360CFE7A" box="[454,534,413,437]" captionStart="Fig" captionStartId="27.[136,167,1926,1945]" captionTargetBox="[147,1405,1034,1897]" captionTargetId="figure-6@27.[136,1413,1023,1905]" captionTargetPageId="27" captionText="Fig. 21. A–D, Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak; E–G, B. bario sp. nov., holotype male (29.6 × 23.1 mm) (ZRC 2021.0692), Bario, Sarawak. Scale bars:A, D, E, H = 1.0 mm; B, C, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074572" httpUri="https://zenodo.org/record/8074572/files/figure.png" pageId="11" pageNumber="12">21A–D</figureCitation>
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)
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<uri id="3316FE293C39FFA8FF02FE263612FE25" pageId="11" pageNumber="12">
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urn:lsid:zoobank.org:act:
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<uuid id="3321C8FE3C39FFA8FE2DFE263612FE25" pageId="11" pageNumber="12">5EB6A04C-E114-4F4E-ACA4- C9D76BC5F8A4</uuid>
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</uri>
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</paragraph>
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<paragraph id="4738F22B3C39FFA8FF65FD8E3526FD66" blockId="11.[136,738,528,1961]" pageId="11" pageNumber="12">
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<emphasis id="75F32E393C39FFA8FF65FD8E3596FDE7" box="[197,396,528,552]" italics="true" pageId="11" pageNumber="12">Material examined</emphasis>
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:
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<materialsCitation id="F7EFF8763C39FFA8FE39FD8F3526FD66" collectingDate="2002-11" collectorName="I. Das" country="Malaysia" latitude="3.8000" location="Malaysia" longitude="113.7666" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<specimenCount id="518139A23C39FFA8FE39FD8F35C5FDE7" box="[409,479,528,553]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
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(22.6 ×
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<quantity id="807F5FCE3C39FFA8FD98FD8F368DFDE6" box="[568,663,529,553]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="11" pageNumber="12" unit="mm" value="19.1">19.1 mm</quantity>
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) (ZRC 2006.064), Bukit Kasut, in river along trail to jetty, in isolated hill some distance from Niah massif, Niah, ca. 3.80°N 113.77'E, Sarawak,
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<collectingCountry id="3F90B2BB3C39FFA8FE48FDEE3655FD47" box="[488,591,624,648]" name="Malaysia" pageId="11" pageNumber="12">Malaysia</collectingCountry>
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, coll.
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<collectorName id="EA7297FD3C39FFA8FD36FDEF36C6FD47" box="[662,732,625,648]" pageId="11" pageNumber="12">I. Das</collectorName>
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,
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<date id="3339D4EB3C39FFA8FF28FD0E3522FD66" box="[136,312,656,681]" pageId="11" pageNumber="12" value="2002-11">
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<collectingDate id="237D2D033C39FFA8FF28FD0E3522FD66" box="[136,312,656,681]" pageId="11" pageNumber="12" value="2002-11">November 2002</collectingDate>
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</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="4738F22B3C39FFA8FF65FD2F35E2FA47" blockId="11.[136,738,528,1961]" pageId="11" pageNumber="12">
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<emphasis id="75F32E393C39FFA8FF65FD2F3534FD07" box="[197,302,689,712]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
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: Adult carapace transversely subquadrate, width to length ratio 1.18 (
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<figureCitation id="DFBCEEAE3C39FFA8FE0DFD4E3639FD26" box="[429,547,720,745]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8A, B</figureCitation>
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); striae on lateral parts of carapace distinct (
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<figureCitation id="DFBCEEAE3C39FFA8FE12FD6E3613FCC6" box="[434,521,752,777]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8B</figureCitation>
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); branchial regions gently convex in frontal view (
