281 lines
50 KiB
XML
281 lines
50 KiB
XML
<document id="48ADD95BB30BC29BB39C3BA4B4EB1324" ID-CLB-Dataset="54577" ID-DOI="10.6620/ZS.2022.61-13" ID-GBIF-Dataset="854b8390-f85f-4b0a-8c7a-2bfc1da39225" ID-PMC="PMC9579960" ID-PubMed="36330021" ID-Zenodo-Dep="8074520" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="tatiana" checkinTime="1687184145743" checkinUser="valdenar" docAuthor="Ng, Peter K L & Grinang, Jongkar" docDate="2022" docId="CF2E433D3C34FFA4FC25FDA330A8FE9A" docLanguage="en" docName="ZoolStud.61.13.1-35.pdf" docOrigin="Zoological studies 61 (13)" docSource="http://dx.doi.org/10.6620/ZS.2022.61-13" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Borneosa aspera" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="33173B453C32FFA3FFA0FF9E341AFFCF" masterDocTitle="Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species." masterLastPageNumber="35" masterPageNumber="1" pageNumber="7" updateTime="1716798126574" updateUser="ExternalLinkService">
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<mods:title id="99A359AFF8C03E5044F3BD7F55F51E35">Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species.</mods:title>
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<mods:namePart id="D3FD46E9F3E82214005714EFBF4F5F55">Grinang, Jongkar</mods:namePart>
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<mods:title id="696B3A624423C1F41C34033E8CE39C8D">Zoological studies</mods:title>
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<treatment id="CF2E433D3C34FFA4FC25FDA330A8FE9A" ID-DOI="http://doi.org/10.5281/zenodo.8055962" ID-GBIF-Taxon="210385854" ID-Zenodo-Dep="8055962" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A" httpUri="http://treatment.plazi.org/id/CF2E433D3C34FFA4FC25FDA330A8FE9A" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection id="0F9DA1A03C34FFA5FC25FDA33163FD99" box="[901,1401,572,598]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="4738F22B3C34FFA5FC25FDA33163FD99" blockId="6.[901,1401,572,629]" box="[901,1401,572,598]" pageId="6" pageNumber="7">
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<taxonomicName id="808789A83C34FFA5FC25FDA33163FD99" ID-CoL="99435" authority="(Ng & Stuebing, 1989)" baseAuthorityName="Ng & Stuebing" baseAuthorityYear="1989" box="[901,1401,572,598]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aspera">
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<emphasis id="75F32E393C34FFA5FC25FDA33163FD99" bold="true" box="[901,1401,572,598]" pageId="6" pageNumber="7">
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<emphasis id="75F32E393C34FFA5FC25FDA33044FD99" bold="true" box="[901,1118,573,598]" italics="true" pageId="6" pageNumber="7">Borneosa aspera</emphasis>
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(
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<bibRefCitation id="23168FDA3C34FFA5FBCDFDA3316BFD99" author="Ng PKL & Stuebing RB" box="[1133,1393,572,598]" pageId="6" pageNumber="7" pagination="13 - 19" refId="ref19121" refString="Ng PKL, Stuebing RB. 1989. Description of a new species of montane freshwater crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Decapoda: Brachyura: Gecarcinucoidea) from Borneo. Malay Nat J 43 (1): 13 - 19." type="journal article" year="1989">Ng & Stuebing, 1989</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0F9DA1A03C34FFA5FC18FDC3315CFDBA" box="[952,1350,605,629]" pageId="6" pageNumber="7" type="description">
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<paragraph id="4738F22B3C34FFA5FC18FDC3315CFDBA" blockId="6.[901,1401,572,629]" box="[952,1350,605,629]" pageId="6" pageNumber="7">
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(
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<figureCitation id="DFBCEEAE3C34FFA5FC60FDC33027FDBA" box="[960,1085,605,629]" captionStart="Fig" captionStartId="2.[174,205,1898,1917]" captionTargetBox="[191,1434,1032,1873]" captionTargetId="figure-481@2.