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<document id="BB230D941C3285566D64C5A492F1C813" ID-DOI="10.5281/zenodo.280590" ID-GBIF-Dataset="35f6ecad-68d3-44cb-9183-d1c9617c2c57" ID-ISSN="1175-5326" ID-Zenodo-Dep="280590" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460462301487" checkinUser="plazi" docAuthor="Cárdenas, Paco &amp; Rapp, Hans Tore" docDate="2012" docId="A61787B7DC42FF9BFF33FB2CFE50FC61" docLanguage="en" docName="zt03253p053.pdf" docOrigin="Zootaxa 3253" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thenea valdiviae" docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="5A2EFFCFDC69FFB6FFA4FFC9FFB8FFE8" masterDocTitle="A review of Norwegian streptaster-bearing Astrophorida (Porifera: Demospongiae: Tetractinellida), new records and a new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="44" updateTime="1698304409925" updateUser="plazi">
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<mods:title id="CA9204EC7DDF247C9C5B5848FE907EFB">A review of Norwegian streptaster-bearing Astrophorida (Porifera: Demospongiae: Tetractinellida), new records and a new species</mods:title>
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<mods:namePart id="DE86FB875868755D4BE557BDB5EEB4E1">Cárdenas, Paco</mods:namePart>
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<mods:namePart id="C10309F37F42ED706F54D2BA0F253435">Rapp, Hans Tore</mods:namePart>
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<mods:date id="3C200D4C7C554F222BD185073BA7283C">2012</mods:date>
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<treatment id="A61787B7DC42FF9BFF33FB2CFE50FC61" ID-DOI="http://doi.org/10.5281/zenodo.6166259" ID-GBIF-Taxon="119512844" ID-Zenodo-Dep="6166259" LSID="urn:lsid:plazi:treatment:A61787B7DC42FF9BFF33FB2CFE50FC61" httpUri="http://treatment.plazi.org/id/A61787B7DC42FF9BFF33FB2CFE50FC61" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<subSubSection id="66A4652ADC42FF9DFF33FB2CFD76FA8E" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="2E0136A1DC42FF9DFF33FB2CFDC0FB17" blockId="43.[151,632,1253,1312]" box="[151,632,1253,1279]" pageId="43" pageNumber="44">
<heading id="754981CDDC42FF9DFF33FB2CFDC0FB17" bold="true" box="[151,632,1253,1279]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<emphasis id="1CCAEAB3DC42FF9DFF33FB2CFDC0FB17" bold="true" box="[151,632,1253,1279]" pageId="43" pageNumber="44">
<taxonomicName id="E9BE4D22DC42FF9DFF33FB2CFEDCFB17" box="[151,356,1253,1279]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC42FF9DFF33FB2CFEDCFB17" bold="true" box="[151,356,1253,1279]" italics="true" pageId="43" pageNumber="44">Thenea valdiviae</emphasis>
</taxonomicName>
von
<bibRefCitation id="4A2F4B50DC42FF9DFE05FB2CFDC0FB17" author="Lendenfeld" box="[417,632,1253,1279]" pageId="43" pageNumber="44" refString="von Lendenfeld, R. (1907) Die Tetraxonia. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf der Dampfer Valdivia 1898 - 1899, 11, i - iv, 59 - 374, pls IX - XLVI." type="journal article" year="1907">Lendenfeld, 1907</bibRefCitation>
</emphasis>
</heading>
</paragraph>
<paragraph id="2E0136A1DC42FF9DFF33FAC1FE5AFAC8" blockId="43.[151,632,1253,1312]" box="[151,482,1288,1312]" pageId="43" pageNumber="44">
(
<figureCitation id="B6852A24DC42FF9DFF3AFAC1FEADFAC8" box="[158,277,1288,1312]" captionStart="FIGURE 18" captionStartId="34.[151,250,1922,1945]" captionTargetBox="[216,1384,193,1910]" captionTargetId="figure@34.[216,1385,193,1911]" captionTargetPageId="34" captionText="FIGURE 18. Streptasters of Norwegian species of Thenea. Streptasters of Thenea abyssorum Koltun, 1964 [ZMBN 85228]. A. Plesiaster; B. Spirasters; Streptasters of Thenea levis von Lendenfeld, 1907 [ZMBN 85230]. C. Plesiaster. Same scale as A; D. spirasters. Streptasters of Thenea muricata (Bowerbank, 1858) [ZMBN 85231]. E. Plesiaster; F. Spiraster; Streptasters of Thenea valdiviae von Lendenfeld, 1907. G. Plesiaster [ZMBN 85257]; H. Spirasters. Same scale as E. [ZMBN 85233]." httpUri="https://zenodo.org/record/280608/files/figure.png" pageId="43" pageNumber="44">Figures 18</figureCitation>
GH, 24,
<tableCitation id="633C031ADC42FF9DFE22FAC1FE60FAC8" box="[390,472,1288,1312]" captionStart="TABLE 5" captionStartId="42.[151,239,979,1003]" captionTargetBox="[197,1424,1176,1971]" captionTargetPageId="42" captionText="TABLE 5. Depth, individual spicule dimensions of streptasters (in µm) for specimens of Thenea valdiviae, Thenea muricata, Thenea schmidti and Thenea bojeadori. Means are in bold; other values are ranges; n = 30 unless stated otherwise between parentheses. - = not referred." httpUri="http://table.plazi.org/id/7AC16629DC43FF9CFF33FC1AFD20FBC6" pageId="43" pageNumber="44" tableUuid="7AC16629DC43FF9CFF33FC1AFD20FBC6">Table 5</tableCitation>
)
</paragraph>
<paragraph id="2E0136A1DC42FF9DFF33FA98FD76FA8E" blockId="43.[151,1436,1360,1566]" box="[151,718,1360,1382]" pageId="43" pageNumber="44">
<emphasis id="1CCAEAB3DC42FF9DFF33FA98FF46FA8E" box="[151,254,1361,1382]" italics="true" pageId="43" pageNumber="44">Synonymy</emphasis>
(modified from
<bibRefCitation id="4A2F4B50DC42FF9DFE01FA99FD72FA8E" author="Steenstrup" box="[421,714,1360,1382]" pageId="43" pageNumber="44" refString="Steenstrup, E. &amp; Tendal, O. S. (1982) The genus Thenea (Porifera, Demospongiae, Choristida) in the Norwegian Sea and adjacent waters; an annotated key. Sarsia, 67, 259 - 268." type="journal article" year="1982">Steenstrup &amp; Tendal (1982))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66A4652ADC42FF9DFF33FAA6FD4AF9F6" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="2E0136A1DC42FF9DFF33FAA6FCDFFA4B" blockId="43.[151,1436,1360,1566]" pageId="43" pageNumber="44">
<taxonomicName id="E9BE4D22DC42FF9DFF33FAA6FEFEFA6D" box="[151,326,1391,1413]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC42FF9DFF33FAA6FEFEFA6D" box="[151,326,1391,1413]" italics="true" pageId="43" pageNumber="44">Thenea valdiviae</emphasis>
</taxonomicName>
von
<bibRefCitation id="4A2F4B50DC42FF9DFED8FAA7FD8CFA6C" author="Lendenfeld" box="[380,564,1390,1412]" pageId="43" pageNumber="44" refString="von Lendenfeld, R. (1907) Die Tetraxonia. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf der Dampfer Valdivia 1898 - 1899, 11, i - iv, 59 - 374, pls IX - XLVI." type="journal article" year="1907">Lendenfeld, 1907</bibRefCitation>
: von
<bibRefCitation id="4A2F4B50DC42FF9DFDCAFAA7FCA7FA6C" author="Lendenfeld" box="[622,799,1390,1412]" pageId="43" pageNumber="44" refString="von Lendenfeld, R. (1907) Die Tetraxonia. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf der Dampfer Valdivia 1898 - 1899, 11, i - iv, 59 - 374, pls IX - XLVI." type="journal article" year="1907">Lendenfeld 1907</bibRefCitation>
, p. 190;
<bibRefCitation id="4A2F4B50DC42FF9DFCDFFAA7FBA5FA6C" author="Hentschel" box="[891,1053,1390,1412]" pageId="43" pageNumber="44" refString="Hentschel, E. (1929) Die Kiesel- und Hornschwamme des Nordlichen Eismeers. In: Romer, F., Schaudinn, F., Brauer, A. &amp; Arndt, W. (Eds.) Fauna Arctica. Eine Zusammenstellung der arktischen Tierformen mit besonderer Berucksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nordliche Eismeer im Jahre 1898. G. Fischer, Jena, pp. 857 - 1042, pls XII - XIV." type="journal article" year="1929">Hentschel 1929</bibRefCitation>
, p. 863, p. 918;
<bibRefCitation id="4A2F4B50DC42FF9DFB6DFAA6FF44FA4B" author="Steenstrup" pageId="43" pageNumber="44" refString="Steenstrup, E. &amp; Tendal, O. S. (1982) The genus Thenea (Porifera, Demospongiae, Choristida) in the Norwegian Sea and adjacent waters; an annotated key. Sarsia, 67, 259 - 268." type="journal article" year="1982">Steenstrup &amp; Tendal 1982</bibRefCitation>
, p. 259;
<bibRefCitation id="4A2F4B50DC42FF9DFEF7FA44FE57FA4B" author="Klitgaard" box="[339,495,1421,1443]" pageId="43" pageNumber="44" refString="Klitgaard, A. B. (1995) The fauna associated with outer shelf and upper slope sponges (Porifera, Demospongiae) at the Faroe Islands, northeastern Atlantic. Sarsia, 80, 1 - 22." type="journal article" year="1995">Klitgaard 1995</bibRefCitation>
; p. 1;
<bibRefCitation id="4A2F4B50DC42FF9DFD88FA44FD45FA4B" author="Cardenas" box="[556,765,1421,1443]" pageId="43" pageNumber="44" refString="Cardenas, P., Xavier, J. R., Reveillaud, J., Schander, C. &amp; Rapp, H. T. (2011) Molecular phylogeny of the Astrophorida (Porifera, Demospongiae p) reveals an unexpected high level of spicule homoplasy. PLoS ONE, 6, e 18318. doi: 10.1371 / journal. pone. 0018318." type="journal article" year="2011">
Cárdenas
<emphasis id="1CCAEAB3DC42FF9DFD34FA47FD05FA4B" box="[656,701,1421,1443]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
. 2011
</bibRefCitation>
, Table S1.
</paragraph>
<paragraph id="2E0136A1DC42FF9DFF33FA65FE21FA08" blockId="43.[151,1436,1360,1566]" pageId="43" pageNumber="44">
<taxonomicName id="E9BE4D22DC42FF9DFF33FA65FE5CFA29" authority="Vosmaer 1882" authorityName="Vosmaer" authorityYear="1882" box="[151,484,1452,1474]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC42FF9DFF33FA65FEFBFA2A" box="[151,323,1452,1474]" italics="true" pageId="43" pageNumber="44">Thenea muricata</emphasis>
:
<bibRefCitation id="4A2F4B50DC42FF9DFEF5FA64FE5CFA29" author="Vosmaer" box="[337,484,1452,1474]" pageId="43" pageNumber="44" refString="Vosmaer, G. C. J. (1882) Report on the sponges dredged up in the Arctic Sea by the ' Willem Barents' in the years 1878 and 1879. Niederlandisches Archiv fur Zoologie Supplement, 1, 1 - 58, pls I - IV." type="journal article" year="1882">Vosmaer 1882</bibRefCitation>
</taxonomicName>
, p. 5;
<bibRefCitation id="4A2F4B50DC42FF9DFD86FA64FD0EFA2A" author="Vosmaer" box="[546,694,1452,1474]" pageId="43" pageNumber="44" refString="Vosmaer, G. C. J. (1885) The Sponges of the ' Willem Barents' Expedition 1880 and 1881. Bijdragen tot de Dierkunde, 12, 1 - 47, pls I - V." type="journal article" year="1885">Vosmaer 1885</bibRefCitation>
, p. 4;
<bibRefCitation id="4A2F4B50DC42FF9DFD57FA65FC34FA2A" author="Levinsen" box="[755,908,1452,1474]" pageId="43" pageNumber="44" refString="Levinsen, G. M. R. (1886) Kara-Havets Svampe (Porifera). Dijmphna-Togtets zoologisk-botaniske Udbytte, 341 - 372, pls. XXIX - XXXI." type="book chapter" year="1886">Levinsen 1886</bibRefCitation>
, p. 343;
<bibRefCitation id="4A2F4B50DC42FF9DFC47FA65FB3BFA29" author="Lundbeck" box="[995,1155,1452,1474]" pageId="43" pageNumber="44" refString="Lundbeck, W. (1909) The Porifera of East-Greenland. Meddelelser om Gronland, 29, 423 - 464, pl XIV." type="journal article" year="1909">Lundbeck 1909</bibRefCitation>
, p. 427;
<bibRefCitation id="4A2F4B50DC42FF9DFB7FFA65FAC6FA29" author="Brondsted" box="[1243,1406,1452,1474]" pageId="43" pageNumber="44" refString="Brondsted, H. V. (1933) The Godthaab expedition 1928. Porifera. Meddelelser om Gronland, 79, 1 - 25." type="journal article" year="1933">Brøndsted 1933</bibRefCitation>
, p. 6;
<bibRefCitation id="4A2F4B50DC42FF9DFF45FA05FEDAFA08" author="Burton" box="[225,354,1483,1505]" pageId="43" pageNumber="44" refString="Burton, M. (1934) Report on the sponges of the Norwegian expeditions to East-Greenland (1930, 1931, and 1932). Skrifter om Svaldbard og Ishavet, 61, 1 - 33." type="journal article" year="1934">Burton 1934</bibRefCitation>
, p. 6.
</paragraph>
<paragraph id="2E0136A1DC42FF9DFF33FA20FD3AFA17" blockId="43.[151,1436,1360,1566]" box="[151,642,1513,1535]" pageId="43" pageNumber="44">
<taxonomicName id="E9BE4D22DC42FF9DFF33FA20FEFBFA17" box="[151,323,1513,1535]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC42FF9DFF33FA20FEFBFA17" box="[151,323,1513,1535]" italics="true" pageId="43" pageNumber="44">Thenea muricata</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="4A2F4B50DC42FF9DFE0DFA20FD87FA17" author="Stephens" box="[425,575,1513,1535]" pageId="43" pageNumber="44" refString="Stephens, J. (1915) Sponges of the Coasts of Ireland. I. - The Triaxonia and part of the Tetraxonida. Fisheries, Ireland Scientific Investigations, 1914, 1 - 43, pls I - V." type="journal article" year="1915">Stephens 1915</bibRefCitation>
, p. 11.
</paragraph>
<paragraph id="2E0136A1DC42FF9DFF33F9C0FD4AF9F6" blockId="43.[151,1436,1360,1566]" box="[151,754,1544,1566]" pageId="43" pageNumber="44">
?
<taxonomicName id="E9BE4D22DC42FF9DFF06F9C1FE5AF9F6" authority="Hansen 1885" authorityName="Hansen" authorityYear="1885" box="[162,482,1544,1566]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC42FF9DFF06F9C1FEF6F9F6" box="[162,334,1544,1566]" italics="true" pageId="43" pageNumber="44">Thenea muricata</emphasis>
:
<bibRefCitation id="4A2F4B50DC42FF9DFEFFF9C0FE5AF9F6" author="Hansen" box="[347,482,1544,1566]" pageId="43" pageNumber="44" refString="Hansen, G. A. (1885) Spongiadae. The Norwegian North-Atlantic Expedition 1876 - 1878. Zoologi, 13, 1 - 26, pls I - VII, 1 map." type="book chapter" year="1885">Hansen 1885</bibRefCitation>
</taxonomicName>
, p. 18;
<bibRefCitation id="4A2F4B50DC42FF9DFD89F9C1FD16F9F6" author="Koltun" box="[557,686,1544,1566]" pageId="43" pageNumber="44" refString="Koltun, V. M. (1966) Four-rayed sponges of Northern and Far Eastern seas of the USSR (order Tetraxonida). Opredeliti Faunei SSSR 90. (Zoological Institute of the Academy of Sciences of the USSR: Moscow, Leningrad), 1 - 112, pls I - XXXVIII." type="book chapter" year="1966">Koltun 1966</bibRefCitation>
, p. 36.
</paragraph>
</subSubSection>
<subSubSection id="66A4652ADC42FF9AFF33F982FCDAF87A" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="2E0136A1DC42FF9DFF33F982FB87F8D2" blockId="43.[151,1436,1610,1994]" pageId="43" pageNumber="44">
<emphasis id="1CCAEAB3DC42FF9DFF33F982FEB8F98B" box="[151,256,1611,1635]" italics="true" pageId="43" pageNumber="44">Material.</emphasis>
<collectionCode id="48AFAE64DC42FF9DFEA3F982FEE0F98A" box="[263,344,1611,1634]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="43" pageNumber="44">ZMBN</collectionCode>
85255, west of Marstein, western
<collectingCountry id="56A97631DC42FF9DFD44F982FC83F98B" box="[736,827,1611,1635]" name="Norway" pageId="43" pageNumber="44">Norway</collectingCountry>
,
<geoCoordinate id="4B8A5066DC42FF9DFCE3F983FC06F98A" box="[839,958,1610,1634]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="15" value="60.138336">60°8'18''N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFC6FF983FB87F98A" box="[971,1087,1610,1634]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="15" value="4.846389">4°50'47''E</geoCoordinate>
,
<quantity id="E9469B44DC42FF9DFBEFF983FB2DF98A" box="[1099,1173,1610,1634]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="43" pageNumber="44" unit="m" value="300.0">300 m</quantity>
, Agassiz trawl;
<collectionCode id="48AFAE64DC42FF9DFAEFF982FA24F98A" box="[1355,1436,1611,1634]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="43" pageNumber="44">ZMBN</collectionCode>
85256, Freifjorden, Møre, western
<collectingCountry id="56A97631DC42FF9DFD87F9B9FDC6F960" box="[547,638,1648,1672]" name="Norway" pageId="43" pageNumber="44">Norway</collectingCountry>
,
<geoCoordinate id="4B8A5066DC42FF9DFD2EF9A6FCB4F96F" box="[650,780,1647,1671]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="9" value="63.034832">63°02.09'N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFCBCF9A6FC31F96F" box="[792,905,1647,1671]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="9" value="7.8636665">7°51.82'E</geoCoordinate>
,
<quantity id="E9469B44DC42FF9DFC31F9A6FC67F96F" box="[917,991,1647,1671]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="m" value="100.0">100 m</quantity>
, Møre 2006 cruise;
<collectionCode id="48AFAE64DC42FF9DFB67F9B9FAFFF96F" box="[1219,1351,1648,1671]" httpUri="http://grbio.org/cool/pnpi-f6fs" name="Norwegian University of Science and Technology - University Museum" pageId="43" pageNumber="44">NTNU-VM</collectionCode>
55620, Sandøy, Risvaerfjorden,
<geoCoordinate id="4B8A5066DC42FF9DFE12F95BFDE9F942" box="[438,593,1682,1706]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="65.004166">65°0'15.00&quot;N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFDC1F95BFCB3F942" box="[613,779,1682,1706]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="11.491667">11°29'30.00&quot;E</geoCoordinate>
,
<quantity id="E9469B44DC42FF9DFCBAF95BFC10F942" box="[798,936,1682,1706]" metricMagnitude="2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="43" pageNumber="44" unit="m" value="115.0" valueMax="130.0" valueMin="100.0">100130 m</quantity>
;
<collectionCode id="48AFAE64DC42FF9DFC18F95AFBB5F942" box="[956,1037,1683,1706]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="43" pageNumber="44">ZMBN</collectionCode>
85257, near Røst reef, northern
<collectingCountry id="56A97631DC42FF9DFF33F971FF4AF938" box="[151,242,1720,1744]" name="Norway" pageId="43" pageNumber="44">Norway</collectingCountry>
,
<geoCoordinate id="4B8A5066DC42FF9DFEA4F97EFE3AF927" box="[256,386,1719,1743]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="9" value="67.587166">67°35.23'N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFE34F97EFDB8F927" box="[400,512,1719,1743]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="9" value="9.482">9°28.92'E</geoCoordinate>
, Polarstern ARK-XXII/1a 2007 cruise,
<quantity id="E9469B44DC42FF9DFC6FF97EFBAFF927" box="[971,1047,1719,1743]" metricMagnitude="2" metricUnit="m" metricValue="5.9799999999999995" pageId="43" pageNumber="44" unit="m" value="598.0">598 m</quantity>
;
<collectionCode id="48AFAE64DC42FF9DFB81F971FB12F927" box="[1061,1194,1720,1743]" httpUri="http://grbio.org/cool/pnpi-f6fs" name="Norwegian University of Science and Technology - University Museum" pageId="43" pageNumber="44">NTNU-VM</collectionCode>
55545, Vågsfjorden, Vesterålen, northern
<collectingCountry id="56A97631DC42FF9DFE36F912FE55F91B" box="[402,493,1755,1779]" name="Norway" pageId="43" pageNumber="44">Norway</collectingCountry>
,
<geoCoordinate id="4B8A5066DC42FF9DFDA6F913FD25F91A" box="[514,669,1754,1778]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="69.008606">69°0'30.99&quot;N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFD16F913FCE1F91A" box="[690,857,1754,1778]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="16.681385">16°40'52.99&quot;E</geoCoordinate>
,
<quantity id="E9469B44DC42FF9DFCC9F913FC41F91A" box="[877,1017,1754,1778]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="43" pageNumber="44" unit="m" value="275.0" valueMax="300.0" valueMin="250.0">250300 m</quantity>
;
<collectionCode id="48AFAE64DC42FF9DFBA9F912FB2CF91A" box="[1037,1172,1755,1778]" httpUri="http://grbio.org/cool/pnpi-f6fs" name="Norwegian University of Science and Technology - University Museum" pageId="43" pageNumber="44">NTNU-VM</collectionCode>
, 54950, Barents Sea,
<geoCoordinate id="4B8A5066DC42FF9DFF33F936FF4DF8FF" box="[151,245,1791,1815]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="925" value="70.71667">70°43'N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFEA6F936FEE5F8FF" box="[258,349,1791,1815]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="925" value="16.933332">16°56'E</geoCoordinate>
,
<quantity id="E9469B44DC42FF9DFECEF936FE0DF8FF" box="[362,437,1791,1815]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="43" pageNumber="44" unit="m" value="740.0">740 m</quantity>
;
<collectionCode id="48AFAE64DC42FF9DFE65F8C9FDFDF8FF" box="[449,581,1792,1815]" httpUri="http://grbio.org/cool/pnpi-f6fs" name="Norwegian University of Science and Technology - University Museum" pageId="43" pageNumber="44">NTNU-VM</collectionCode>
54975, Barents Sea,
<geoCoordinate id="4B8A5066DC42FF9DFC93F936FC2EF8FF" box="[823,918,1791,1815]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="925" value="70.76667">70°46'N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFC06F936FC46F8FF" box="[930,1022,1791,1815]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="925" value="17.0">17°00'E</geoCoordinate>
,
<quantity id="E9469B44DC42FF9DFBAEF936FBEDF8FF" box="[1034,1109,1791,1815]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="43" pageNumber="44" unit="m" value="910.0">910 m</quantity>
;
<collectionCode id="48AFAE64DC42FF9DFBC6F8C9FB0BF8FF" box="[1122,1203,1792,1815]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="43" pageNumber="44">ZMBN</collectionCode>
85233, Barents Sea, off Finmark, Ecosystem Barents Sea 2007 cruise,
<geoCoordinate id="4B8A5066DC42FF9DFD7EF8EBFCE4F8D2" box="[730,860,1826,1850]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="9" value="71.805336">71°48.32'N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFCC3F8EBFC5EF8D2" box="[871,998,1826,1850]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="9" value="28.662666">28°39.76'E</geoCoordinate>
,
<specimenCount id="38B8FD28DC42FF9DFC55F8EBFB87F8D2" box="[1009,1087,1826,1850]" pageId="43" pageNumber="44" type="generic">
<quantity id="E9469B44DC42FF9DFC55F8EBFB83F8D2" box="[1009,1083,1826,1850]" metricMagnitude="2" metricUnit="m" metricValue="2.9699999999999998" pageId="43" pageNumber="44" unit="m" value="297.0">297 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="2E0136A1DC42FF9DFF63F881FE7EF8B7" blockId="43.[151,1436,1610,1994]" box="[199,454,1863,1887]" pageId="43" pageNumber="44">
<emphasis id="1CCAEAB3DC42FF9DFF63F881FE7EF8B7" box="[199,454,1863,1887]" italics="true" pageId="43" pageNumber="44">Comparative material.</emphasis>
</paragraph>
<paragraph id="2E0136A1DC42FF9DFF63F8A2FAECF822" blockId="43.[151,1436,1610,1994]" pageId="43" pageNumber="44">
<taxonomicName id="E9BE4D22DC42FF9DFF63F8A2FE3FF86B" box="[199,391,1899,1923]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC42FF9DFF63F8A2FE3FF86B" box="[199,391,1899,1923]" italics="true" pageId="43" pageNumber="44">Thenea valdiviae</emphasis>
</taxonomicName>
, ZMAPOR 2385, Willem Barents Sea Expedition
<date id="5A001061DC42FF9DFC62F8A3FB92F86A" box="[966,1066,1898,1922]" pageId="43" pageNumber="44" value="1878" valueMax="1884">187884</date>
, off Finnmark, northern
<collectingCountry id="56A97631DC42FF9DFAE5F8A2FA23F86B" box="[1345,1435,1899,1923]" name="Norway" pageId="43" pageNumber="44">Norway</collectingCountry>
(
<geoCoordinate id="4B8A5066DC42FF9DFF3BF846FE83F84F" box="[159,315,1935,1959]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="71.8667">71°52'0.11&quot;N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFEEDF846FE49F84F" box="[329,497,1935,1959]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="19.7833">19°46'59.88&quot;E</geoCoordinate>
),
<quantity id="E9469B44DC42FF9DFDA3F846FDEBF84F" box="[519,595,1935,1959]" metricMagnitude="2" metricUnit="m" metricValue="3.2399999999999998" pageId="43" pageNumber="44" unit="m" value="324.0">324 m</quantity>
, identified by G. C. J. Vosmaer as
<taxonomicName id="E9BE4D22DC42FF9DFC48F859FBD5F84F" box="[1004,1133,1936,1959]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC42FF9DFC48F859FBD5F84F" box="[1004,1133,1936,1959]" italics="true" pageId="43" pageNumber="44">T. muricata</emphasis>
</taxonomicName>
, det. P. Cárdenas;
<collectionCode id="48AFAE64DC42FF9DFAEEF859FA23F84F" box="[1354,1435,1936,1959]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="43" pageNumber="44">ZMBN</collectionCode>
85234, Schultz Massive Seamount,
<collectingCountry id="56A97631DC42FF9DFD8CF87BFD25F822" box="[552,669,1970,1994]" name="Greenland" pageId="43" pageNumber="44">Greenland</collectingCountry>
Sea,
<geoCoordinate id="4B8A5066DC42FF9DFD7EF87BFC81F822" box="[730,825,1970,1994]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="925" value="73.833336">73°50'N</geoCoordinate>
,
<geoCoordinate id="4B8A5066DC42FF9DFCE0F87BFC1FF822" box="[836,935,1970,1994]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="92" value="7.5383334">7°32.3'E</geoCoordinate>
, BIODEEP 2007 cruise,
<quantity id="E9469B44DC42FF9DFB64F87BFAB1F822" box="[1216,1289,1970,1994]" metricMagnitude="2" metricUnit="m" metricValue="5.779999999999999" pageId="43" pageNumber="44" unit="m" value="578.0">578 m</quantity>
, ROV.
</paragraph>
<paragraph id="2E0136A1DC45FF9AFF63FF51FCCBFE8C" blockId="44.[151,1436,151,357]" pageId="44" pageNumber="45">
<emphasis id="1CCAEAB3DC45FF9AFF63FF51FE21FF58" box="[199,409,152,176]" italics="true" pageId="44" pageNumber="45">Outer morphology</emphasis>
(
<figureCitation id="B6852A24DC45FF9AFE0FFF5EFE46FF47" box="[427,510,151,175]" captionStart="FIGURE 24" captionStartId="44.[151,250,1328,1351]" captionTargetBox="[176,1436,406,1304]" captionTargetId="figure@44.[151,1436,375,1304]" captionTargetPageId="44" captionText="FIGURE 24. Thenea valdiviae von Lendenfeld, 1907. A. Whole specimen from Barents Sea, top view. Note the sieve on the oscule [ZMBN 85233]. Scale: 2 cm; B. Whole specimen, side view showing the poral area, also with sieve [ZMBN 85233]; C. Whole specimen from Freisfjorden (Møre), top view. Note the sieve on the oscule. Scale: 1 cm [ZMBN 85256]; D. Top surface of a young specimen covered with buds. p: pores. os: oscule. [ZMBN 85257]; E. Thick section showing the skeletal organization. Scale: 1 mm [ZMBN 85233]." httpUri="https://zenodo.org/record/280614/files/figure.png" pageId="44" pageNumber="45">Fig. 24</figureCitation>
AD). Massive sponge. Young specimens (
<collectionCode id="48AFAE64DC45FF9AFC48FF5EFB87FF47" box="[1004,1087,151,175]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="44" pageNumber="45">ZMBN</collectionCode>
85257) have cribriporal pores and a uniporal oscule on opposite sides (as in young
<taxonomicName id="E9BE4D22DC45FF9AFD51FF74FCCEFF3C" box="[757,886,189,212]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="44" pageNumber="45" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC45FF9AFD51FF74FCCEFF3C" box="[757,886,189,212]" italics="true" pageId="44" pageNumber="45">T. muricata</emphasis>
</taxonomicName>
), with a flat top surface. A fringe surrounds the oscule. Budding was observed on the top surface of these young specimens (
<figureCitation id="B6852A24DC45FF9AFC5CFF16FBF3FF1F" box="[1016,1099,223,247]" captionStart="FIGURE 24" captionStartId="44.[151,250,1328,1351]" captionTargetBox="[176,1436,406,1304]" captionTargetId="figure@44.[151,1436,375,1304]" captionTargetPageId="44" captionText="FIGURE 24. Thenea valdiviae von Lendenfeld, 1907. A. Whole specimen from Barents Sea, top view. Note the sieve on the oscule [ZMBN 85233]. Scale: 2 cm; B. Whole specimen, side view showing the poral area, also with sieve [ZMBN 85233]; C. Whole specimen from Freisfjorden (Møre), top view. Note the sieve on the oscule. Scale: 1 cm [ZMBN 85256]; D. Top surface of a young specimen covered with buds. p: pores. os: oscule. [ZMBN 85257]; E. Thick section showing the skeletal organization. Scale: 1 mm [ZMBN 85233]." httpUri="https://zenodo.org/record/280614/files/figure.png" pageId="44" pageNumber="45">Fig. 24</figureCitation>
D). Larger specimens are subcircular (
<collectionCode id="48AFAE64DC45FF9AFF5FFECDFEF6FEF4" box="[251,334,260,284]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="44" pageNumber="45">ZMBN</collectionCode>
85255, 85233) or slightly flattened (
<collectionCode id="48AFAE64DC45FF9AFD4FFECDFC85FEF4" box="[747,829,260,284]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="44" pageNumber="45">ZMBN</collectionCode>
85256) with a triangular shape. They look very much like
<taxonomicName id="E9BE4D22DC45FF9AFF6EFEE0FEF4FEA8" box="[202,332,297,320]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="44" pageNumber="45" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC45FF9AFF6EFEE0FEF4FEA8" box="[202,332,297,320]" italics="true" pageId="44" pageNumber="45">T. muricata</emphasis>
</taxonomicName>
but the oscule, separated from the poral area and situated on the top, always has a coarse sieve covering it. Surface is minutely to very hispid. Not compressible.
</paragraph>
<caption id="7AC16629DC45FF9AFF33FAF9FDF5FA29" httpUri="https://zenodo.org/record/280614/files/figure.png" pageId="44" pageNumber="45" targetBox="[176,1436,406,1304]" targetPageId="44">
<paragraph id="2E0136A1DC45FF9AFF33FAF9FDF5FA29" blockId="44.[151,1437,1328,1473]" pageId="44" pageNumber="45">
<emphasis id="1CCAEAB3DC45FF9AFF33FAF9FE99FAAE" bold="true" box="[151,289,1328,1351]" pageId="44" pageNumber="45">FIGURE 24.</emphasis>
<taxonomicName id="E9BE4D22DC45FF9AFE83FAF9FE6EFAAE" box="[295,470,1328,1350]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="44" pageNumber="45" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC45FF9AFE83FAF9FE6EFAAE" box="[295,470,1328,1350]" italics="true" pageId="44" pageNumber="45">Thenea valdiviae</emphasis>
</taxonomicName>
von Lendenfeld, 1907. A. Whole specimen from Barents Sea, top view. Note the sieve on the oscule [ZMBN 85233]. Scale: 2 cm; B. Whole specimen, side view showing the poral area, also with sieve [ZMBN 85233]; C. Whole specimen from Freisfjorden (Møre), top view. Note the sieve on the oscule. Scale: 1 cm [ZMBN 85256]; D. Top surface of a young specimen covered with buds. p: pores. os: oscule. [ZMBN 85257]; E. Thick section showing the skeletal organization. Scale: 1 mm [ZMBN 85233].
</paragraph>
</caption>
<paragraph id="2E0136A1DC45FF9AFF63FA27FD3CF9EE" blockId="44.[151,1436,1518,2008]" box="[199,644,1518,1542]" pageId="44" pageNumber="45">
<emphasis id="1CCAEAB3DC45FF9AFF63FA27FE9EF9EE" box="[199,294,1518,1542]" italics="true" pageId="44" pageNumber="45">Skeleton</emphasis>
(
<figureCitation id="B6852A24DC45FF9AFE91FA27FE30F9EE" box="[309,392,1518,1542]" captionStart="FIGURE 24" captionStartId="44.[151,250,1328,1351]" captionTargetBox="[176,1436,406,1304]" captionTargetId="figure@44.[151,1436,375,1304]" captionTargetPageId="44" captionText="FIGURE 24. Thenea valdiviae von Lendenfeld, 1907. A. Whole specimen from Barents Sea, top view. Note the sieve on the oscule [ZMBN 85233]. Scale: 2 cm; B. Whole specimen, side view showing the poral area, also with sieve [ZMBN 85233]; C. Whole specimen from Freisfjorden (Møre), top view. Note the sieve on the oscule. Scale: 1 cm [ZMBN 85256]; D. Top surface of a young specimen covered with buds. p: pores. os: oscule. [ZMBN 85257]; E. Thick section showing the skeletal organization. Scale: 1 mm [ZMBN 85233]." httpUri="https://zenodo.org/record/280614/files/figure.png" pageId="44" pageNumber="45">Fig. 24</figureCitation>
E). Like
<taxonomicName id="E9BE4D22DC45FF9AFE4CFA26FDC5F9EE" box="[488,637,1518,1542]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="44" pageNumber="45" phylum="Porifera" rank="species" species="abyssorum">
<emphasis id="1CCAEAB3DC45FF9AFE4CFA26FDC5F9EE" box="[488,637,1518,1542]" italics="true" pageId="44" pageNumber="45">T. abyssorum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="2E0136A1DC45FF9AFF63F9DBFBBEF8CE" blockId="44.[151,1436,1518,2008]" pageId="44" pageNumber="45">
<emphasis id="1CCAEAB3DC45FF9AFF63F9DBFE9DF9C2" box="[199,293,1554,1578]" italics="true" pageId="44" pageNumber="45">Spicules</emphasis>
(
<collectionCode id="48AFAE64DC45FF9AFE93F9DBFE32F9C2" box="[311,394,1554,1578]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="44" pageNumber="45">ZMBN</collectionCode>
85233) (
<figureCitation id="B6852A24DC45FF9AFE53F9DBFDF2F9C2" box="[503,586,1554,1578]" captionStart="FIGURE 18" captionStartId="34.[151,250,1922,1945]" captionTargetBox="[216,1384,193,1910]" captionTargetId="figure@34.[216,1385,193,1911]" captionTargetPageId="34" captionText="FIGURE 18. Streptasters of Norwegian species of Thenea. Streptasters of Thenea abyssorum Koltun, 1964 [ZMBN 85228]. A. Plesiaster; B. Spirasters; Streptasters of Thenea levis von Lendenfeld, 1907 [ZMBN 85230]. C. Plesiaster. Same scale as A; D. spirasters. Streptasters of Thenea muricata (Bowerbank, 1858) [ZMBN 85231]. E. Plesiaster; F. Spiraster; Streptasters of Thenea valdiviae von Lendenfeld, 1907. G. Plesiaster [ZMBN 85257]; H. Spirasters. Same scale as E. [ZMBN 85233]." httpUri="https://zenodo.org/record/280608/files/figure.png" pageId="44" pageNumber="45">Fig. 18</figureCitation>
GH). (a) oxeas, slightly bent, length:
<quantity id="E9469B44DC45FF9AFBB6F9DBFB3CF9C2" box="[1042,1156,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="44" pageNumber="45" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">410 mm</quantity>
; width: 20
<emphasis id="1CCAEAB3DC45FF9AFAB4F9DBFAF9F9C2" box="[1296,1345,1554,1578]" italics="true" pageId="44" pageNumber="45">48.1</emphasis>
70 µm (N=8). (b) dichotriaenes, rhabdome is with a single bend, more rarely flexuous, most rhabdomes have a swelling just below the cladome, rhabdome length: 620
<emphasis id="1CCAEAB3DC45FF9AFD1FF993FCB0F99A" box="[699,776,1626,1650]" italics="true" pageId="44" pageNumber="45">3933.7</emphasis>
5075 µm (N=8); width: 23
<emphasis id="1CCAEAB3DC45FF9AFBFCF993FB31F99A" box="[1112,1161,1626,1650]" italics="true" pageId="44" pageNumber="45">58.1</emphasis>
90 µm; many irregular cladomes, protoclad length: 75
<emphasis id="1CCAEAB3DC45FF9AFDA8F9B7FDF3F97E" box="[524,587,1662,1686]" italics="true" pageId="44" pageNumber="45">236.8</emphasis>
350 µm; deuteroclad length: 125
<emphasis id="1CCAEAB3DC45FF9AFC42F9B7FB9DF97E" box="[998,1061,1662,1686]" italics="true" pageId="44" pageNumber="45">600.2</emphasis>
1175 µm. (c) anatriaenes, rare, rhabdome straight or bent, length: 40004125 µm (N=2); rhabdome width: 1520 µm; bent clads (similar to
<taxonomicName id="E9BE4D22DC45FF9AFA23F96AFF44F936" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="44" pageNumber="45" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC45FF9AFA23F96AFF44F936" italics="true" pageId="44" pageNumber="45">T. muricata</emphasis>
</taxonomicName>
anatriaenes), clad length: 170220 µm. (d) protriaenes, not observed in this specimen. (e) spirasters (and rare metasters) (
<figureCitation id="B6852A24DC45FF9AFEF5F923FE1CF8EA" box="[337,420,1770,1794]" captionStart="FIGURE 18" captionStartId="34.[151,250,1922,1945]" captionTargetBox="[216,1384,193,1910]" captionTargetId="figure@34.[216,1385,193,1911]" captionTargetPageId="34" captionText="FIGURE 18. Streptasters of Norwegian species of Thenea. Streptasters of Thenea abyssorum Koltun, 1964 [ZMBN 85228]. A. Plesiaster; B. Spirasters; Streptasters of Thenea levis von Lendenfeld, 1907 [ZMBN 85230]. C. Plesiaster. Same scale as A; D. spirasters. Streptasters of Thenea muricata (Bowerbank, 1858) [ZMBN 85231]. E. Plesiaster; F. Spiraster; Streptasters of Thenea valdiviae von Lendenfeld, 1907. G. Plesiaster [ZMBN 85257]; H. Spirasters. Same scale as E. [ZMBN 85233]." httpUri="https://zenodo.org/record/280608/files/figure.png" pageId="44" pageNumber="45">Fig. 18</figureCitation>
H), spined, length: 11
<emphasis id="1CCAEAB3DC45FF9AFD08F923FD65F8EA" box="[684,733,1770,1794]" italics="true" pageId="44" pageNumber="45">18.2</emphasis>
27.5 µm. (f) plesiasters (
<figureCitation id="B6852A24DC45FF9AFBA3F923FBE2F8EA" box="[1031,1114,1770,1794]" captionStart="FIGURE 18" captionStartId="34.[151,250,1922,1945]" captionTargetBox="[216,1384,193,1910]" captionTargetId="figure@34.[216,1385,193,1911]" captionTargetPageId="34" captionText="FIGURE 18. Streptasters of Norwegian species of Thenea. Streptasters of Thenea abyssorum Koltun, 1964 [ZMBN 85228]. A. Plesiaster; B. Spirasters; Streptasters of Thenea levis von Lendenfeld, 1907 [ZMBN 85230]. C. Plesiaster. Same scale as A; D. spirasters. Streptasters of Thenea muricata (Bowerbank, 1858) [ZMBN 85231]. E. Plesiaster; F. Spiraster; Streptasters of Thenea valdiviae von Lendenfeld, 1907. G. Plesiaster [ZMBN 85257]; H. Spirasters. Same scale as E. [ZMBN 85233]." httpUri="https://zenodo.org/record/280608/files/figure.png" pageId="44" pageNumber="45">Fig. 18</figureCitation>
G), few, 46 actines, faintly spined, actine length: 18.5
<emphasis id="1CCAEAB3DC45FF9AFE68F8C7FE45F8CE" box="[460,509,1806,1830]" italics="true" pageId="44" pageNumber="45">41.4</emphasis>
75 µm, actine width: 3
<emphasis id="1CCAEAB3DC45FF9AFCB2F8C7FC81F8CE" box="[790,825,1806,1830]" italics="true" pageId="44" pageNumber="45">7.4</emphasis>
12.5 µm (N=23).
</paragraph>
<paragraph id="2E0136A1DC45FF9AFF63F8FBFCDAF87A" blockId="44.[151,1436,1518,2008]" pageId="44" pageNumber="45">
<emphasis id="1CCAEAB3DC45FF9AFF63F8FBFEDEF8A2" box="[199,358,1842,1866]" italics="true" pageId="44" pageNumber="45">Reproduction.</emphasis>
Oocytes in vitellogenetic phase were observed in the thick sections of
<collectionCode id="48AFAE64DC45FF9AFB2EF8FAFB63F8A2" box="[1162,1243,1843,1866]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="44" pageNumber="45">ZMBN</collectionCode>
85233, collected in mid
<date id="5A001061DC45FF9AFF4AF89FFE3AF886" box="[238,386,1878,1902]" pageId="44" pageNumber="45" value="2007-08">August 2007</date>
. Subglobular whitish buds were occasionally observed on specimens from the Røst reef (
<figureCitation id="B6852A24DC45FF9AFACBF89FFF0CF87A" captionStart="FIGURE 24" captionStartId="44.[151,250,1328,1351]" captionTargetBox="[176,1436,406,1304]" captionTargetId="figure@44.[151,1436,375,1304]" captionTargetPageId="44" captionText="FIGURE 24. Thenea valdiviae von Lendenfeld, 1907. A. Whole specimen from Barents Sea, top view. Note the sieve on the oscule [ZMBN 85233]. Scale: 2 cm; B. Whole specimen, side view showing the poral area, also with sieve [ZMBN 85233]; C. Whole specimen from Freisfjorden (Møre), top view. Note the sieve on the oscule. Scale: 1 cm [ZMBN 85256]; D. Top surface of a young specimen covered with buds. p: pores. os: oscule. [ZMBN 85257]; E. Thick section showing the skeletal organization. Scale: 1 mm [ZMBN 85233]." httpUri="https://zenodo.org/record/280614/files/figure.png" pageId="44" pageNumber="45">Fig. 24</figureCitation>
D) and from the Schultz Massive Seamount (
<collectingCountry id="56A97631DC45FF9AFD0FF8B3FC99F87A" box="[683,801,1914,1938]" name="Greenland" pageId="44" pageNumber="45">Greenland</collectingCountry>
Sea).
</paragraph>
</subSubSection>
<subSubSection id="66A4652ADC45FF9BFF63F857FCDEFF47" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="distribution">
<paragraph id="2E0136A1DC45FF9AFF63F857FC74F830" blockId="44.[151,1436,1518,2008]" pageId="44" pageNumber="45">
<emphasis id="1CCAEAB3DC45FF9AFF63F857FEE8F85E" box="[199,336,1950,1974]" italics="true" pageId="44" pageNumber="45">Distribution</emphasis>
.
<collectingCountry id="56A97631DC45FF9AFEC1F857FE64F85E" box="[357,476,1950,1974]" name="Greenland" pageId="44" pageNumber="45">Greenland</collectingCountry>
,
<collectingCountry id="56A97631DC45FF9AFE4AF857FDFBF85E" box="[494,579,1950,1974]" name="Iceland" pageId="44" pageNumber="45">Iceland</collectingCountry>
, Faeroe Islands,
<collectingCountry id="56A97631DC45FF9AFCB0F856FCD7F85F" box="[788,879,1951,1975]" name="Norway" pageId="44" pageNumber="45">Norway</collectingCountry>
, Norwegian Sea,
<collectingCountry id="56A97631DC45FF9AFBEDF857FB06F85E" box="[1097,1214,1950,1974]" name="Greenland" pageId="44" pageNumber="45">Greenland</collectingCountry>
Sea, Barents Sea, Spitzbergen (
<bibRefCitation id="4A2F4B50DC45FF9AFE8EF809FDEDF830" author="Steenstrup" box="[298,597,1984,2008]" pageId="44" pageNumber="45" refString="Steenstrup, E. &amp; Tendal, O. S. (1982) The genus Thenea (Porifera, Demospongiae, Choristida) in the Norwegian Sea and adjacent waters; an annotated key. Sarsia, 67, 259 - 268." type="journal article" year="1982">Steenstrup &amp; Tendal 1982</bibRefCitation>
); SW of
<collectingCountry id="56A97631DC45FF9AFD1CF809FCBFF830" box="[696,775,1984,2008]" name="Ireland" pageId="44" pageNumber="45">Ireland</collectingCountry>
(
<bibRefCitation id="4A2F4B50DC45FF9AFCB2F809FC07F830" author="Stephens" box="[790,959,1984,2008]" pageId="44" pageNumber="45" refString="Stephens, J. (1915) Sponges of the Coasts of Ireland. I. - The Triaxonia and part of the Tetraxonida. Fisheries, Ireland Scientific Investigations, 1914, 1 - 43, pls I - V." type="journal article" year="1915">Stephens 1915</bibRefCitation>
).
</paragraph>
<paragraph id="2E0136A1DC44FF9BFF63FF51FCDEFF47" blockId="45.[151,1437,151,905]" box="[199,870,151,176]" pageId="45" pageNumber="46">
<emphasis id="1CCAEAB3DC44FF9BFF63FF51FEB3FF58" box="[199,267,152,176]" italics="true" pageId="45" pageNumber="46">Depth</emphasis>
.
<quantity id="E9469B44DC44FF9BFEBDFF5EFE1EFF47" box="[281,422,151,175]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.9" metricValueMin="0.1" pageId="45" pageNumber="46" unit="m" value="1000.0" valueMax="1900.0" valueMin="100.0">1001900 m</quantity>
(
<bibRefCitation id="4A2F4B50DC44FF9BFE11FF5EFCE2FF47" author="Steenstrup" box="[437,858,151,175]" pageId="45" pageNumber="46" refString="Steenstrup, E. &amp; Tendal, O. S. (1982) The genus Thenea (Porifera, Demospongiae, Choristida) in the Norwegian Sea and adjacent waters; an annotated key. Sarsia, 67, 259 - 268." type="journal article" year="1982">Steenstrup &amp; Tendal 1982; this study</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="66A4652ADC44FF9BFF63FF74FE50FC61" pageId="45" pageNumber="46" type="discussion">
<paragraph id="2E0136A1DC44FF9BFF63FF74FC3FFEF1" blockId="45.[151,1437,151,905]" pageId="45" pageNumber="46">
<emphasis id="1CCAEAB3DC44FF9BFF63FF74FEFAFF3C" box="[199,322,189,212]" italics="true" pageId="45" pageNumber="46">Discussion</emphasis>
. The specimens from the Korsfjord have more anatriaenes and regular dichotriaenes than ZMBN 85233 from the Barents Sea. Protriaenes are very common in ZMBN 85234 from the
<collectingCountry id="56A97631DC44FF9BFBE8FF17FB79FF1E" box="[1100,1217,222,246]" name="Greenland" pageId="45" pageNumber="46">Greenland</collectingCountry>
Sea. Plesiasters are rare (ZMBN 85233) to moderately present (ZMBN 85255, 85257).
</paragraph>
<paragraph id="2E0136A1DC44FF9BFF33FEECFE49FE4D" blockId="45.[151,1437,151,905]" pageId="45" pageNumber="46">
<taxonomicName id="E9BE4D22DC44FF9BFF33FEECFEA6FED4" box="[151,286,292,316]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFF33FEECFEA6FED4" box="[151,286,292,316]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
is known to be at least dimorphic and our Norwegian specimens correspond to the arctic form described by
<bibRefCitation id="4A2F4B50DC44FF9BFE95FE8FFDCAFEB6" author="Steenstrup" box="[305,626,326,350]" pageId="45" pageNumber="46" refString="Steenstrup, E. &amp; Tendal, O. S. (1982) The genus Thenea (Porifera, Demospongiae, Choristida) in the Norwegian Sea and adjacent waters; an annotated key. Sarsia, 67, 259 - 268." type="journal article" year="1982">Steenstrup &amp; Tendal (1982)</bibRefCitation>
. Our new records show that
<taxonomicName id="E9BE4D22DC44FF9BFC64FE81FBFDFEB6" box="[960,1093,326,351]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFC64FE81FBFDFEB6" box="[960,1093,326,351]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
must be present all along the Norwegian coast whereas, to our knowledge,
<taxonomicName id="E9BE4D22DC44FF9BFD03FEA3FC91FE69" box="[679,809,362,385]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC44FF9BFD03FEA3FC91FE69" box="[679,809,362,385]" italics="true" pageId="45" pageNumber="46">T. muricata</emphasis>
</taxonomicName>
has not been formally identified north of Trollsteinen (
<geoCoordinate id="4B8A5066DC44FF9BFF3BFE45FEBCFE4C" box="[159,260,396,420]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="64.583336">64°35N</geoCoordinate>
), except for the
<typeStatus id="F1058803DC44FF9BFE1DFE44FE55FE4D" box="[441,493,397,421]" pageId="45" pageNumber="46">type</typeStatus>
.
</paragraph>
<paragraph id="2E0136A1DC44FF9BFF63FE67FE8AFD7E" blockId="45.[151,1437,151,905]" pageId="45" pageNumber="46">
The easiest way to discriminate adult specimens is to look at the oscule morphology: large cribriporal irregular shaped oscules with a surrounding fringe in
<taxonomicName id="E9BE4D22DC44FF9BFD39FE1BFC9AFE01" box="[669,802,465,489]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFD39FE1BFC9AFE01" box="[669,802,465,489]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
, small uniporal round “naked” oscules in
<taxonomicName id="E9BE4D22DC44FF9BFAB5FE1BFA2BFE01" box="[1297,1427,466,489]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC44FF9BFAB5FE1BFA2BFE01" box="[1297,1427,466,489]" italics="true" pageId="45" pageNumber="46">T. muricata</emphasis>
</taxonomicName>
. Young specimens are, without genetic markers, very difficult to differentiate since they both have opposite oscule/ pore areas on their sides. As for the spicules, they are fairly similar in both species (
<tableCitation id="633C031ADC44FF9BFBF8FDDFFB0CFDC6" box="[1116,1204,534,558]" captionStart="TABLE 5" captionStartId="42.[151,239,979,1003]" captionTargetBox="[197,1424,1176,1971]" captionTargetPageId="42" captionText="TABLE 5. Depth, individual spicule dimensions of streptasters (in µm) for specimens of Thenea valdiviae, Thenea muricata, Thenea schmidti and Thenea bojeadori. Means are in bold; other values are ranges; n = 30 unless stated otherwise between parentheses. - = not referred." httpUri="http://table.plazi.org/id/7AC16629DC43FF9CFF33FC1AFD20FBC6" pageId="45" pageNumber="46" tableUuid="7AC16629DC43FF9CFF33FC1AFD20FBC6">Table 5</tableCitation>
). There seems to be more metasters in
<taxonomicName id="E9BE4D22DC44FF9BFECFFDF3FE54FDB9" box="[363,492,570,593]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC44FF9BFECFFDF3FE54FDB9" box="[363,492,570,593]" italics="true" pageId="45" pageNumber="46">T. muricata</emphasis>
</taxonomicName>
and
<taxonomicName id="E9BE4D22DC44FF9BFD8CFDF3FD1BFDB9" box="[552,675,569,593]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="schmidti">
<emphasis id="1CCAEAB3DC44FF9BFD8CFDF3FD1BFDB9" box="[552,675,569,593]" italics="true" pageId="45" pageNumber="46">T. schmidti</emphasis>
</taxonomicName>
than in
<taxonomicName id="E9BE4D22DC44FF9BFCA2FDF3FC32FDB9" box="[774,906,569,593]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFCA2FDF3FC32FDB9" box="[774,906,569,593]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
, but this need to be confirmed. Moreover, the rarity of plesiasters in
<taxonomicName id="E9BE4D22DC44FF9BFE3FFD94FD98FD9C" box="[411,544,604,628]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFE3FFD94FD98FD9C" box="[411,544,604,628]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
seems to be a general rule, observed by others (
<bibRefCitation id="4A2F4B50DC44FF9BFBF4FD95FB40FD9C" author="Vosmaer" box="[1104,1272,604,628]" pageId="45" pageNumber="46" refString="Vosmaer, G. C. J. (1885) The Sponges of the ' Willem Barents' Expedition 1880 and 1881. Bijdragen tot de Dierkunde, 12, 1 - 47, pls I - V." type="journal article" year="1885">Vosmaer 1885</bibRefCitation>
;
<bibRefCitation id="4A2F4B50DC44FF9BFAA3FD95FE9EFD7E" author="Steenstrup" pageId="45" pageNumber="46" refString="Steenstrup, E. &amp; Tendal, O. S. (1982) The genus Thenea (Porifera, Demospongiae, Choristida) in the Norwegian Sea and adjacent waters; an annotated key. Sarsia, 67, 259 - 268." type="journal article" year="1982">Steenstrup &amp; Tendal 1982</bibRefCitation>
).
</paragraph>
<paragraph id="2E0136A1DC44FF9BFF63FD6BFE50FC61" blockId="45.[151,1437,151,905]" pageId="45" pageNumber="46">
As sister-species, Norwegian
<taxonomicName id="E9BE4D22DC44FF9BFDB1FD6BFD21FD51" box="[533,665,673,697]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFDB1FD6BFD21FD51" box="[533,665,673,697]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
are genetically clearly separated from
<taxonomicName id="E9BE4D22DC44FF9BFBF4FD6BFB77FD51" box="[1104,1231,674,697]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC44FF9BFBF4FD6BFB77FD51" box="[1104,1231,674,697]" italics="true" pageId="45" pageNumber="46">T. muricata</emphasis>
</taxonomicName>
(
<figureCitation id="B6852A24DC44FF9BFB46FD68FA8EFD51" box="[1250,1334,673,697]" captionStart="FIGURE 25" captionStartId="45.[151,250,1900,1923]" captionTargetBox="[228,1377,963,1881]" captionTargetId="figure@45.[215,1384,959,1885]" captionTargetPageId="45" captionText="FIGURE 25. Simplified molecular phylogeny of the Astrophorida focusing on the relationships within the Theneidae, modified from Cárdenas et al. (2011, Fig. 2). Maximum-likelihood tree made from concatenated sequences of COI (Folmer fragment) and 28 S (C 1 D 2 domains). Bootstrap nodal support values&gt; 50 % are given above the nodes (2,000 replicates)." httpUri="https://zenodo.org/record/280615/files/figure.png" pageId="45" pageNumber="46">Fig. 25</figureCitation>
), but our specimen from the
<collectingCountry id="56A97631DC44FF9BFECAFD0DFE5BFD34" box="[366,483,708,732]" name="Greenland" pageId="45" pageNumber="46">Greenland</collectingCountry>
Sea seamount is somewhat different (ZMBN 85234): it has a COI sequence identical to
<taxonomicName id="E9BE4D22DC44FF9BFF10FD21FE8AFD16" box="[180,306,743,767]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC44FF9BFF10FD21FE8AFD16" box="[180,306,743,767]" italics="true" pageId="45" pageNumber="46">T. muricata</emphasis>
</taxonomicName>
, and on the other hand a 28S sequence identical to
<taxonomicName id="E9BE4D22DC44FF9BFCD0FD21FC4EFD16" box="[884,1014,742,767]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFCD0FD21FC4EFD16" box="[884,1014,742,767]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
. This specimen, and others examined from the same seamount are all very hispid (hairy-like), with cribriporal oscule and pores on opposite sides, and rare plesiasters. So they are not morphologically different from the Norwegian
<taxonomicName id="E9BE4D22DC44FF9BFBBEFCE4FB25FCAC" box="[1050,1181,812,836]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="valdiviae">
<emphasis id="1CCAEAB3DC44FF9BFBBEFCE4FB25FCAC" box="[1050,1181,812,836]" italics="true" pageId="45" pageNumber="46">T. valdiviae</emphasis>
</taxonomicName>
, except maybe for the larger spirasters (
<tableCitation id="633C031ADC44FF9BFEFAFC87FE00FC8E" box="[350,440,846,870]" captionStart="TABLE 5" captionStartId="42.[151,239,979,1003]" captionTargetBox="[197,1424,1176,1971]" captionTargetPageId="42" captionText="TABLE 5. Depth, individual spicule dimensions of streptasters (in µm) for specimens of Thenea valdiviae, Thenea muricata, Thenea schmidti and Thenea bojeadori. Means are in bold; other values are ranges; n = 30 unless stated otherwise between parentheses. - = not referred." httpUri="http://table.plazi.org/id/7AC16629DC43FF9CFF33FC1AFD20FBC6" pageId="45" pageNumber="46" tableUuid="7AC16629DC43FF9CFF33FC1AFD20FBC6">Table 5</tableCitation>
). This shared COI could be the result of ongoing hybridization with
<taxonomicName id="E9BE4D22DC44FF9BFB40FC99FADFFC8E" box="[1252,1383,847,871]" class="Demospongiae" family="Theneidae" genus="Thenea" kingdom="Animalia" order="Astrophorida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="muricata">
<emphasis id="1CCAEAB3DC44FF9BFB40FC99FADFFC8E" box="[1252,1383,847,871]" italics="true" pageId="45" pageNumber="46">T. muricata</emphasis>
</taxonomicName>
and deserves to be further studied.
</paragraph>
</subSubSection>
</treatment>
</document>