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<figureCitation id="DFBCEEAE3C39FFA8FDA0FC8E369AFCE6" box="[512,640,784,809]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8C, F</figureCitation>
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); dorsal margin of frontal median triangle contiguous with lateral margins (
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<figureCitation id="DFBCEEAE3C39FFA8FE9CFCCE35A8FCA6" box="[316,434,848,873]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8C–E</figureCitation>
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); exorbital tooth with outer margin almost entire (
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<figureCitation id="DFBCEEAE3C39FFA8FE32FCEE35EAFC46" box="[402,496,880,905]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8B</figureCitation>
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); epibranchial tooth distinct, separated from rest of margin by low cleft (
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<figureCitation id="DFBCEEAE3C39FFA8FF30FC2E34FFFC06" box="[144,229,944,969]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8B</figureCitation>
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); median lobe of posterior margin of epistome obtusely triangular (
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<figureCitation id="DFBCEEAE3C39FFA8FE21FC4E361EFC26" box="[385,516,976,1001]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8C–E</figureCitation>
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); ischium of third maxilliped subquadrate (
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<figureCitation id="DFBCEEAE3C39FFA8FE33FC6E35F1FBC6" box="[403,491,1008,1033]" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="11" pageNumber="12">Fig. 9A</figureCitation>
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); fingers of adult male cheliped closing along entire cutting margins (
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<figureCitation id="DFBCEEAE3C39FFA8FD25FB8E3486FB86" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="11" pageNumber="12">Fig. 9D, E</figureCitation>
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); P3 and P4 dactyli subequal in length (
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<figureCitation id="DFBCEEAE3C39FFA8FDF8FBAE36CEFB86" box="[600,724,1072,1097]" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="11" pageNumber="12">Fig. 9G, H</figureCitation>
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); male pleonal somite 6 subquadrate, slightly longer than broad (
|
||
<figureCitation id="DFBCEEAE3C39FFA8FF75FBEE3531FB46" box="[213,299,1136,1161]" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="11" pageNumber="12">Fig. 9C</figureCitation>
|
||
); male sternopleonal cavity prominently anterior, distance between tip of cavity and suture of thoracic sternites 2 and 3 distinctly shorter than length of thoracic sternite 2 (
|
||
<figureCitation id="DFBCEEAE3C39FFA8FE27FB4E35FAFB26" box="[391,480,1232,1257]" captionStart="Fig" captionStartId="13.[136,167,1925,1944]" captionTargetBox="[136,1413,730,1899]" captionTargetId="figure-282@13.[136,1413,730,1899]" captionTargetPageId="13" captionText="Fig. 9. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074544" httpUri="https://zenodo.org/record/8074544/files/figure.png" pageId="11" pageNumber="12">Fig. 9B</figureCitation>
|
||
); adult Gl subterminal segment relatively slender; terminal segment subcylindrical, distal half gently curved outwards, tapering gradually to truncate tip, 0.30 times length of subterminal segment, distal opening large, subovate (
|
||
<figureCitation id="DFBCEEAE3C39FFA8FF30FAEE350EFA46" box="[144,276,1392,1417]" captionStart="Fig" captionStartId="27.[136,167,1926,1945]" captionTargetBox="[147,1405,1034,1897]" captionTargetId="figure-6@27.[136,1413,1023,1905]" captionTargetPageId="27" captionText="Fig. 21. A–D, Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak; E–G, B. bario sp. nov., holotype male (29.6 × 23.1 mm) (ZRC 2021.0692), Bario, Sarawak. Scale bars:A, D, E, H = 1.0 mm; B, C, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074572" httpUri="https://zenodo.org/record/8074572/files/figure.png" pageId="11" pageNumber="12">Fig. 21A–C</figureCitation>
|
||
); females not known.
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C39FFA8FF65FA0E3587FA67" blockId="11.[136,738,528,1961]" box="[197,413,1424,1448]" pageId="11" pageNumber="12">
|
||
<emphasis id="75F32E393C39FFA8FF65FA0E350BFA67" box="[197,273,1424,1448]" italics="true" pageId="11" pageNumber="12">Colour</emphasis>
|
||
: Not known.