[191,1434,1032,1873]" captionTargetPageId="2" captionText="Fig. 1. A, B, frontal margin and median triangle; C, D, male anterior thoracic sternum and sternopleonal cavity.A, C, Sundathelphusa philippina (von Martens, 1868), paralectotype male (51.4 × 42.8 mm) (ZMB 1050), Samar, Philippines; B, D, Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah." figureDoi="http://doi.org/10.5281/zenodo.8074522" httpUri="https://zenodo.org/record/8074522/files/figure.png" pageId="6" pageNumber="7">Figs. 1B, D</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C34FFA5FBE9FDC33042FDBA" box="[1097,1112,605,629]" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">4</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C34FFA5FBC3FDC33068FDBA" box="[1123,1138,605,629]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">5</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C34FFA5FBDEFDC330B7FDBA" box="[1150,1197,605,629]" captionStart="Fig" captionStartId="25.[136,168,1870,1889]" captionTargetBox="[136,1413,729,1841]" captionTargetId="figure-349@25.[136,1413,729,1841]" captionTargetPageId="25" captionText="Fig. 19. Male sternopleonal cavity and pleonal locking tubercle on sternite 5. A, Borneosa tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; C, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Barito, Kalimantan Tengah; D, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074568" httpUri="https://zenodo.org/record/8074568/files/figure.png" pageId="6" pageNumber="7">19B</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C34FFA5FB18FDC3311DFDBA" box="[1208,1287,605,629]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng & Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="6" pageNumber="7">20M–P</figureCitation>
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,
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<figureCitation id="DFBCEEAE3C34FFA5FAB1FDC33125FDBA" box="[1297,1343,605,629]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng & Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="6" pageNumber="7">23B</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0F9DA1A03C34FFA5FCF2FD063157FD2E" pageId="6" pageNumber="7" type="reference_group">
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<paragraph id="4738F22B3C34FFA5FCF2FD063157FD2E" blockId="6.[850,1452,664,737]" pageId="6" pageNumber="7">
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<treatmentCitationGroup id="6797D5053C34FFA5FCF2FD063157FD2E" pageId="6" pageNumber="7">
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<treatmentCitation id="C626D43A3C34FFA5FCF2FD06311FFD64" author="Ng PKL & Stuebing RB" box="[850,1285,664,684]" page="14" pageId="6" pageNumber="7" year="1989">
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<taxonomicName id="808789A83C34FFA5FCF2FD06311FFD64" ID-CoL="53GBC" authority="Ng & Stuebing, 1989: 14" authorityName="Ng & Stuebing" authorityPageNumber="14" authorityYear="1989" box="[850,1285,664,684]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aspera">
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<emphasis id="75F32E393C34FFA5FCF2FD063007FD64" box="[850,1053,664,683]" italics="true" pageId="6" pageNumber="7">Sundathelphusa aspera</emphasis>
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<bibRefCitation id="23168FDA3C34FFA5FB84FD07311FFD64" author="Ng PKL & Stuebing RB" box="[1060,1285,664,684]" pageId="6" pageNumber="7" pagination="13 - 19" refId="ref19121" refString="Ng PKL, Stuebing RB. 1989. Description of a new species of montane freshwater crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Decapoda: Brachyura: Gecarcinucoidea) from Borneo. Malay Nat J 43 (1): 13 - 19." type="journal article" year="1989">Ng & Stuebing, 1989: 14</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs. 1, 2;
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<treatmentCitation id="C626D43A3C34FFA5FACBFD0737E5FD09" author="Ng PKL & Yeo DCJ" page="12" pageId="6" pageNumber="7" year="2007">
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<bibRefCitation id="23168FDA3C34FFA5FACBFD0737E5FD09" author="Ng PKL & Yeo DCJ" pageId="6" pageNumber="7" pagination="95 - 120" refId="ref19171" refString="Ng PKL, Yeo DCJ. 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua L (ed.) Proceedings of the Seminar on the Status of Biological Diversity in Malaysia & Threat Assessment of Plant Species in Malaysia. 28 - 30 June 2005, Forest Research Institute Malaysia, Kepong, pp. 95 - 120." type="book chapter" year="2007">Ng and Yeo 2007: 12</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="C626D43A3C34FFA5FBABFD2D30A2FD09" author="Ng PKL & Guinot D & Davie PJF" box="[1035,1208,691,710]" page="72" pageId="6" pageNumber="7" year="2008">
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<bibRefCitation id="23168FDA3C34FFA5FBABFD2D30A2FD09" author="Ng PKL & Guinot D & Davie PJF" box="[1035,1208,691,710]" pageId="6" pageNumber="7" pagination="1 - 286" refId="ref18968" refString="Ng PKL, Guinot D, Davie PJF. 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull Zool Suppl 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008: 72</bibRefCitation>
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</treatmentCitation>
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;
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<bibRefCitation id="23168FDA3C34FFA5FB64FD2D31B2FD09" author="Cumberlidge N & Ng PKL & Yeo DCJ & Magalhaes C & Campos MR & Alvarez F & Naruse T & Daniels SR & Esser LJ & Attipoe FYK & Clotilde-Ba F-L & Darwall W & Mcivor A & Collen B & Ram M." box="[1220,1448,691,710]" pageId="6" pageNumber="7" pagination="1665 - 1673" refId="ref17167" refString="Cumberlidge N, Ng PKL, Yeo DCJ, Magalhaes C, Campos MR, Alvarez F, Naruse T, Daniels SR, Esser LJ, Attipoe FYK, Clotilde-Ba F-L, Darwall W, Mcivor A, Collen B, Ram M. 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biol Conserv 142: 1665 - 1673. doi: 10.1016 / j. biocon. 2009.02.038." type="journal article" year="2009">Cumberlidge et al. 2009</bibRefCitation>
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: unpaginated appendix;
|
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<treatmentCitation id="C626D43A3C34FFA5FBE7FD5030A4FD2E" author="Ng PKL" box="[1095,1214,717,737]" page="497" pageId="6" pageNumber="7" year="2015">
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<bibRefCitation id="23168FDA3C34FFA5FBE7FD5030A4FD2E" author="Ng PKL" box="[1095,1214,717,737]" pageId="6" pageNumber="7" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng 2015: 497</bibRefCitation>
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||
</treatmentCitation>
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||
, figs. 4–6, 7I–K.
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</treatmentCitationGroup>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="0F9DA1A03C34FFA4FC2FFC9730A8FE9A" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
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<paragraph id="4738F22B3C34FFA5FC2FFC97313DFB0E" blockId="6.[850,1452,777,1953]" pageId="6" pageNumber="7">
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<emphasis id="75F32E393C34FFA5FC2FFC973063FCEE" box="[911,1145,777,801]" italics="true" pageId="6" pageNumber="7">Material examined</emphasis>
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:
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<materialsCitation id="F7EFF8763C34FFA5FB2EFC973096FC6E" ID-GBIF-Occurrence="4134795312" collectingDate="2012-07-06" collectingDateMax="2014-08-07" collectingDateMin="2012-07-06" collectionCode="ZRC" country="Malaysia" county="Sungei Rabergan" elevation="750" latitude="4.95" location="Mendalong" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" pageId="6" pageNumber="7" specimenCode="ZRC 1989.2143" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="holotype">
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<typeStatus id="983C4C893C34FFA5FB2EFC9730E4FCEE" box="[1166,1278,777,801]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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male (42.6 × 32.8 mm) (
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<specimenCode id="17215A503C34FFA5FC6DFCB73098FC8E" box="[973,1154,809,833]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC 1989.2143</specimenCode>
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), clear, swift forest stream,
|
||
<collectingCounty id="AE598AA73C34FFA5FCF2FCD73004FCAE" box="[850,1054,841,865]" pageId="6" pageNumber="7">Sungei Rabergan</collectingCounty>
|
||
, tributary of
|
||
<collectingMunicipality id="A75C68513C34FFA5FB1EFCD73171FCAE" box="[1214,1387,841,865]" pageId="6" pageNumber="7">Sungei Muaya</collectingMunicipality>
|
||
, near
|
||
<location id="4258A4F03C34FFA5FCF2FCF737C3FC4E" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FCF2FCF737C3FC4E" box="[850,985,873,897]" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="Mendalong" pageId="6" pageNumber="7" stateProvince="Temburong">Mendalong</location>
|
||
,
|
||
<geoCoordinate id="22B394EC3C34FFA5FC49FCF73023FC4E" box="[1001,1081,873,897]" degrees="4" direction="north" minutes="57" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="4.95">4°57'N</geoCoordinate>
|
||
<geoCoordinate id="22B394EC3C34FFA5FBE5FCF730A8FC4E" box="[1093,1202,873,897]" degrees="115" direction="east" minutes="43" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="115.71667">115°43'E</geoCoordinate>
|
||
,
|
||
<elevation id="CCAA15183C34FFA5FB62FCF73120FC4E" box="[1218,1338,873,897]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="7" unit="m" value="750.0">
|
||
<quantity id="807F5FCE3C34FFA5FB62FCF73116FC4F" box="[1218,1292,873,897]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="7" unit="m" value="750.0">750 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<location id="4258A4F03C34FFA5FAEAFCF737B5FC6E" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FAEAFCF737B5FC6E" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="Sipitang District" pageId="6" pageNumber="7" stateProvince="Temburong">Sipitang District</location>
|
||
, Sabah,
|
||
<collectingCountry id="3F90B2BB3C34FFA5FBB8FC17309DFC6E" box="[1048,1159,905,929]" name="Malaysia" pageId="6" pageNumber="7">Malaysia</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="F7EFF8763C34FFA5FB37FC1737D8FC0E" ID-GBIF-Occurrence="4867371304" collectingDate="2012-07-06" collectingDateMax="2014-08-07" collectingDateMin="2012-07-06" collectionCode="ZRC" collectorName="Jangorun" country="Malaysia" county="Sungei Rabergan" elevation="750" latitude="4.95" location="Mendalong" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Temburong">
|
||
<location id="4258A4F03C34FFA5FB37FC1730F0FC6E" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FB37FC1730F0FC6E" box="[1175,1258,905,929]" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="Others" pageId="6" pageNumber="7" stateProvince="Temburong">Others</location>
|
||
:
|
||
<specimenCount id="518139A23C34FFA5FB58FC173153FC6E" box="[1272,1353,905,929]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
|
||
(32.3 × 25.4 mm),
|
||
</materialsCitation>
|
||
<materialsCitation id="F7EFF8763C34FFA5FC69FC373037FBEE" ID-GBIF-Occurrence="4867371306" collectingDate="2012-07-06" collectingDateMax="2014-08-07" collectingDateMin="2012-07-06" collectionCode="ZRC" collectorName="Jangorun" country="Malaysia" county="Sungei Rabergan" elevation="750" latitude="4.95" location="Mendalong" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" pageId="6" pageNumber="7" specimenCode="ZRC 2014.845" specimenCount="1" specimenCount-female="1" stateProvince="Temburong">
|
||
<specimenCount id="518139A23C34FFA5FC69FC37303DFC0E" box="[969,1063,937,961]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
|
||
(44.1 × 33.1 mm) (
|
||
<specimenCode id="17215A503C34FFA5FB5BFC3731BAFC0E" box="[1275,1440,937,961]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC 2014.845</specimenCode>
|
||
),
|
||
<collectingRegion id="85433CC93C34FFA5FCF2FC5737C1FC2E" box="[850,987,969,993]" country="Brunei" name="Temburong" pageId="6" pageNumber="7">Temburong</collectingRegion>
|
||
,
|
||
<location id="4258A4F03C34FFA5FC4AFC57303CFC2E" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FC4AFC57303CFC2E" box="[1002,1062,969,993]" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="Amo" pageId="6" pageNumber="7" stateProvince="Temburong">Amo</location>
|
||
,
|
||
<location id="4258A4F03C34FFA5FB95FC57316BFC2F" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FB95FC57316BFC2F" box="[1077,1393,969,993]" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="Batu Apoi Forest Reserve" pageId="6" pageNumber="7" stateProvince="Temburong">Batu Apoi Forest Reserve</location>
|
||
,
|
||
<location id="4258A4F03C34FFA5FA21FC5731B0FC2E" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FA21FC5731B0FC2E" box="[1409,1450,969,993]" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="Ulu" pageId="6" pageNumber="7" stateProvince="Temburong">Ulu</location>
|
||
Temburong
|
||
<location id="4258A4F03C34FFA5FC77FC77306AFBCE" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FC77FC77306AFBCE" box="[983,1136,1001,1025]" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="National Park" pageId="6" pageNumber="7" stateProvince="Temburong">National Park</location>
|
||
,
|
||
<collectingCountry id="3F90B2BB3C34FFA5FBDDFC7730D3FBCE" box="[1149,1225,1001,1025]" name="Brunei" pageId="6" pageNumber="7">Brunei</collectingCountry>
|
||
, coll.
|
||
<collectorName id="EA7297FD3C34FFA5FAAEFC77316AFBCE" box="[1294,1392,1001,1025]" pageId="6" pageNumber="7">Jangorun</collectorName>
|
||
anak
|
||
<location id="4258A4F03C34FFA5FCF2FB97376CFBEE" LSID="urn:lsid:plazi:treatment:CF2E433D3C34FFA4FC25FDA330A8FE9A:4258A4F03C34FFA5FCF2FB97376CFBEE" box="[850,886,1033,1057]" country="Malaysia" county="Sungei Rabergan" latitude="4.95" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" name="Eri" pageId="6" pageNumber="7" stateProvince="Temburong">Eri</location>
|
||
,
|
||
<date id="3339D4EB3C34FFA5FC23FB973033FBEE" box="[899,1065,1033,1057]" pageId="6" pageNumber="7" value="2014-08-07">
|
||
<collectingDate id="237D2D033C34FFA5FC23FB973033FBEE" box="[899,1065,1033,1057]" pageId="6" pageNumber="7" value="2014-08-07">7 August 2014</collectingDate>
|
||
</date>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="F7EFF8763C34FFA5FB96FB9730E0FBAE" ID-GBIF-Occurrence="4134795306" collectingDate="2012-07-06" collectingDateMax="2014-08-07" collectingDateMin="2012-07-06" collectionCode="ZRC" collectorName="S. Salam" country="Malaysia" county="Sungei Rabergan" elevation="750" latitude="4.95" location="Mendalong" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" pageId="6" pageNumber="7" specimenCode="ZRC 2013.746" specimenCount="1" specimenCount-female="1" stateProvince="Temburong">
|
||
<specimenCount id="518139A23C34FFA5FB96FB97308CFBEE" box="[1078,1174,1033,1057]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
|
||
(44.5 × 33.7 mm) (
|
||
<specimenCode id="17215A503C34FFA5FAD5FB9737A4FB8E" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC 2013.746</specimenCode>
|
||
), Bukit Pagon,
|
||
<collectingRegion id="85433CC93C34FFA5FBCEFBB73156FB8E" box="[1134,1356,1065,1089]" country="Brunei" name="Temburong" pageId="6" pageNumber="7">Temburong District</collectingRegion>
|
||
,
|
||
<collectingCountry id="3F90B2BB3C34FFA5FAF9FBB731BCFB8E" box="[1369,1446,1065,1089]" name="Brunei" pageId="6" pageNumber="7">Brunei</collectingCountry>
|
||
,
|
||
<elevation id="CCAA15183C34FFA5FCF2FBD737DAFBAE" box="[850,960,1097,1121]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="7" unit="m" value="900.0">
|
||
<quantity id="807F5FCE3C34FFA5FCF2FBD73782FBAF" box="[850,920,1097,1121]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="7" unit="m" value="900.0">900 m</quantity>
|
||
asl
|
||
</elevation>
|
||
, coll.
|
||
<collectorName id="EA7297FD3C34FFA5FBA4FBD7307CFBAE" box="[1028,1126,1097,1121]" pageId="6" pageNumber="7">S. Salam</collectorName>
|
||
,
|
||
<date id="3339D4EB3C34FFA5FBD3FBD730EDFBAE" box="[1139,1271,1097,1121]" pageId="6" pageNumber="7" value="2012-07-06">
|
||
<collectingDate id="237D2D033C34FFA5FBD3FBD730EDFBAE" box="[1139,1271,1097,1121]" pageId="6" pageNumber="7" value="2012-07-06">6 July 2012</collectingDate>
|
||
</date>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="F7EFF8763C34FFA5FAA3FBD7313DFB0E" ID-GBIF-Occurrence="4134795301" collectingDate="2012-07-06" collectingDateMax="2014-08-07" collectingDateMin="2012-07-06" collectionCode="ZRC" collectorName="H. Mohammod" country="Malaysia" county="Sungei Rabergan" elevation="750" latitude="4.95" location="Mendalong" longLatPrecision="1306" longitude="115.71667" municipality="Sungei Muaya" pageId="6" pageNumber="7" specimenCode="ZRC 2012.1225" specimenCount="2" specimenCount-female="2" stateProvince="Temburong">
|
||
<specimenCount id="518139A23C34FFA5FAA3FBD73176FBAE" box="[1283,1388,1097,1121]" pageId="6" pageNumber="7" type="female">2 females</specimenCount>
|
||
(29.3 × 23.0 mm, 19.7 × 15.6 mm) (
|
||
<specimenCode id="17215A503C34FFA5FB1EFBF73165FB4E" box="[1214,1407,1129,1153]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="7">ZRC 2012.1225</specimenCode>
|
||
), in primary forest stream, in area near Sarawak border,
|
||
<collectingCountry id="3F90B2BB3C34FFA5FCF2FB373787FB0E" box="[850,925,1193,1217]" name="Brunei" pageId="6" pageNumber="7">Brunei</collectingCountry>
|
||
, coll.
|
||
<collectorName id="EA7297FD3C34FFA5FC7EFB373099FB0E" box="[990,1155,1193,1217]" pageId="6" pageNumber="7">H. Mohammod</collectorName>
|
||
,
|
||
<date id="3339D4EB3C34FFA5FB2EFB37313EFB0E" box="[1166,1316,1193,1217]" pageId="6" pageNumber="7" value="2012-10">
|
||
<collectingDate id="237D2D033C34FFA5FB2EFB37313EFB0E" box="[1166,1316,1193,1217]" pageId="6" pageNumber="7" value="2012-10">October 2012</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C34FFA5FC2FFB5431BDF84E" blockId="6.[850,1452,777,1953]" pageId="6" pageNumber="7">
|
||
<emphasis id="75F32E393C34FFA5FC2FFB543019FB2E" box="[911,1027,1226,1249]" italics="true" pageId="6" pageNumber="7">Diagnosis</emphasis>
|
||
: Adult carapace transversely ovate, width to length ratio 1.27–1.30 (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FB7CFB773171FACF" box="[1244,1387,1257,1281]" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">Figs. 4A, B</figureCitation>
|
||
,
|
||
<figureCitation id="DFBCEEAE3C34FFA5FADBFB773186FACE" box="[1403,1436,1257,1281]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">5F</figureCitation>
|
||
); striae on lateral parts of carapace strong (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FAB3FA973169FAEE" box="[1299,1395,1289,1313]" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">Figs. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="DFBCEEAE3C34FFA5FADFFA973184FAEE" box="[1407,1438,1289,1313]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">5F</figureCitation>
|
||
); branchial regions gently convex in frontal view (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FAF5FAB7377EFAAE" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">Fig. 4C, F</figureCitation>
|
||
); dorsal margin of frontal median triangle contiguous with lateral margins (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FBE2FAF730A0FA4E" box="[1090,1210,1385,1409]" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">Fig. 4C–E</figureCitation>
|
||
); exorbital tooth with outer margin almost entire (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FB24FA1730FFFA6E" box="[1156,1253,1417,1441]" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">Figs. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="DFBCEEAE3C34FFA5FB52FA17310BFA6E" box="[1266,1297,1417,1441]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">5F</figureCitation>
|
||
); epibranchial tooth low, separated from rest of margin by shallow low cleft (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FC32FA5737EBFA2E" box="[914,1009,1481,1505]" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">Figs. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="DFBCEEAE3C34FFA5FC5DFA573006FA2E" box="[1021,1052,1481,1505]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">5F</figureCitation>
|
||
); median lobe of posterior margin of epistome obtusely triangular (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FB04FA773104F9CE" box="[1188,1310,1513,1537]" captionStart="Fig" captionStartId="7.[137,168,1898,1917]" captionTargetBox="[136,1413,661,1871]" captionTargetId="figure-203@7.[136,1413,661,1871]" captionTargetPageId="7" captionText="Fig. 4. Borneosa aspera (Ng & Stuebing), 1989, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074530" httpUri="https://zenodo.org/record/8074530/files/figure.png" pageId="6" pageNumber="7">Fig. 4C–E</figureCitation>
|
||
); ischium of third maxilliped subquadrate (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FB39F99730EBF9EE" box="[1177,1265,1545,1569]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
|
||
); fingers of adult male cheliped closing along entire cutting margins (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FA20F9B7378DF9AE" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">Fig. 5D, E</figureCitation>
|
||
); P3 dactylus longer than P4 dactylus (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FAEFF9D73747F94E" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">Fig. 5H, I</figureCitation>
|
||
); male pleonal somite 6 rectangular, longer than broad (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FCFBF91737AEF96E" box="[859,948,1673,1697]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">Fig. 5C</figureCitation>
|
||
); male sternopleonal cavity not prominently anterior, distance between tip of cavity and suture of thoracic sternites 2 and 3 subequal to length of thoracic sternite 2 (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FC67F9773007F8CE" box="[967,1053,1769,1793]" captionStart="Fig" captionStartId="8.[174,205,1898,1917]" captionTargetBox="[174,1451,574,1872]" captionTargetId="figure-232@8.[174,1451,574,1872]" captionTargetPageId="8" captionText="Fig. 5. Borneosa aspera (Ng & Stuebing, 1989). A–C, holotype male (42.6 × 32.8 mm) (ZRC 1989.2143), Sipitang, Sabah; D–J, male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of right chela; E, outer view of left chela; F, dorsal view of carapace; G–J, right P2–P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074532" httpUri="https://zenodo.org/record/8074532/files/figure.png" pageId="6" pageNumber="7">Fig. 5B</figureCitation>
|
||
); adult Gl subterminal segment very slender; terminal segment cylindrical, distal half almost straight, tip wide, truncate, 0.35–0.37 times length of subterminal segment, distal opening very wide, ovate (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FCFAF8F737F9F84E" box="[858,995,1897,1921]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng & Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="6" pageNumber="7">Fig. 20M–O</figureCitation>
|
||
); vulvae transversely subovate (
|
||
<figureCitation id="DFBCEEAE3C34FFA5FA99F8F73181F84E" box="[1337,1435,1897,1921]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng & Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="6" pageNumber="7">Fig. 23B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C34FFA4FC2FF8173674FE99" blockId="6.[850,1452,777,1953]" lastBlockId="7.[136,738,253,598]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis id="75F32E393C34FFA5FC2FF81737C4F86E" box="[911,990,1929,1953]" italics="true" pageId="6" pageNumber="7">Colour</emphasis>
|
||
: In life, the carapace and pereopods of the species are dark to dull brown, with the ventral and frontal surfaces light brown to yellow and pale yellow; the pleon being reddish-brown (
|
||
<figureCitation id="DFBCEEAE3C35FFA4FE7DFEA33678FE99" box="[477,610,317,342]" captionStart="Fig" captionStartId="30.[174,205,1841,1860]" captionTargetBox="[253,1372,253,1815]" captionTargetId="figure-117@30.[253,1372,249,1815]" captionTargetPageId="30" captionText="Fig. 24. Colour in life. A–C, Borneosa aspera (Ng & Stuebing, 1989), female (44.5 × 33.7 mm (ZRC 2013.746), Temburong, Brunei; D, E, B. aspera (Ng & Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; F, G, B. aspera (Ng & Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; H, B. bario sp. nov., holotype male (29.6 × 23.1 mm) (ZRC 2021.0692), Bario, Sarawak; I, B. bario sp. nov., paratype male (31.1 × 23.9 mm) (UNIMAS.C.00011), Bario, Sarawak; J, B. bario sp. nov., Sungei Lawan, tributary of Sungei Tutoh, Baram basin, Miri, specimen not caught. Photographs: A, B, Mohd.Abdul Majid; C–F, Low Yee Wen." figureDoi="http://doi.org/10.5281/zenodo.8074578" httpUri="https://zenodo.org/record/8074578/files/figure.png" pageId="7" pageNumber="8">Fig. 24A–G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<footnote id="249CEE253C34FFA5FB3CF87B31B1F838" box="[1180,1451,2020,2039]" pageId="6" pageNumber="7">
|
||
<paragraph id="4738F22B3C34FFA5FB3CF87B31B1F838" blockId="6.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="6" pageNumber="7">© 2022 Academia Sinica, Taiwan</paragraph>
|
||
</footnote>
|
||
<paragraph id="4738F22B3C35FFA4FF65FEC33033FED9" blockId="7.[136,738,253,598]" lastBlockId="7.[813,1413,253,341]" pageId="7" pageNumber="8">
|
||
<emphasis id="75F32E393C35FFA4FF65FEC33532FEBA" box="[197,296,349,373]" italics="true" pageId="7" pageNumber="8">Remarks</emphasis>
|
||
: Described on the basis of one large male by
|
||
<bibRefCitation id="23168FDA3C35FFA4FF48FEE035EAFE59" author="Ng PKL & Stuebing RB" box="[232,496,381,406]" pageId="7" pageNumber="8" pagination="13 - 19" refId="ref19121" refString="Ng PKL, Stuebing RB. 1989. Description of a new species of montane freshwater crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Decapoda: Brachyura: Gecarcinucoidea) from Borneo. Malay Nat J 43 (1): 13 - 19." type="journal article" year="1989">Ng and Stuebing (1989)</bibRefCitation>
|
||
from northern Sabah, the species was subsequently recorded and redescribed at length on the basis of a larger series of specimens from
|
||
<collectingCountry id="3F90B2BB3C35FFA4FF67FE43350BFE3A" box="[199,273,477,501]" name="Brunei" pageId="7" pageNumber="8">Brunei</collectingCountry>
|
||
(
|
||
<bibRefCitation id="23168FDA3C35FFA4FE82FE403593FE3A" author="Ng PKL" box="[290,393,477,502]" pageId="7" pageNumber="8" pagination="494 - 502" refId="ref18785" refString="Ng PKL. 2015. On a new species of Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from southeastern Borneo. Raffles Bull Zool 63: 494 - 502." type="journal article" year="2015">Ng 2015</bibRefCitation>
|
||
).
|
||
<taxonomicName id="808789A83C35FFA4FE3FFE403641FE3A" baseAuthorityName="Ng & Stuebing" baseAuthorityYear="1989" box="[415,603,478,501]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="aspera">
|
||
<emphasis id="75F32E393C35FFA4FE3FFE403641FE3A" box="[415,603,478,501]" italics="true" pageId="7" pageNumber="8">Borneosa aspera</emphasis>
|
||
</taxonomicName>
|
||
is the most distinctive member of the genus as it reaches the largest sizes (a female, 44.5 ×
|
||
<quantity id="807F5FCE3C35FFA4FE30FD8035ECFDFA" box="[400,502,542,566]" metricMagnitude="-2" metricUnit="m" metricValue="3.37" pageId="7" pageNumber="8" unit="mm" value="33.7">33.7 mm</quantity>
|
||
,
|
||
<collectionCode id="21966AEE3C35FFA4FDA3FD80362DFDF9" box="[515,567,542,566]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
|
||
2013.746) and males also have proportionately, the longest G1 terminal segment (
|
||
<figureCitation id="DFBCEEAE3C35FFA4FC34FF633007FED9" box="[916,1053,253,278]" captionStart="Fig" captionStartId="26.[174,205,1813,1832]" captionTargetBox="[179,1444,490,1784]" captionTargetId="figure-6@26.[174,1451,472,1795]" captionTargetPageId="26" captionText="Fig. 20. A–E, Borneosa tenebrosa (Holthuis, 1979), male (16.1 × 13.0 mm) (ZRC 2013.1293), Gunong Mulu National Park, Sarawak; F–H, B. tenebrosa (Holthuis, 1979), paratype male (21.9 × 17.8 mm) (NHM 1979.136), Gunong Mulu National Park, Sarawak; I–L, B. tenebrosa (Holthuis, 1979), holotype male (23.5 × 19.1 mm) (NHM 1979.135), Gunong Mulu National Park, Sarawak (structures laterally inverted for comparative purposes); M–P, B. aspera (Ng & Stuebing, 1989), male (32.3 × 25.4 mm) (ZRC 2014.845), Temburong, Brunei; Q–T, B. brachyphallus (Ng, 2015), holotype male (27.1 × 22.0 mm) (MZB Cru 4396), Sungei Barito, central Kalimantan. Scale bars: A, D, E, F, G, J, K, N, O, R, S = 0.5 mm; B, C = 0.1 mm; H, I, L, M, P, Q, T = 1.0 mm" figureDoi="http://doi.org/10.5281/zenodo.8074570" httpUri="https://zenodo.org/record/8074570/files/figure.png" pageId="7" pageNumber="8">Fig. 20M–O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4738F22B3C35FFA4FCC9FE8330A8FE9A" blockId="7.[813,1413,253,341]" pageId="7" pageNumber="8">
|
||
<emphasis id="75F32E393C35FFA4FCC9FE8337A5FEFA" box="[873,959,285,309]" italics="true" pageId="7" pageNumber="8">Biology</emphasis>
|
||
: All the specimens were collected from primary highland rainforest streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |