treatments-xml/data/CF/2E/43/CF2E433D3C35FFA8FCDAFEE3361AFE99.xml

<document id="F8CE3E3B18F9EDB9B51960DA686ED688" ID-CLB-Dataset="54577" ID-DOI="10.6620/ZS.2022.61-13" ID-GBIF-Dataset="854b8390-f85f-4b0a-8c7a-2bfc1da39225" ID-PMC="PMC9579960" ID-PubMed="36330021" ID-Zenodo-Dep="8074520" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687184145743" checkinUser="valdenar" docAuthor="Ng, Peter K L &amp; Grinang, Jongkar" docDate="2022" docId="CF2E433D3C35FFA8FCDAFEE3361AFE99" docLanguage="en" docName="ZoolStud.61.13.1-35.pdf" docOrigin="Zoological studies 61 (13)" docSource="http://dx.doi.org/10.6620/ZS.2022.61-13" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Borneosa brachyphallus" docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="33173B453C32FFA3FFA0FF9E341AFFCF" masterDocTitle="Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species." masterLastPageNumber="35" masterPageNumber="1" pageNumber="8" updateTime="1687519204558" updateUser="ExternalLinkService">
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<mods:title id="FFCF1734788259E1FAE29740D295DE28">Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species.</mods:title>
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<mods:namePart id="CFE20E4AB9289E2D2C039C80B91DAE54">Ng, Peter K L</mods:namePart>
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<mods:namePart id="F3EFD73847355E44DA306B4F998EB120">Grinang, Jongkar</mods:namePart>
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<mods:date id="06A0CDFADBC38F4C0A31FD7DB0287748">2022</mods:date>
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<treatment id="CF2E433D3C35FFA8FCDAFEE3361AFE99" ID-DOI="http://doi.org/10.5281/zenodo.8055964" ID-Zenodo-Dep="8055964" LSID="urn:lsid:plazi:treatment:CF2E433D3C35FFA8FCDAFEE3361AFE99" httpUri="http://treatment.plazi.org/id/CF2E433D3C35FFA8FCDAFEE3361AFE99" lastPageId="11" lastPageNumber="11" pageId="7" pageNumber="8">
<subSubSection id="0F9DA1A03C35FFA4FCDAFEE33122FE59" box="[890,1336,381,406]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="4738F22B3C35FFA4FCDAFEE33122FE59" blockId="7.[890,1336,381,437]" box="[890,1336,381,406]" pageId="7" pageNumber="8">
<heading id="1C7045473C35FFA4FCDAFEE33122FE59" bold="true" box="[890,1336,381,406]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="808789A83C35FFA4FCDAFEE33122FE59" ID-CoL="9GDCL" authority="(Ng, 2015)" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[890,1336,381,406]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brachyphallus">
<emphasis id="75F32E393C35FFA4FCDAFEE33122FE59" bold="true" box="[890,1336,381,406]" pageId="7" pageNumber="8">
<emphasis id="75F32E393C35FFA4FCDAFEE330ABFE59" bold="true" box="[890,1201,381,406]" italics="true" pageId="7" pageNumber="8">Borneosa brachyphallus</emphasis>
(
<bibRefCitation id="23168FDA3C35FFA4FB61FEE3312BFE59" author="Ng PKL" box="[1217,1329,381,406]" pageId="7" pageNumber="8" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng, 2015</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C35FFA4FC75FE0330C7FE7A" box="[981,1245,413,437]" pageId="7" pageNumber="8" type="description">
<paragraph id="4738F22B3C35FFA4FC75FE0330C7FE7A" blockId="7.[890,1336,381,437]" box="[981,1245,413,437]" pageId="7" pageNumber="8">
(
<figureCitation id="DFBCEEAE3C35FFA4FC7DFE033032FE7A" box="[989,1064,413,437]" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
, 
<figureCitation id="DFBCEEAE3C35FFA4FB94FE003059FE7A" box="[1076,1091,414,437]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="7" pageNumber="8">7</figureCitation>