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C39FFA8FF65FA2E34E5F9C7" blockId="11.[136,738,528,1961]" pageId="11" pageNumber="12">
|
||
<emphasis id="75F32E393C39FFA8FF65FA2E3552FA07" box="[197,328,1456,1480]" italics="true" pageId="11" pageNumber="12">Etymology</emphasis>
|
||
: The species is named after its
|
||
<typeStatus id="983C4C893C39FFA8FF28FA4F34ADFA26" box="[136,183,1489,1513]" pageId="11" pageNumber="12">type</typeStatus>
|
||
locality in Niah. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C39FFA8FF65F98E37D3FE5A" blockId="11.[136,738,528,1961]" lastBlockId="11.[813,1414,253,534]" pageId="11" pageNumber="12">
|
||
<emphasis id="75F32E393C39FFA8FF65F98E353CF9E7" box="[197,294,1552,1576]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
|
||
:
|
||
<taxonomicName id="808789A83C39FFA8FE97F98F35C6F9E7" authority="Ng & Grinang, 2022" authorityName="Ng & Grinang" authorityYear="2022" box="[311,476,1552,1576]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="niah" status="sp. nov.">
|
||
<emphasis id="75F32E393C39FFA8FE97F98F35C6F9E7" box="[311,476,1552,1576]" italics="true" pageId="11" pageNumber="12">Borneosa niah</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EC093423C39FFA8FE46F98F3625F9E7" box="[486,575,1553,1576]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a G1 that is similar to that of
|
||
<taxonomicName id="808789A83C39FFA8FEC5F9AF35F4F987" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[357,494,1584,1608]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="75F32E393C39FFA8FEC5F9AF35F4F987" box="[357,494,1584,1608]" italics="true" pageId="11" pageNumber="12">B. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
although the terminal segment is relatively longer (
|
||
<figureCitation id="DFBCEEAE3C39FFA8FE58F9CE3694F9A6" box="[504,654,1616,1641]" captionStart="Fig" captionStartId="27.[136,167,1926,1945]" captionTargetBox="[147,1405,1034,1897]" captionTargetId="figure-6@27.[136,1413,1023,1905]" captionTargetPageId="27" captionText="Fig. 21. A–D, Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak; E–G, B. bario sp. nov., holotype male (29.6 × 23.1 mm) (ZRC 2021.0692), Bario, Sarawak. Scale bars:A, D, E, H = 1.0 mm; B, C, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074572" httpUri="https://zenodo.org/record/8074572/files/figure.png" pageId="11" pageNumber="12">Fig. 21B, C</figureCitation>
|
||
) when compared to specimens of
|
||
<taxonomicName id="808789A83C39FFA8FE0CF9EF362EF947" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[428,564,1648,1672]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="75F32E393C39FFA8FE0CF9EF362EF947" box="[428,564,1648,1672]" italics="true" pageId="11" pageNumber="12">B. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
of similar sizes (
|
||
<figureCitation id="DFBCEEAE3C39FFA8FF30F90E3515F966" box="[144,271,1680,1705]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng & Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="11" pageNumber="12">Fig. 20F, G</figureCitation>
|
||
). In addition, the frontal median triangle of
|
||
<taxonomicName id="808789A83C39FFA8FF28F92F34C3F907" authorityName="Ng & Grinang" authorityYear="2022" box="[136,217,1712,1736]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="niah">
|
||
<emphasis id="75F32E393C39FFA8FF28F92F34C3F907" box="[136,217,1712,1736]" italics="true" pageId="11" pageNumber="12">B. niah</emphasis>
|
||
</taxonomicName>
|
||
is complete, with the dorsal margin confluent with the lateral margins (
|
||
<figureCitation id="DFBCEEAE3C39FFA8FE12F94E3636F926" box="[434,556,1744,1769]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8C–E</figureCitation>
|
||
) (versus dorsal margin of frontal median triangle separated from lateral margins by a small gap or connected by small granules in
|
||
