treatments-xml/data/CF/2E/43/CF2E433D3C21FFB5FF48FEE3350BFBBA.xml
2024-06-21 12:52:23 +02:00

283 lines
42 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="12650B272E876DD4655F49D1BDE1BE9E" ID-CLB-Dataset="54577" ID-DOI="10.6620/ZS.2022.61-13" ID-GBIF-Dataset="854b8390-f85f-4b0a-8c7a-2bfc1da39225" ID-PMC="PMC9579960" ID-PubMed="36330021" ID-Zenodo-Dep="8074520" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687184145743" checkinUser="valdenar" docAuthor="Ng, Peter K L &amp; Grinang, Jongkar" docDate="2022" docId="CF2E433D3C21FFB5FF48FEE3350BFBBA" docLanguage="en" docName="ZoolStud.61.13.1-35.pdf" docOrigin="Zoological studies 61 (13)" docSource="http://dx.doi.org/10.6620/ZS.2022.61-13" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Borneosa sarawakensis Ng &amp; Grinang 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="23" masterDocId="33173B453C32FFA3FFA0FF9E341AFFCF" masterDocTitle="Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species." masterLastPageNumber="35" masterPageNumber="1" pageNumber="20" updateTime="1687519204558" updateUser="ExternalLinkService">
<mods:mods id="2D919771881D931C78F456DE49C6AB0C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6A9F59F4855CB902BAD964093FB5236C">
<mods:title id="54787AA1DCEEB88F944503F15A29D3AC">Establishment of A New Bornean Genus of Gecarcinucidae (Crustacea: Brachyura), with Descriptions of Five New Species.</mods:title>
</mods:titleInfo>
<mods:name id="F9920AB4B79D5E1AB7BD29B507A511E2" type="personal">
<mods:role id="2FC15A667CBFB9B04A9C5866981FBC65">
<mods:roleTerm id="62A8631386D6C7538DDDB566F2A4A999">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF151EA358138B1A8B903B53030B4B0F">Ng, Peter K L</mods:namePart>
</mods:name>
<mods:name id="C5DB187378938269CAAFB22985A6FED2" type="personal">
<mods:role id="D1011F24C27578B1CF4720BFD53C4AF3">
<mods:roleTerm id="DE4FC1F3823D6388392422DAE5D7E8BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2962F887C16E6F9A771EE68A511160F">Grinang, Jongkar</mods:namePart>
</mods:name>
<mods:typeOfResource id="41371F7098D388582403D9A58EFC1CF5">text</mods:typeOfResource>
<mods:relatedItem id="23CBAA94B7E2630BBD3FE34064CD6235" type="host">
<mods:titleInfo id="55F25A74E585680E21C514BC98EC958C">
<mods:title id="99A241FCFEEFCE15856596A03204CEFE">Zoological studies</mods:title>
</mods:titleInfo>
<mods:part id="E07EA61EA03C5DA4D2F7F5419D3B7E23">
<mods:date id="971697A8124EB176331E642825B5118A">2022</mods:date>
<mods:detail id="3719E2F6D10059133C1EC5668DFC2589" type="pubDate">
<mods:number id="45C6E5642C4B0865B333D95B95FFDB1C">2022-05-05</mods:number>
</mods:detail>
<mods:detail id="7C3B2BC07941A4ADA42642BD0EB1B09F" type="volume">
<mods:number id="3769380742030C34B806F9E87A87D789">61</mods:number>
</mods:detail>
<mods:detail id="75C5703AE609EB85204661C0205A8B85" type="issue">
<mods:number id="99060310266DD0231FA06FAEE4716072">13</mods:number>
</mods:detail>
<mods:extent id="DA7E82D42C0DEDEF367D3F8A31A9D631" unit="page">
<mods:start id="E0EC11B5AFB5B9C73CCAFB6DBB0EC25F">1</mods:start>
<mods:end id="2ECA3E540EA3910D3F7D36C7E4594944">35</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="30721A6240102309CC1FBB312E2030A5">
<mods:url id="89AEA87CD2A1367B712D4CD518AD5042">http://dx.doi.org/10.6620/ZS.2022.61-13</mods:url>
</mods:location>
<mods:classification id="E27B79BA52E0B2B19E291B6C8ECCC524">journal article</mods:classification>
<mods:identifier id="DB173692BE198904BAA06C1B100B5616" type="CLB-Dataset">54577</mods:identifier>
<mods:identifier id="FFB7D9F8EF4A388D3F8387C7802A28D8" type="DOI">10.6620/ZS.2022.61-13</mods:identifier>
<mods:identifier id="8C5C98EB45FB2B72DA9B2093B7C680F1" type="GBIF-Dataset">854b8390-f85f-4b0a-8c7a-2bfc1da39225</mods:identifier>
<mods:identifier id="91B6CBD4212A1A2727F930A8B07CC23A" type="PMC">PMC9579960</mods:identifier>
<mods:identifier id="9411B2632AA1F670599ACB876D035A73" type="PubMed">36330021</mods:identifier>
<mods:identifier id="0DAF95D5D96C9D1439B37A698C088680" type="Zenodo-Dep">8074520</mods:identifier>
</mods:mods>
<treatment id="CF2E433D3C21FFB5FF48FEE3350BFBBA" ID-DOI="http://doi.org/10.5281/zenodo.8055972" ID-Zenodo-Dep="8055972" LSID="urn:lsid:plazi:treatment:CF2E433D3C21FFB5FF48FEE3350BFBBA" httpUri="http://treatment.plazi.org/id/CF2E433D3C21FFB5FF48FEE3350BFBBA" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20">
<subSubSection id="0F9DA1A03C21FFB0FF48FEE3369BFE59" box="[232,641,381,406]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="4738F22B3C21FFB0FF48FEE3369BFE59" blockId="19.[165,708,381,490]" box="[232,641,381,406]" pageId="19" pageNumber="20">
<heading id="1C7045473C21FFB0FF48FEE3369BFE59" bold="true" box="[232,641,381,406]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="20" reason="2">
<emphasis id="75F32E393C21FFB0FF48FEE3369BFE59" bold="true" box="[232,641,381,406]" pageId="19" pageNumber="20">
<taxonomicName id="808789A83C21FFB0FF48FEE3360DFE59" ID-CoL="993S5" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[232,535,381,406]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="sarawakensis" status="sp. nov.">
<emphasis id="75F32E393C21FFB0FF48FEE3360DFE59" bold="true" box="[232,535,381,406]" italics="true" pageId="19" pageNumber="20">Borneosa sarawakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C21FFB0FDBFFEE3369BFE59" box="[543,641,381,406]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C21FFB0FE89FE03361EFE25" pageId="19" pageNumber="20" type="description">
<paragraph id="4738F22B3C21FFB0FE89FE03361EFE25" blockId="19.[165,708,381,490]" pageId="19" pageNumber="20">
(
<figureCitation id="DFBCEEAE3C21FFB0FE91FE033590FE7A" box="[305,394,413,437]" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="19" pageNumber="20">Figs. 15</figureCitation>
,
<figureCitation id="DFBCEEAE3C21FFB0FE35FE0335A8FE7A" box="[405,434,413,437]" captionStart="Fig" captionStartId="21.[136,168,1926,1945]" captionTargetBox="[136,1413,566,1899]" captionTargetId="figure-238@21.[136,1413,566,1899]" captionTargetPageId="21" captionText="Fig. 16. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of chelae; EH, right P2P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074562" httpUri="https://zenodo.org/record/8074562/files/figure.png" pageId="19" pageNumber="20">16</figureCitation>
,
<figureCitation id="DFBCEEAE3C21FFB0FE1DFE03361BFE7A" box="[445,513,413,437]" captionStart="Fig" captionStartId="28.[174,205,1869,1888]" captionTargetBox="[239,1395,251,1840]" captionTargetId="figure-5@28.[227,1398,240,1871]" captionTargetPageId="28" captionText="Fig. 22. AD, Borneosa kapit sp. nov., paratype male (23.5 × 19.1 mm) (ZRC 2021.0518), Belaga, Sarawak; EG, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak; H, B. kapit sp. nov., paratype male (35.8 × 28.0 mm) (ZRC 2021.0521), Kapit, Sarawak; I L, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak; MP, S. serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. Scale bars: A, D, E, H, I, L, M, P = 1.0 mm; B, C, F, G, J, K, N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074574" httpUri="https://zenodo.org/record/8074574/files/figure.png" pageId="19" pageNumber="20">22IL</figureCitation>
,
<figureCitation id="DFBCEEAE3C21FFB0FDACFE033622FE7A" box="[524,568,413,437]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="19" pageNumber="20">23E</figureCitation>
)
<uri id="3316FE293C21FFB0FF05FE26361EFE25" pageId="19" pageNumber="20">
urn:lsid:zoobank.org:act:
<uuid id="3321C8FE3C21FFB0FE33FE26361EFE25" pageId="19" pageNumber="20">1A80E5BF-EB74-42B0-AF74- 98E5023A91FA</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C21FFB6FF65FD8E36DAFEF9" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="4738F22B3C21FFB6FF65FD8E36DAFEF9" blockId="19.[136,737,528,585]" lastBlockId="21.[136,737,253,501]" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<emphasis id="75F32E393C21FFB0FF65FD8E35B0FDE7" box="[197,426,528,552]" italics="true" pageId="19" pageNumber="20">Material examined</emphasis>
:
<materialsCitation id="F7EFF8763C21FFB0FE60FD8E311EFE9A" collectingDate="2015-01-30" collectionCode="ZRC" collectorName="J. Grinang" country="Malaysia" location="Malaysia" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="983C4C893C21FFB0FE60FD8E362EFDE7" box="[448,564,528,552]" pageId="19" pageNumber="20">Holotype</typeStatus>
:
<specimenCount id="518139A23C21FFB0FDE3FD8E3666FDE7" box="[579,636,528,552]" pageId="19" pageNumber="20" type="male">male</specimenCount>
(35.5 ×
<quantity id="807F5FCE3C21FFB0FF28FDAF34F0FD86" box="[136,234,561,585]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="19" pageNumber="20" unit="mm" value="28.1">28.1 mm</quantity>
), 1 female (11.8 ×
<quantity id="807F5FCE3C21FFB0FE10FDAF360AFD86" box="[432,528,561,585]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="19" pageNumber="20" unit="mm" value="10.3">10.3 mm</quantity>
) (
<collectionCode id="21966AEE3C21FFB0FD85FDAF3640FD86" box="[549,602,561,585]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="19" pageNumber="20">ZRC</collectionCode>
2021.0516), first order stream, primary forest, Sungei Musing, Batang Ai National Park, Sri Aman, Sarawak,
<collectingCountry id="3F90B2BB3C21FFB0FABDFE83319BFEFA" box="[1309,1409,285,309]" name="Malaysia" pageId="19" pageNumber="20">Malaysia</collectingCountry>
, coll.
<collectorName id="EA7297FD3C21FFB0FCCAFEA037F9FE9A" box="[874,995,317,341]" pageId="19" pageNumber="20">J. Grinang</collectorName>
et al.,
<date id="3339D4EB3C21FFB0FB9BFEA0311AFE9A" box="[1083,1280,317,342]" pageId="19" pageNumber="20" value="2015-01-30">
<collectingDate id="237D2D033C21FFB0FB9BFEA0311AFE9A" box="[1083,1280,317,342]" pageId="19" pageNumber="20" value="2015-01-30">30 January 2015</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="F7EFF8763C21FFB0FAAEFEA037EEFE5A" pageId="19" pageNumber="20">
<typeStatus id="983C4C893C21FFB0FAAEFEA0319BFE99" box="[1294,1409,318,342]" pageId="19" pageNumber="20">Paratypes</typeStatus>
:
<specimenCount id="518139A23C21FFB0FC8DFEC0378AFEBA" box="[813,912,349,374]" pageId="19" pageNumber="20" type="female">1 female</specimenCount>
(11.8 ×
<quantity id="807F5FCE3C21FFB0FC58FEC03045FEB9" box="[1016,1119,350,374]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="19" pageNumber="20" unit="mm" value="10.3">10.3 mm</quantity>
) (ZRC 2021.0517), same data as
<typeStatus id="983C4C893C21FFB0FC28FEE337EAFE5A" box="[904,1008,381,405]" pageId="19" pageNumber="20">holotype</typeStatus>
.
</materialsCitation>
<materialsCitation id="F7EFF8763C21FFB7FC5EFEE33083FED9" lastPageId="20" lastPageNumber="21" latitude="1.6833167" longitude="112.16513" pageId="19" pageNumber="20">
Others:
<specimenCount id="518139A23C21FFB0FBFDFEE030B6FE5A" box="[1117,1196,381,406]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
(30.8 ×
<quantity id="807F5FCE3C21FFB0FAB5FEE33164FE59" box="[1301,1406,381,406]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="19" pageNumber="20" unit="mm" value="24.6">24.6 mm</quantity>
) (ZRC 2020.0338), station THH19-34, in fast flowing water over bedrock, rocks and gravel, feeder stream to Sungei Mujok, survey up to ca.
<quantity id="807F5FCE3C21FFB0FB25FE4030D3FE3A" box="[1157,1225,478,502]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="m" value="800.0">800 m</quantity>
upstream, Nanga Sepulau, Kanowit basin in Rejang Basin,
<geoCoordinate id="22B394EC3C21FFB0FAA8FE60319FFDD9" box="[1288,1413,510,534]" degrees="1" direction="north" minutes="40.999" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="1.6833167">1°40.999'N</geoCoordinate>
<geoCoordinate id="22B394EC3C21FFB0FC8DFD8037DBFDF9" box="[813,961,542,566]" degrees="112" direction="east" minutes="09.908" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="112.16513">112°09.908'E</geoCoordinate>
,
<collectingCountry id="3F90B2BB3C21FFB0FC6FFD833035FDFA" box="[975,1071,541,565]" name="Malaysia" pageId="19" pageNumber="20">Sarawak</collectingCountry>
,
<collectingCountry id="3F90B2BB3C21FFB0FB9DFD8330B9FDFA" box="[1085,1187,541,565]" name="Malaysia" pageId="19" pageNumber="20">Malaysia</collectingCountry>
, electrofishing, coll. H.H. Tan et al.,
<date id="3339D4EB3C21FFB0FC49FDA330B7FD99" box="[1001,1197,573,598]" pageId="19" pageNumber="20" value="2019-10-15">15 October 2019</date>
;
<specimenCount id="518139A23C21FFB0FB1BFDA03132FD9A" box="[1211,1320,573,598]" pageId="19" pageNumber="20" type="female">3 females</specimenCount>
(24.6 ×
<quantity id="807F5FCE3C26FFB7FF0EFF603508FEDA" box="[174,274,254,278]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="20" pageNumber="21" unit="mm" value="20.9">20.9 mm</quantity>
, 25.3 ×
<quantity id="807F5FCE3C26FFB7FED0FF6035CFFEDA" box="[368,469,254,278]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="20" pageNumber="21" unit="mm" value="21.2">21.2 mm</quantity>
, 27.2 ×
<quantity id="807F5FCE3C26FFB7FD93FF60368FFEDA" box="[563,661,254,278]" metricMagnitude="-2" metricUnit="m" metricValue="2.27" pageId="20" pageNumber="21" unit="mm" value="22.7">22.7 mm</quantity>
[with
<specimenCount id="518139A23C26FFB7FD40FF60351CFEFA" pageId="20" pageNumber="21" type="juvenile">one juvenile</specimenCount>
crab under pleon]) (ZRC 2020.0336), station DZ-01, along first small stream on right side of Sungei Mujok, upstream from base camp, at night, Lanjak-Entimau National Park,
<collectingCountry id="3F90B2BB3C26FFB7FE1AFEE33600FE5A" box="[442,538,381,405]" name="Malaysia" pageId="20" pageNumber="21">Sarawak</collectingCountry>
,
<collectingCountry id="3F90B2BB3C26FFB7FD87FEE33697FE5A" box="[551,653,381,405]" name="Malaysia" pageId="20" pageNumber="21">Malaysia</collectingCountry>
, coll. K.O. Chan et al.,
<date id="3339D4EB3C26FFB7FC7AFF60308FFED9" box="[986,1173,253,278]" pageId="20" pageNumber="21" value="2019-10-12">12 October 2019</date>
;
</materialsCitation>
<materialsCitation id="F7EFF8763C26FFB6FB02FF6036DAFEF9" lastPageId="21" lastPageNumber="22" latitude="1.68195" longitude="112.169365" pageId="20" pageNumber="21">
1 brooding female (30.0 × 24.0 mm) (ZRC 2020.0337), station THH19-33, in clear fast flowing water over bedrock, rocks, gravel, upstream from Nanga Sepulau (ca.
<quantity id="807F5FCE3C26FFB7FB53FEC0312AFEBA" box="[1267,1328,349,374]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="km" value="1.0">1 km</quantity>
) to Nanga Sepuna, Kanowit basin in Rejang Basin,
<geoCoordinate id="22B394EC3C26FFB7FA8CFEE031B1FE59" box="[1324,1451,382,406]" degrees="1" direction="north" minutes="40.917" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="1.68195">1°40.917'N</geoCoordinate>
<geoCoordinate id="22B394EC3C27FFB6FF28FF60352EFEDA" box="[136,308,253,278]" degrees="112" direction="east" minutes="10. 162" orientation="longitude" pageId="21" pageNumber="22" precision="1" value="112.169365">112 °10.162 ' E</geoCoordinate>
,
<quantity id="807F5FCE3C27FFB6FEEBFF603581FEDA" box="[331,411,254,278]" metricMagnitude="2" metricUnit="m" metricValue="1.04" pageId="21" pageNumber="22" unit="m" value="104.0">104 m</quantity>
asl,
<collectingCountry id="3F90B2BB3C27FFB6FE42FF603657FEDA" box="[482,589,253,278]" name="Malaysia" pageId="21" pageNumber="22">Sarawak</collectingCountry>
,
<collectingCountry id="3F90B2BB3C27FFB6FDC4FF6036C2FEDA" box="[612,728,253,278]" name="Malaysia" pageId="21" pageNumber="22">Malaysia</collectingCountry>
, electrofishing, coll. H.H. Tan et al.,
<date id="3339D4EB3C27FFB6FDA5FE8036A6FEF9" box="[517,700,285,310]" pageId="21" pageNumber="22" value="2019-10-14">14 October 2019</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="13F8A2A33C21FFB0FF28F8D0377BF87A" ID-DOI="http://doi.org/10.5281/zenodo.8074558" ID-Zenodo-Dep="8074558" httpUri="https://zenodo.org/record/8074558/files/figure.png" pageId="19" pageNumber="20" startId="19.[136,167,1870,1889]" targetBox="[136,1413,633,1844]" targetPageId="19" targetType="figure">
<paragraph id="4738F22B3C21FFB0FF28F8D0377BF87A" blockId="19.[136,1414,1870,1973]" pageId="19" pageNumber="20">
<emphasis id="75F32E393C21FFB0FF28F8D034D6F8AE" bold="true" box="[136,204,1870,1889]" pageId="19" pageNumber="20">Fig. 14.</emphasis>
<taxonomicName id="808789A83C21FFB0FF78F8D13541F8AE" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[216,347,1870,1889]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kapit" status="sp. nov.">
<emphasis id="75F32E393C21FFB0FF78F8D13541F8AE" box="[216,347,1870,1889]" italics="true" pageId="19" pageNumber="20">Borneosa kapit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C21FFB0FEC1F8D035BDF8AE" box="[353,423,1870,1889]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, overall dorsal view. A, paratype male (35.8 × 28.6 mm) (UNIMAS.C.00102), Sungei Yong, Sarawak; B, paratype male (36.3 × 27.6 mm) (UNIMAS.C.00103), Benin, Kapit, Sarawak; C, paratype female (27.8 × 22.3 mm) (UNIMAS.C.00103), Benin, Kapit, Sarawak; D, paratype male (12.8 × 9.9 mm) (UNIMAS.C.00039), Belaga, Sarawak; E, paratype male (20.6 × 16.9 mm) (UNIMAS.C.00039), Belaga, Sarawak; F, paratype female (12.4 × 10.9 mm) (UNIMAS.C.00101), Kapit, Sarawak.
</paragraph>
</caption>
<footnote id="249CEE253C21FFB0FF28F87B3582F838" box="[136,408,2020,2039]" pageId="19" pageNumber="20">
<paragraph id="4738F22B3C21FFB0FF28F87B3582F838" blockId="19.[136,408,2020,2039]" box="[136,408,2020,2039]" pageId="19" pageNumber="20">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="13F8A2A33C26FFB7FF0EF8F635B3F87C" ID-DOI="http://doi.org/10.5281/zenodo.8074560" ID-Zenodo-Dep="8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="20" pageNumber="21" startId="20.[174,205,1896,1915]" targetBox="[184,1441,456,1871]" targetPageId="20" targetType="figure">
<paragraph id="4738F22B3C26FFB7FF0EF8F635B3F87C" blockId="20.[174,1451,1896,1972]" pageId="20" pageNumber="21">
<emphasis id="75F32E393C26FFB7FF0EF8F634EBF8B4" bold="true" box="[174,241,1896,1915]" pageId="20" pageNumber="21">Fig. 15.</emphasis>
<taxonomicName id="808789A83C26FFB7FF5DF8F735D2F8B4" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[253,456,1896,1915]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sarawakensis" status="sp. nov.">
<emphasis id="75F32E393C26FFB7FF5DF8F735D2F8B4" box="[253,456,1896,1915]" italics="true" pageId="20" pageNumber="21">Borneosa sarawakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C26FFB7FE6EF8F73609F8B3" box="[462,531,1897,1916]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax.
</paragraph>
</caption>
<footnote id="249CEE253C26FFB7FB3CF87B31B1F838" box="[1180,1451,2020,2039]" pageId="20" pageNumber="21">
<paragraph id="4738F22B3C26FFB7FB3CF87B31B1F838" blockId="20.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="20" pageNumber="21">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<subSubSection id="0F9DA1A03C27FFB5FF65FEA03569FD1A" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="4738F22B3C27FFB5FF65FEA03618FD99" blockId="21.[136,737,253,501]" lastBlockId="22.[174,775,253,1141]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="75F32E393C27FFB6FF65FEA03550FE9A" box="[197,330,318,341]" italics="true" pageId="21" pageNumber="22">Diagnosis</emphasis>
: Adult carapace transversely subquadrate, width to length ratio 1.181.26 (
<figureCitation id="DFBCEEAE3C27FFB6FD12FEC334FBFE59" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15A, B</figureCitation>
); striae on lateral parts of carapace strong (
<figureCitation id="DFBCEEAE3C27FFB6FF30FE0334ECFE7A" box="[144,246,413,437]" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15B</figureCitation>
); branchial regions gently convex in frontal view (
<figureCitation id="DFBCEEAE3C27FFB6FF70FE233540FE19" box="[208,346,445,470]" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15C, F</figureCitation>
); dorsal margin of frontal median triangle contiguous with lateral margins (
<figureCitation id="DFBCEEAE3C27FFB6FDF1FE4336CEFE3A" box="[593,724,477,501]" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15CE</figureCitation>
); exorbital tooth with outer margin almost entire (
<figureCitation id="DFBCEEAE3C27FFB6FAF8FF633747FEFA" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15B</figureCitation>
); epibranchial tooth very low to absent, separated from rest of margin by very low cleft when present (
<figureCitation id="DFBCEEAE3C27FFB6FAFAFEA33747FEBA" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15B</figureCitation>
), anterolateral margin almost entire (
<figureCitation id="DFBCEEAE3C27FFB6FAAEFEC3316DFEBA" box="[1294,1399,349,373]" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15B</figureCitation>
); median lobe of posterior margin of epistome obtusely triangular (
<figureCitation id="DFBCEEAE3C27FFB6FC0CFE03302FFE7A" box="[940,1077,413,437]" captionStart="Fig" captionStartId="20.[174,205,1896,1915]" captionTargetBox="[184,1441,456,1871]" captionTargetId="figure-140@20.[184,1441,456,1871]" captionTargetPageId="20" captionText="Fig. 15. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, frontal median triangle, eyes, epistome, antennae and antennules; E, F, different views of frontal part of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8074560" httpUri="https://zenodo.org/record/8074560/files/figure.png" pageId="21" pageNumber="22">Fig. 15CE</figureCitation>
); ischium of third maxilliped rectangular (
<figureCitation id="DFBCEEAE3C27FFB6FC60FE233036FE1A" box="[960,1068,445,469]" captionStart="Fig" captionStartId="21.[136,168,1926,1945]" captionTargetBox="[136,1413,566,1899]" captionTargetId="figure-238@21.[136,1413,566,1899]" captionTargetPageId="21" captionText="Fig. 16. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of chelae; EH, right P2P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074562" httpUri="https://zenodo.org/record/8074562/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
); fingers of larger adult male cheliped with gape at base of when closed (
<figureCitation id="DFBCEEAE3C27FFB6FAB2FE433163FE3A" box="[1298,1401,477,501]" captionStart="Fig" captionStartId="21.[136,168,1926,1945]" captionTargetBox="[136,1413,566,1899]" captionTargetId="figure-238@21.[136,1413,566,1899]" captionTargetPageId="21" captionText="Fig. 16. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of chelae; EH, right P2P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074562" httpUri="https://zenodo.org/record/8074562/files/figure.png" pageId="21" pageNumber="22">Fig. 16D</figureCitation>
); P3 and P4 dactyli subequal in length (
<figureCitation id="DFBCEEAE3C24FFB5FDCFFF6336E3FED9" box="[623,761,253,278]" captionStart="Fig" captionStartId="21.[136,168,1926,1945]" captionTargetBox="[136,1413,566,1899]" captionTargetId="figure-238@21.[136,1413,566,1899]" captionTargetPageId="21" captionText="Fig. 16. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of chelae; EH, right P2P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074562" httpUri="https://zenodo.org/record/8074562/files/figure.png" pageId="22" pageNumber="23">Fig. 16F, G</figureCitation>
); male pleonal somite 6 rectangular, longer than broad (
<figureCitation id="DFBCEEAE3C24FFB5FF16FEA33506FE9A" box="[182,284,317,341]" captionStart="Fig" captionStartId="21.[136,168,1926,1945]" captionTargetBox="[136,1413,566,1899]" captionTargetId="figure-238@21.[136,1413,566,1899]" captionTargetPageId="21" captionText="Fig. 16. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of chelae; EH, right P2P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074562" httpUri="https://zenodo.org/record/8074562/files/figure.png" pageId="22" pageNumber="23">Fig. 16C</figureCitation>
); male sternopleonal cavity not prominently anterior, distance between tip of cavity and suture of thoracic sternites 2 and 3 longer than length of thoracic sternite 2 (
<figureCitation id="DFBCEEAE3C24FFB5FE8DFE03358DFE7A" box="[301,407,413,437]" captionStart="Fig" captionStartId="21.[136,168,1926,1945]" captionTargetBox="[136,1413,566,1899]" captionTargetId="figure-238@21.[136,1413,566,1899]" captionTargetPageId="21" captionText="Fig. 16. Borneosa sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of chelae; EH, right P2P5, respectively, all to same scale." figureDoi="http://doi.org/10.5281/zenodo.8074562" httpUri="https://zenodo.org/record/8074562/files/figure.png" pageId="22" pageNumber="23">Fig. 16B</figureCitation>
); adult Gl subterminal segment relatively slender; terminal segment subcylindrical, distal half distinctly curved outwards, with distinct rounded tip, 0.300.32 times length of subterminal segment, distal opening large, subovate (
<figureCitation id="DFBCEEAE3C24FFB5FDDAFD8336E0FDF9" box="[634,762,541,566]" captionStart="Fig" captionStartId="28.[174,205,1869,1888]" captionTargetBox="[239,1395,251,1840]" captionTargetId="figure-5@28.[227,1398,240,1871]" captionTargetPageId="28" captionText="Fig. 22. AD, Borneosa kapit sp. nov., paratype male (23.5 × 19.1 mm) (ZRC 2021.0518), Belaga, Sarawak; EG, B. kapit sp. nov., holotype male (37.4 × 28.4 mm) (SMF 48750), Rajang River, Sarawak; H, B. kapit sp. nov., paratype male (35.8 × 28.0 mm) (ZRC 2021.0521), Kapit, Sarawak; I L, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak; MP, S. serrata sp. nov., holotype male (25.5 × 22.0 mm) (ZRC 2006.57), Sungei Pangean, Kalimantan Utara. Scale bars: A, D, E, H, I, L, M, P = 1.0 mm; B, C, F, G, J, K, N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8074574" httpUri="https://zenodo.org/record/8074574/files/figure.png" pageId="22" pageNumber="23">Fig. 22IK</figureCitation>
); vulvae almost round (
<figureCitation id="DFBCEEAE3C24FFB5FE36FDA335ECFD99" box="[406,502,573,598]" captionStart="Fig" captionStartId="29.[136,167,1841,1860]" captionTargetBox="[136,1413,418,1816]" captionTargetId="figure-171@29.[136,1413,418,1816]" captionTargetPageId="29" captionText="Fig. 23. Female thoracic sternum and vulvae. A, Borneosa tenebrosa (Holthuis, 1979), female (30.8 × 24.6 mm) (ZRC 1989.3420), Gunong Mulu National Park, Sarawak; B, B. aspera (Ng &amp; Stuebing, 1989), female (44.1 × 33.1 mm) (ZRC 2014.845), Temburong, Brunei; C, female (39.8 × 31.2 mm) (ZRC 2021.0840), Barito, Kalimantan Tengah; D, B. bario sp. nov., paratype female (24.3 × 19.3 mm) (ZRC 2021.0693), Bario, Sarawak; E, B. kapit sp. nov., paratype female (40.6 × 31.4 mm) (ZRC 2021.0521), Kapit, Sarawak; F, B. sarawakensis sp. nov., female (27.2 × 22.7 mm) (ZRC 2020.336), Lanjak-Entimau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.8074576" httpUri="https://zenodo.org/record/8074576/files/figure.png" pageId="22" pageNumber="23">Fig. 23F</figureCitation>
).
</paragraph>
<caption id="13F8A2A33C27FFB6FF28F81830F6F87A" ID-DOI="http://doi.org/10.5281/zenodo.8074562" ID-Zenodo-Dep="8074562" httpUri="https://zenodo.org/record/8074562/files/figure.png" pageId="21" pageNumber="22" startId="21.[136,168,1926,1945]" targetBox="[136,1413,566,1899]" targetPageId="21" targetType="figure">
<paragraph id="4738F22B3C27FFB6FF28F81830F6F87A" blockId="21.[136,1414,1926,2039]" pageId="21" pageNumber="22">
<emphasis id="75F32E393C27FFB6FF28F81834D4F856" bold="true" box="[136,206,1926,1945]" pageId="21" pageNumber="22">Fig. 16.</emphasis>
<taxonomicName id="808789A83C27FFB6FF7DF81835B6F856" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[221,428,1926,1945]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sarawakensis" status="sp. nov.">
<emphasis id="75F32E393C27FFB6FF7DF81835B6F856" box="[221,428,1926,1945]" italics="true" pageId="21" pageNumber="22">Borneosa sarawakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C27FFB6FE14F81835E6F856" box="[436,508,1926,1945]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak. A, left third maxilliped; B, anterior male thoracic sternum and pleon; C, male pleon; D, outer view of chelae; EH, right P2P5, respectively, all to same scale.
</paragraph>
</caption>
<footnote id="249CEE253C27FFB6FF28F87B3582F838" box="[136,408,2020,2039]" pageId="21" pageNumber="22">
<paragraph id="4738F22B3C27FFB6FF28F87B3582F838" blockId="21.[136,1414,1926,2039]" box="[136,408,2020,2039]" pageId="21" pageNumber="22">© 2022 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4738F22B3C24FFB5FF4AFDC33569FD1A" blockId="22.[174,775,253,1141]" pageId="22" pageNumber="23">
<emphasis id="75F32E393C24FFB5FF4AFDC3352CFDBA" box="[234,310,605,629]" italics="true" pageId="22" pageNumber="23">Colour</emphasis>
: In life, the carapace is maroon to reddish brown, darker on the anterior parts; the ambulatory legs and chela are olive brown with the ventral surface pale yellow (
<figureCitation id="DFBCEEAE3C24FFB5FEA5FD23357DFD1A" box="[261,359,701,725]" captionStart="Fig" captionStartId="31.[136,168,1897,1916]" captionTargetBox="[136,1413,1067,1872]" captionTargetId="figure-479@31.[136,1413,1067,1872]" captionTargetPageId="31" captionText="Fig. 25. A, type locality of B. sarawakensis sp. nov., Sungei Musing, first order stream feeding Batang Ai reservoir, Sri Aman, Sarawak; B, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak (freshly preserved); C, B. kapit sp. nov., paratype male (32.3 × 25.6 mm) (ZRC 2021.0520), Sungei Lebau, Pelagus, Kapit, Sarawak (freshly preserved)." figureDoi="http://doi.org/10.5281/zenodo.8074580" httpUri="https://zenodo.org/record/8074580/files/figure.png" pageId="22" pageNumber="23">Fig. 25B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C24FFB5FF4AFD43350BFCDA" pageId="22" pageNumber="23" type="etymology">
<paragraph id="4738F22B3C24FFB5FF4AFD43350BFCDA" blockId="22.[174,775,253,1141]" pageId="22" pageNumber="23">
<emphasis id="75F32E393C24FFB5FF4AFD433546FD3A" box="[234,348,733,757]" italics="true" pageId="22" pageNumber="23">Etymology</emphasis>
: The species is named after the state of
<collectingCountry id="3F90B2BB3C24FFB5FF0EFD633517FCDA" box="[174,269,765,789]" name="Malaysia" pageId="22" pageNumber="23">Sarawak</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C24FFB5FF4AFC8335AAFCBA" pageId="22" pageNumber="23" type="discussion">
<paragraph id="4738F22B3C24FFB5FF4AFC8335AAFCBA" blockId="22.[174,775,253,1141]" pageId="22" pageNumber="23">
<emphasis id="75F32E393C24FFB5FF4AFC83355DFCFA" box="[234,327,797,821]" italics="true" pageId="22" pageNumber="23">Remarks</emphasis>
: For differences between
<taxonomicName id="808789A83C24FFB5FDFAFC80371DFCFA" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[602,775,797,821]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sarawakensis" status="sp. nov.">
<emphasis id="75F32E393C24FFB5FDFAFC80371DFCFA" box="[602,775,797,821]" italics="true" pageId="22" pageNumber="23">B. sarawakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C24FFB5FF0EFCA0351CFC9A" box="[174,262,830,853]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
and its closest species,
<taxonomicName id="808789A83C24FFB5FDBAFCA03668FC9A" authority="Ng &amp; Grinang, 2022" authorityName="Ng &amp; Grinang" authorityYear="2022" box="[538,626,829,853]" class="Malacostraca" family="Gecarcinucidae" genus="Borneosa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kapit" status="sp. nov.">
<emphasis id="75F32E393C24FFB5FDBAFCA03668FC9A" box="[538,626,829,853]" italics="true" pageId="22" pageNumber="23">B. kapit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EC093423C24FFB5FDDCFCA036CCFC9A" box="[636,726,830,853]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, see remarks for latter taxon.
</paragraph>
</subSubSection>
<subSubSection id="0F9DA1A03C24FFB5FF4AFCE3350BFBBA" pageId="22" pageNumber="23" type="biology_ecology">
<paragraph id="4738F22B3C24FFB5FF4AFCE3350BFBBA" blockId="22.[174,775,253,1141]" pageId="22" pageNumber="23">
<emphasis id="75F32E393C24FFB5FF4AFCE33525FC5A" box="[234,319,893,917]" italics="true" pageId="22" pageNumber="23">Biology</emphasis>
: All the habitats in the
<typeStatus id="983C4C893C24FFB5FDF0FCE03665FC59" box="[592,639,894,918]" pageId="22" pageNumber="23">type</typeStatus>
locality are feeder streams of the Batang Ai hydroelectric reservoir, on a relatively high gradient, with rocky substrates and fast flowing clear, cold water (
<figureCitation id="DFBCEEAE3C24FFB5FE52FC43364CFC3A" box="[498,598,989,1013]" captionStart="Fig" captionStartId="31.[136,168,1897,1916]" captionTargetBox="[136,1413,1067,1872]" captionTargetId="figure-479@31.[136,1413,1067,1872]" captionTargetPageId="31" captionText="Fig. 25. A, type locality of B. sarawakensis sp. nov., Sungei Musing, first order stream feeding Batang Ai reservoir, Sri Aman, Sarawak; B, B. sarawakensis sp. nov., holotype male (35.5 × 28.1 mm) (ZRC 2021.0516), Batang Ai, Sarawak (freshly preserved); C, B. kapit sp. nov., paratype male (32.3 × 25.6 mm) (ZRC 2021.0520), Sungei Lebau, Pelagus, Kapit, Sarawak (freshly preserved)." figureDoi="http://doi.org/10.5281/zenodo.8074580" httpUri="https://zenodo.org/record/8074580/files/figure.png" pageId="22" pageNumber="23">Fig. 25A</figureCitation>
). The specimens from Lanjak-Entimau Wildlife Sanctuary, Kanowit, were collected from pristine forest streams with shallow water and a rocky substrate. These sites share the same drainage.
</paragraph>
</subSubSection>
</treatment>
</document>