, 
<figureCitation id="DFBCEEAE3C35FFA4FBEFFE033067FE7A" box="[1103,1149,413,437]" captionStart="Fig" captionStartId="25.[136,168,1870,1889]" captionTargetBox="[136,1413,729,1841]" captionTargetId="figure-349@25.[136,1413,729,1841]" captionTargetPageId="25" captionText="Fig. 19. Male sternopleonal cavity and pleonal locking tubercle on sternite 5. A, Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; C, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; D, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074568" httpUri="https://zenodo.org/record/8074568/files/figure.png" pageId="7" pageNumber="8">19C</figureCitation>
, 
<figureCitation id="DFBCEEAE3C35FFA4FB29FE0330CFFE7A" box="[1161,1237,413,437]" captionStart="Fig" captionStartId="28.[174,205,1869,1888]" captionTargetBox="[239,1395,251,1840]" captionTargetId="figure-5@28.[227,1398,240,1871]" captionTargetPageId="28" captionText="Fig. 22. A–D, Borneosa kapit sp. nov., paratype male (23.5 × 19.1 mm) (ZRC 2021.0518), Belaga, Sarawak; E–G, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak; H, B. kapit sp. nov., paratype male (35.8 × 28.0 mm) (ZRC 2021.0521), Kapit, Sarawak; I– L, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak; M–P, S. serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. Scale bars: A, D, E, H, I, L, M, P = 1.0 mm; B, C, F, G, J, K, N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074574" httpUri="https://zenodo.org/record/8074574/files/figure.png" pageId="7" pageNumber="8">22Q–T</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C35FFA4FC8DFE463156FE24" box="[813,1356,472,492]" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="4738F22B3C35FFA4FC8DFE463156FE24" blockId="7.[813,1356,472,492]" box="[813,1356,472,492]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="6797D5053C35FFA4FC8DFE463156FE24" box="[813,1356,472,492]" pageId="7" pageNumber="8">
<treatmentCitation id="C626D43A3C35FFA4FC8DFE4630ACFE24" author="Ng PKL" box="[813,1206,472,492]" page="494" pageId="7" pageNumber="8" year="2015">
<taxonomicName id="808789A83C35FFA4FC8DFE4630ACFE24" ID-CoL="93754" authority="Ng, 2015: 494" authorityName="Ng" authorityPageNumber="494" authorityYear="2015" box="[813,1206,472,492]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brachyphallus">
<emphasis id="75F32E393C35FFA4FC8DFE46302EFE24" box="[813,1076,472,491]" italics="true" pageId="7" pageNumber="8">Sundathelphusa brachyphallus</emphasis>
<bibRefCitation id="23168FDA3C35FFA4FB9AFE4730ACFE24" author="Ng PKL" box="[1082,1206,472,492]" pageId="7" pageNumber="8" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng, 2015: 494</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1, 2, 7A–E.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C35FFA8FCC9FD8A361AFE99" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="4738F22B3C35FFA4FCC9FD8A3199FDE4" blockId="7.[813,1413,532,588]" box="[873,1411,532,556]" pageId="7" pageNumber="8">
<emphasis id="75F32E393C35FFA4FCC9FD8A3055FDE3" box="[873,1103,532,556]" italics="true" pageId="7" pageNumber="8">Material examined</emphasis>
: 
<typeStatus id="983C4C893C35FFA4FBC4FD8A30C2FDE3" box="[1124,1240,532,556]" pageId="7" pageNumber="8">Holotype</typeStatus>
: male (27.1 ×
</paragraph>
<paragraph id="4738F22B3C35FFABFC8DFDAA36CCFE39" blockId="7.[813,1413,532,588]" lastBlockId="8.[174,775,253,502]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
22.0 mm) (
<collectionCode id="21966AEE3C35FFA4FC04FDAA37C5FD83" box="[932,991,564,588]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="8" type="Museum">MZB</collectionCode>
Cru 4396), Liang Hintan, Hulu Sungei Barito, 
<geoCoordinate id="22B394EC3C3AFFABFEADFF603590FED9" box="[269,394,254,278]" degrees="0" direction="north" minutes="27" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="33" value="0.45916665">0°27'33&quot;N</geoCoordinate>
<geoCoordinate id="22B394EC3C3AFFABFE36FF633623FEDA" box="[406,569,253,277]" degrees="115" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="1.0" value="115.016945">115°01'1.0&quot;E</geoCoordinate>
, 
<quantity id="807F5FCE3C3AFFABFDEAFF6336C8FEDA" box="[586,722,253,277]" metricMagnitude="2" metricUnit="m" metricValue="2.36" metricValueMax="2.5" metricValueMin="2.2199999999999998" pageId="8" pageNumber="9" unit="m" value="236.0" valueMax="250.0" valueMin="222.0">222–250 m</quantity>
asl, 
<collectingRegion id="85433CC93C3AFFABFF0EFE833597FEFA" box="[174,397,285,309]" country="Indonesia" name="Kalimantan Tengah" pageId="8" pageNumber="9">Kalimantan Tengah</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C3AFFABFE3BFE833610FEFA" box="[411,522,285,309]" name="Indonesia" pageId="8" pageNumber="9">Indonesia</collectingCountry>
, coll. C. Rahmadi, 
<date id="3339D4EB3C3AFFABFD4BFE833504FE99" pageId="8" pageNumber="9" value="2004-06-16">16 June 2004</date>
. 
<typeStatus id="983C4C893C3AFFABFE88FEA03590FE99" box="[296,394,318,342]" pageId="8" pageNumber="9">Paratype</typeStatus>
: 
<specimenCount id="518139A23C3AFFABFE33FEA035C1FE9A" box="[403,475,317,342]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(29.3 × 
<quantity id="807F5FCE3C3AFFABFD96FEA0368DFE9A" box="[566,663,318,342]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="8" pageNumber="9" unit="mm" value="23.4">23.4 mm</quantity>
, chelipeds missing) (
<collectionCode id="21966AEE3C3AFFABFE83FEC03541FEB9" box="[291,347,350,374]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC</collectionCode>
2015.0460), same data as 
<typeStatus id="983C4C893C3AFFABFD3BFEC33718FEBA" box="[667,770,349,373]" pageId="8" pageNumber="9">holotype</typeStatus>
. Others: 
<specimenCount id="518139A23C3AFFABFEAAFEE0354DFE5A" box="[266,343,381,406]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(26.1 × 
<quantity id="807F5FCE3C3AFFABFE1DFEE0363DFE59" box="[445,551,382,406]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="8" pageNumber="9" unit="mm" value="21.8">21.8 mm</quantity>
), 
<specimenCount id="518139A23C3AFFABFD9CFEE036B0FE5A" box="[572,682,381,406]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
(39.8 × 
<quantity id="807F5FCE3C3AFFABFF0EFE00350FFE7A" box="[174,277,414,438]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="8" pageNumber="9" unit="mm" value="31.2">31.2 mm</quantity>
, 35.2 × 
<quantity id="807F5FCE3C3AFFABFED6FE0335C7FE79" box="[374,477,413,438]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="8" pageNumber="9" unit="mm" value="27.6">27.6 mm</quantity>
) (
<collectionCode id="21966AEE3C3AFFABFE55FE003636FE79" box="[501,556,414,438]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="9">ZRC</collectionCode>
2021.0840), Liang Hintan, Tumbang Topus, Sumber Barito, 
<collectingRegion id="85433CC93C3AFFABFD26FE2334E5FE3A" country="Indonesia" name="Kalimantan Tengah" pageId="8" pageNumber="9">Kalimantan Tengah</collectingRegion>
, 
<collectingCountry id="3F90B2BB3C3AFFABFEA9FE433569FE3A" box="[265,371,477,501]" name="Indonesia" pageId="8" pageNumber="9">Indonesia</collectingCountry>
, coll. C. Rahmadi, 
<date id="3339D4EB3C3AFFABFDE0FE4036C8FE39" box="[576,722,478,502]" pageId="8" pageNumber="9" value="2004-06-14">14 June 2004</date>
.
</paragraph>
<caption id="13F8A2A33C35FFA4FF29F8F43581F87A" ID-DOI="http://doi.org/10.5281/zenodo.8074530" ID-Zenodo-Dep="8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="7" pageNumber="8" startId="7.[137,168,1898,1917]" targetBox="[136,1413,661,1871]" targetPageId="7" targetType="figure">
<paragraph id="4738F22B3C35FFA4FF29F8F43581F87A" blockId="7.[136,1414,1898,1974]" pageId="7" pageNumber="8">
<emphasis id="75F32E393C35FFA4FF29F8F434D9F8B2" bold="true" box="[137,195,1898,1917]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
<taxonomicName id="808789A83C35FFA4FF71F8F5361BF8B2" authority="(Ng &amp; Stuebing)" baseAuthorityName="Ng &amp; Stuebing" baseAuthorityYear="1989" box="[209,513,1898,1918]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="aspera">
<emphasis id="75F32E393C35FFA4FF71F8F5357EF8B2" box="[209,356,1899,1917]" italics="true" pageId="7" pageNumber="8">Borneosa aspera</emphasis>
(Ng &amp; Stuebing)
</taxonomicName>
, 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax.
</paragraph>
</caption>
<footnote id="249CEE253C35FFA4FF28F87B3582F838" box="[136,408,2020,2039]" pageId="7" pageNumber="8">
<paragraph id="4738F22B3C35FFA4FF28F87B3582F838" blockId="7.[136,408,2020,2039]" box="[136,408,2020,2039]" pageId="7" pageNumber="8">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4738F22B3C3AFFAAFC2FFF6037F2FE39" blockId="8.[850,1452,253,502]" lastBlockId="9.[813,1414,253,533]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="75F32E393C3AFFABFC2FFF60300EFEDA" box="[911,1044,254,277]" italics="true" pageId="8" pageNumber="9">Diagnosis</emphasis>
: Adult carapace transversely subquadrate, width to length ratio 1.23–1.25 (
<figureCitation id="DFBCEEAE3C3AFFABFAE6FE83377CFE9A" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="8" pageNumber="9">Figs. 6A, B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C3AFFABFCD4FEA03796FE99" box="[884,908,318,342]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="8" pageNumber="9">7I</figureCitation>
); striae on lateral parts of carapace distinct (
<figureCitation id="DFBCEEAE3C3AFFABFAD6FEA3376FFEBA" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="8" pageNumber="9">Figs. 6B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C3AFFABFC25FEC037BAFEB9" box="[901,928,350,374]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="8" pageNumber="9">7I</figureCitation>
); branchial regions gently convex in frontal view (
<figureCitation id="DFBCEEAE3C3AFFABFC3CFEE33006FE59" box="[924,1052,381,406]" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="8" pageNumber="9">Fig. 6C, E</figureCitation>
); dorsal margin of frontal median triangle contiguous with lateral margins (
<figureCitation id="DFBCEEAE3C3AFFABFA86FE033184FE7A" box="[1318,1438,413,437]" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="8" pageNumber="9">Fig. 6C–E</figureCitation>
); exorbital tooth with outer margin almost entire (
<figureCitation id="DFBCEEAE3C3AFFABFAD4FE23376EFE3A" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="8" pageNumber="9">Figs. 6B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C3AFFABFCDFFE403782FE39" box="[895,920,478,502]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="8" pageNumber="9">7I</figureCitation>
); epibranchial tooth very low to absent, if visible, separated from rest of margin by shallow low cleft, anterolateral margin may appear entire (
<figureCitation id="DFBCEEAE3C3BFFAAFDEAFE8336B1FEFA" box="[586,683,285,309]" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="9" pageNumber="10">Figs. 6B</figureCitation>
, 
<figureCitation id="DFBCEEAE3C3BFFAAFD1AFE8036C9FEF9" box="[698,723,286,310]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="9" pageNumber="10">7I</figureCitation>
); median lobe of posterior margin of epistome acutely triangular (
<figureCitation id="DFBCEEAE3C3BFFAAFEABFEC33593FEBA" box="[267,393,349,373]" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="9" pageNumber="10">Fig. 6C–E</figureCitation>
); ischium of third maxilliped subquadrate (
<figureCitation id="DFBCEEAE3C3BFFAAFEBEFEE33563FE59" box="[286,377,381,406]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="9" pageNumber="10">Fig. 7A</figureCitation>
); fingers of adult male cheliped closing along entire cutting margins (
<figureCitation id="DFBCEEAE3C3BFFAAFD86FE033665FE79" box="[550,639,413,438]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="9" pageNumber="10">Fig. 7D</figureCitation>
); P3 and P4 dactyli subequal in length (
<figureCitation id="DFBCEEAE3C3BFFAAFE71FE233658FE19" box="[465,578,445,470]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="9" pageNumber="10">Fig. 7F, G</figureCitation>
); male pleonal somite 6 subquadrate, slightly longer than broad (
<figureCitation id="DFBCEEAE3C3BFFAAFD15FE4334B0FDD9" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="9" pageNumber="10">Fig. 7C</figureCitation>
); male sternopleonal cavity not prominently anterior, distance between tip of cavity and suture of thoracic sternites 2 and 3 longer than length of thoracic sternite 2 (
<figureCitation id="DFBCEEAE3C3BFFAAFC95FEA33792FE99" box="[821,904,317,342]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="9" pageNumber="10">Fig. 7B</figureCitation>
); adult Gl subterminal segment relatively stout; terminal segment subconical, distal half gently curved outwards, tapering gradually to subtruncate tip, 0.35 times length of subterminal segment, distal opening relatively smaller, subovate (
<figureCitation id="DFBCEEAE3C3BFFAAFBFFFE2330C5FE19" box="[1119,1247,445,470]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng &amp; Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="9" pageNumber="10">Fig. 20Q–S</figureCitation>
); vulvae almost round (
<figureCitation id="DFBCEEAE3C3BFFAAFCD9FE4337C1FE39" box="[889,987,477,502]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="9" pageNumber="10">Fig. 23C</figureCitation>
).
</paragraph>
<caption id="13F8A2A33C3AFFABFF0EF8F43084F87A" ID-DOI="http://doi.org/10.5281/zenodo.8074532" ID-Zenodo-Dep="8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="8" pageNumber="9" startId="8.[174,205,1898,1917]" targetBox="[174,1451,574,1872]" targetPageId="8" targetType="figure">
<paragraph id="4738F22B3C3AFFABFF0EF8F43084F87A" blockId="8.[174,1452,1898,1974]" pageId="8" pageNumber="9">
<emphasis id="75F32E393C3AFFABFF0EF8F434F2F8B1" bold="true" box="[174,232,1898,1918]" pageId="8" pageNumber="9">Fig. 5.</emphasis>
<taxonomicName id="808789A83C3AFFABFF57F8F53679F8B1" authority="(Ng &amp; Stuebing, 1989)" baseAuthorityName="Ng &amp; Stuebing" baseAuthorityYear="1989" box="[247,611,1898,1918]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aspera">
<emphasis id="75F32E393C3AFFABFF57F8F53597F8B2" box="[247,397,1899,1917]" italics="true" pageId="8" pageNumber="9">Borneosa aspera</emphasis>
(
<bibRefCitation id="23168FDA3C3AFFABFE3AF8F53647F8B1" author="Ng PKL &amp; Stuebing RB" box="[410,605,1898,1918]" pageId="8" pageNumber="9" pagination="13 - 19" refId="ref19121" refString="Ng PKL, Stuebing RB. 1989. Description of a new species of montane freshwater crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Decapoda: Brachyura: Gecarcinucoidea) from Borneo. Malay Nat J 43 (1): 13 - 19." type="journal article" year="1989">Ng &amp; Stuebing, 1989</bibRefCitation>
)
</taxonomicName>
. A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale.
</paragraph>
</caption>
<footnote id="249CEE253C3AFFABFB3CF87B31B1F838" box="[1180,1451,2020,2039]" pageId="8" pageNumber="9">
<paragraph id="4738F22B3C3AFFABFB3CF87B31B1F838" blockId="8.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="8" pageNumber="9">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4738F22B3C3BFFAAFCC9FE63305BFDDA" blockId="9.[813,1414,253,533]" box="[873,1089,509,533]" pageId="9" pageNumber="10">
<emphasis id="75F32E393C3BFFAAFCC9FE6337AFFDDA" box="[873,949,509,533]" italics="true" pageId="9" pageNumber="10">Colour</emphasis>
: Not known.
</paragraph>
<caption id="13F8A2A33C3BFFAAFF28F8F4359EF87A" ID-DOI="http://doi.org/10.5281/zenodo.8074536" ID-Zenodo-Dep="8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,167,1898,1917]" targetBox="[136,1413,588,1872]" targetPageId="9" targetType="figure">
<paragraph id="4738F22B3C3BFFAAFF28F8F4359EF87A" blockId="9.[136,1414,1898,1974]" pageId="9" pageNumber="10">
<emphasis id="75F32E393C3BFFAAFF28F8F434DAF8B2" bold="true" box="[136,192,1898,1917]" pageId="9" pageNumber="10">Fig. 6.</emphasis>
<taxonomicName id="808789A83C3BFFAAFF6BF8F535E5F8B2" authority="(Ng, 2015)" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[203,511,1898,1918]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brachyphallus">
<emphasis id="75F32E393C3BFFAAFF6BF8F53580F8B2" box="[203,410,1898,1917]" italics="true" pageId="9" pageNumber="10">Borneosa brachyphallus</emphasis>
(
<bibRefCitation id="23168FDA3C3BFFAAFE06F8F535E2F8B2" author="Ng PKL" box="[422,504,1898,1918]" pageId="9" pageNumber="10" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng, 2015</bibRefCitation>
)
</taxonomicName>
, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax.
</paragraph>
</caption>
<footnote id="249CEE253C3BFFAAFF28F87B3582F838" box="[136,408,2020,2039]" pageId="9" pageNumber="10">
<paragraph id="4738F22B3C3BFFAAFF28F87B3582F838" blockId="9.[136,408,2020,2039]" box="[136,408,2020,2039]" pageId="9" pageNumber="10">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4738F22B3C38FFA9FF4AFF633043FEB9" blockId="10.[174,775,253,470]" lastBlockId="10.[850,1452,253,469]" pageId="10" pageNumber="11">
<emphasis id="75F32E393C38FFA9FF4AFF633556FEDA" box="[234,332,253,277]" italics="true" pageId="10" pageNumber="11">Remarks</emphasis>
: Of the various characters 
<bibRefCitation id="23168FDA3C38FFA9FD30FF60371CFEDA" author="Ng PKL" box="[656,774,253,278]" pageId="10" pageNumber="11" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng (2015)</bibRefCitation>
used to distinguish this species from 
<taxonomicName id="808789A83C38FFA9FDECFE8036CDFEFA" baseAuthorityName="Holthuis" baseAuthorityYear="1979" box="[588,727,285,309]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="75F32E393C38FFA9FDECFE8036CDFEFA" box="[588,727,285,309]" italics="true" pageId="10" pageNumber="11">B. tenebrosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="808789A83C38FFA9FF0EFEA0350FFE9A" baseAuthorityName="Ng &amp; Stuebing" baseAuthorityYear="1989" box="[174,277,318,341]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aspera">
<emphasis id="75F32E393C38FFA9FF0EFEA0350FFE9A" box="[174,277,318,341]" italics="true" pageId="10" pageNumber="11">B. aspera</emphasis>
</taxonomicName>
, perhaps the most obvious is that the carapace of 
<taxonomicName id="808789A83C38FFA9FF6EFEC03593FEBA" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[206,393,349,373]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachyphallus">
<emphasis id="75F32E393C38FFA9FF6EFEC03593FEBA" box="[206,393,349,373]" italics="true" pageId="10" pageNumber="11">B. brachyphallus</emphasis>
</taxonomicName>
is much smoother, with the striae near the lateral regions relatively lower and more rounded (
<figureCitation id="DFBCEEAE3C38FFA9FEBCFE0335DDFE7A" box="[284,455,413,437]" captionStart="Fig" captionStartId="9.[136,167,1898,1917]" captionTargetBox="[136,1413,588,1872]" captionTargetId="figure-192@9.[136,1413,588,1872]" captionTargetPageId="9" captionText="Fig. 6. Borneosa brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, different view of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074536" httpUri="https://zenodo.org/record/8074536/files/figure.png" pageId="10" pageNumber="11">Figs. 6A, B, E</figureCitation>
, 
<figureCitation id="DFBCEEAE3C38FFA9FE76FE0035EAFE79" box="[470,496,414,438]" captionStart="Fig" captionStartId="10.[174,205,1898,1917]" captionTargetBox="[174,1451,539,1872]" captionTargetId="figure-190@10.[174,1451,539,1872]" captionTargetPageId="10" captionText="Fig. 7. Borneosa brachyphallus (Ng, 2015). A–H, holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; I, paratype male (29.3 × 23.4 mm, chelipeds missing) (ZRC 2015.0460), Barito, Kalimantan Tengah. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of left chela; E–H, right P2–P5, respectively, all to same scale; I, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.8074540" httpUri="https://zenodo.org/record/8074540/files/figure.png" pageId="10" pageNumber="11">7I</figureCitation>
). The short G1 terminal segment is a character now also shared with 
<taxonomicName id="808789A83C38FFA9FD2AFE2036F8FE1A" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[650,738,445,469]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bario" status="sp. nov.">
<emphasis id="75F32E393C38FFA9FD2AFE2036F8FE1A" box="[650,738,445,469]" italics="true" pageId="10" pageNumber="11">B. bario</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C38FFA9FD48FE203799FEDA" box="[744,899,254,469]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="808789A83C38FFA9FC30FF60305BFEDA" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[912,1089,253,277]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sarawakensis" status="sp. nov.">
<emphasis id="75F32E393C38FFA9FC30FF60305BFEDA" box="[912,1089,253,277]" italics="true" pageId="10" pageNumber="11">B. sarawakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C38FFA9FBEAFF6030BAFEDA" box="[1098,1184,254,277]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="808789A83C38FFA9FB78FF60315CFEDA" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[1240,1350,254,277]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="serrata" status="sp. nov.">
<emphasis id="75F32E393C38FFA9FB78FF60315CFEDA" box="[1240,1350,254,277]" italics="true" pageId="10" pageNumber="11">B. serrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C38FFA9FAEFFF6031BDFEDA" box="[1359,1447,254,277]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, although the subterminal segment of 
<taxonomicName id="808789A83C38FFA9FB50FE8031B6FEFA" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[1264,1452,285,309]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachyphallus">
<emphasis id="75F32E393C38FFA9FB50FE8031B6FEFA" box="[1264,1452,285,309]" italics="true" pageId="10" pageNumber="11">B. brachyphallus</emphasis>
</taxonomicName>
is still proportionately the shortest and stoutest among congeners (
<figureCitation id="DFBCEEAE3C38FFA9FC6DFEC33054FEB9" box="[973,1102,349,374]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng &amp; Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="10" pageNumber="11">Fig. 20Q–S</figureCitation>
).
</paragraph>
<paragraph id="4738F22B3C38FFA8FC2FFEE3361AFE99" blockId="10.[850,1452,253,469]" lastBlockId="11.[136,737,253,342]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="75F32E393C38FFA9FC2FFEE337FFFE5A" box="[911,997,381,405]" italics="true" pageId="10" pageNumber="11">Biology</emphasis>
: 
<taxonomicName id="808789A83C38FFA9FC57FEE03113FE5A" baseAuthorityName="Ng" baseAuthorityYear="2015" box="[1015,1289,381,405]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brachyphallus">
<emphasis id="75F32E393C38FFA9FC57FEE03113FE5A" box="[1015,1289,381,405]" italics="true" pageId="10" pageNumber="11">Borneosa brachyphallus</emphasis>
</taxonomicName>
was collected from inside shallow streams in a horizontal cave in 
<collectingRegion id="85433CC93C38FFA9FCF2FE23303DFE1A" box="[850,1063,445,469]" country="Indonesia" name="Kalimantan Tengah" pageId="10" pageNumber="11">central Kalimantan</collectingRegion>
, in the Barito Basin (
<bibRefCitation id="23168FDA3C38FFA8FAB8FE23355DFED9" author="Rahmadi C &amp; Suhardjono Y." lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" refId="ref19236" refString="Rahmadi C, Suhardjono Y. 2004. Gua-gua di Tumbang Topus, Hulu Sungai Barito, Kalimantan Tengah: tinjauan speleologi dan biologi. Laporan Perjalanan 4 - 29 Juni 2004. Bidang Zoologi, Pusat Penelitan Biologi - LIPI, Cibinong, 28 pp." type="book" year="2004">Rahmadi and Suhardjono 2004</bibRefCitation>
), but it is clearly not a cavernicolous species as it has no the troglobitic adaptations (cf. 
<bibRefCitation id="23168FDA3C39FFA8FF28FEA3353EFE9A" author="Holthuis LB" box="[136,292,317,341]" pageId="11" pageNumber="12" pagination="589 - 615" refId="ref17636" refString="Holthuis LB. 1986. Decapoda. In: Botosaneanu L (ed.) Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna Inhabiting Subterranean Waters (Including the Marine Interstitial). E. J. Brill, Leiden, pp. 589 - 615." type="book chapter" year="1986">Holthuis 1986</bibRefCitation>
; 
<bibRefCitation id="23168FDA3C39FFA8FE8EFEA335AEFE99" author="Guinot D." box="[302,436,317,342]" pageId="11" pageNumber="12" pagination="3 - 40" refId="ref17537" refString="Guinot D. 1988. Les crabes cavernicoles du monde. Mem Biospeol 15: 3 - 40." type="journal article" year="1988">Guinot 1988</bibRefCitation>
<bibRefCitation id="23168FDA3C39FFA8FE1BFEA035EFFE99" author="Guinot D." box="[443,501,318,342]" pageId="11" pageNumber="12" pagination="165 - 179" refId="ref17556" refString="Guinot D. 1994. Decapoda Brachyura. In: Juberthie C, Decou V (eds) Encyclopaedia Biospeologica. Tome 1. Societe de Biospeologie, CNRS, Moulis, Ariege, &amp; Academie Roumaine, Bucarest, pp. 165 - 179, pl. 3." type="journal article" year="1994">1994</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>