<taxonomicName id="808789A83C39FFA8FF05F8AF3537F887" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[165,301,1840,1864]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="75F32E393C39FFA8FF05F8AF3537F887" box="[165,301,1840,1864]" italics="true" pageId="11" pageNumber="12">B. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C39FFA8FE9DF8AE35ABF886" box="[317,433,1840,1865]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="11" pageNumber="12">Fig. 2C–E</figureCitation>
|
||
); and the epibranchial tooth is relatively well-developed and strong (
|
||
<figureCitation id="DFBCEEAE3C39FFA8FD94F8CE369CF8A6" box="[564,646,1872,1897]" captionStart="Fig" captionStartId="12.[174,205,1898,1917]" captionTargetBox="[174,1451,434,1872]" captionTargetId="figure-117@12.[174,1451,434,1872]" captionTargetPageId="12" captionText="Fig. 8. Borneosa niah sp. nov., holotype male (22.6 × 19.1 mm) (ZRC 2006.064), Niah, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074542" httpUri="https://zenodo.org/record/8074542/files/figure.png" pageId="11" pageNumber="12">Fig. 8B</figureCitation>
|
||
) (versus low to almost confluent with anterolateral margin in
|
||
<taxonomicName id="808789A83C39FFA8FF08F80F3523F867" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[168,313,1936,1960]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="75F32E393C39FFA8FF08F80F3523F867" box="[168,313,1936,1960]" italics="true" pageId="11" pageNumber="12">B. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="DFBCEEAE3C39FFA8FEEBF80E35ABF866" box="[331,433,1936,1961]" captionStart="Fig" captionStartId="4.[174,205,1897,1916]" captionTargetBox="[174,1451,484,1872]" captionTargetId="figure-151@4.[174,1451,484,1872]" captionTargetPageId="4" captionText="Fig. 2. Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074524" httpUri="https://zenodo.org/record/8074524/files/figure.png" pageId="11" pageNumber="12">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="DFBCEEAE3C39FFA8FE60F80F35F8F866" box="[448,482,1937,1961]" captionStart="Fig" captionStartId="5.[136,167,1869,1888]" captionTargetBox="[136,1413,500,1842]" captionTargetId="figure-175@5.[136,1413,500,1842]" captionTargetPageId="5" captionText="Fig. 3. Borneosa tenebrosa (Holthuis, 1979). A–D, holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; E, paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; F–I, female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, dorsal view of carapace; F–I, left P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074526" httpUri="https://zenodo.org/record/8074526/files/figure.png" pageId="11" pageNumber="12">3E</figureCitation>
|
||
). The two species are adjacent to each other in their distribution, with that of
|
||
<taxonomicName id="808789A83C39FFA8FC8DFE803767FEFA" authorityName="Ng & Grinang" authorityYear="2022" box="[813,893,285,309]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="niah">
|
||
<emphasis id="75F32E393C39FFA8FC8DFE803767FEFA" box="[813,893,285,309]" italics="true" pageId="11" pageNumber="12">B. niah</emphasis>
|
||
</taxonomicName>
|
||
in Niah only just west of the Mulu area where
|
||
<taxonomicName id="808789A83C39FFA8FC8DFEA037DBFE9A" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[813,961,317,341]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="75F32E393C39FFA8FC8DFEA037DBFE9A" box="[813,961,317,341]" italics="true" pageId="11" pageNumber="12">B. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
occurs. Both highlands, however, are separated by lowland rainforests and their drainages are not connected.
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C39FFA8FCC9FE03379CFDD9" blockId="11.[813,1414,253,534]" pageId="11" pageNumber="12">
|
||
<emphasis id="75F32E393C39FFA8FCC9FE0337D4FE7A" box="[873,974,413,437]" italics="true" pageId="11" pageNumber="12">Biology</emphasis>
|
||
: The only known specimen was collected from a stream with rocky substrates in the Niah limestone hills but not from a cave (I. Das, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |