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<document id="73AA19F13D3FF1213701F7D2C2390439" ID-CLB-Dataset="284154" ID-DOI="10.5852/ejt.2023.909.2351" ID-GBIF-Dataset="d0cc9c9f-6e84-4a88-a82f-96565be36a3d" ID-ISSN="2118-9773" ID-Zenodo-Dep="10462860" ID-ZooBank="A48BD2B3-DC40-45BD-9968-F04890A1C5C5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701248827785" checkinUser="plazi" docAuthor="Huber, Bernhard A., Meng, Guanliang, Dupérré, Nadine, Astrin, Jonas &amp; Herrera, Mauricio" docDate="2023" docId="8A6F2C53FFC4A645FD96FB3061A1FCD6" docLanguage="en" docName="EJT.2023.909.1-63.pdf" docOrigin="European Journal of Taxonomy 909" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2351/10271" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.33:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="32" docTitle="Priscula pastaza Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera 2023, sp. nov." docType="treatment" docUuid="DADCE77E-4843-45FC-B14E-B20C35764BE3" docUuidSource="ZooBank" docVersion="3" lastPageNumber="42" masterDocId="7656542BFFE1A66CFFC8FF88626EFF96" masterDocTitle="Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="38" updateTime="1704471815894" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="92F0929EE499EB7EF662D3D06D80EFB8">Andean giants: Priscula spiders from Ecuador, with notes on species groups and egg-sac troglomorphism (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="FC30860BC4D99EA6FD9CF0A55CDC55BF">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="3B0AD602968960CFC30610A0487008E2">Meng, Guanliang</mods:namePart>
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<mods:affiliation id="B36D95F19E7B1BDDCD5AAB456ECB29EC">Zoological Research Museum Alexander Koenig, LIB, Bonn, Germany.</mods:affiliation>
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<mods:namePart id="B43EE28A3955DE87B0A2D3BC8C3156C6">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="AA0C4AF656E721F9A88D9EF6DCBB7A36">Museum of Nature Hamburg, LIB, Hamburg, Germany.</mods:affiliation>
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<mods:namePart id="335D5039C88356B6440CE9E34B363020">Astrin, Jonas</mods:namePart>
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<mods:affiliation id="CF8E9ED38372E03692B221DFE088A325">Instituto Nacional de Biodiversidad, Sección Invertebrados, Quito, Ecuador.</mods:affiliation>
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<taxonomicName id="C5C6E6C6FFC4A649FD96FB30611AFB44" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[606,884,1208,1235]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pastaza" status="sp. nov.">
<emphasis id="30B24157FFC4A649FD96FB30614AFB45" bold="true" box="[606,804,1208,1235]" italics="true" pageId="37" pageNumber="38">Priscula pastaza</emphasis>
Huber
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<paragraph id="02799D45FFC4A649FE6AFB7766FFFA8F" blockId="37.[361,1226,1208,1305]" box="[418,1169,1279,1305]" pageId="37" pageNumber="38">
<figureCitation id="9AFD81C0FFC4A649FE6AFB77604BFA8F" box="[418,549,1279,1305]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[183,1404,633,1854]" captionTargetId="figure-130@12.[183,1404,633,1854]" captionTargetPageId="12" captionText="Fig. 6. Priscula spp., binghamae and andinensis groups, live specimens. A–B. P. pastaza Huber sp. nov.; male and female with egg-sac from Cavernas del Anzu Forest Reserve. C–D. P. bonita Huber sp. nov.; male from La Bonita-Santa Barbara and female with egg-sac from Gruta de los Tayos. E–F. P. lumbaqui Huber sp. nov.; male and female from near Lumbaqui." figureDoi="http://doi.org/10.5281/zenodo.10462882" httpUri="https://zenodo.org/record/10462882/files/figure.png" pageId="37" pageNumber="38">Figs 6A–B</figureCitation>
, 
<figureCitation id="9AFD81C0FFC4A649FDF9FB776013FA8F" box="[561,637,1279,1305]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="38.[191,234,1963,1989]" captionStartId-1="39.[189,232,1928,1954]" captionStartId-2="40.[190,233,1928,1954]" captionStartId-3="41.[189,232,1860,1886]" captionTargetBox-0="[183,1404,1160,1885]" captionTargetBox-1="[199,1395,472,1826]" captionTargetBox-2="[191,1393,1304,1888]" captionTargetBox-3="[183,1404,918,1784]" captionTargetId-0="figure-280@38.[183,1404,1160,1885]" captionTargetId-1="figure-98@39.[186,1401,472,1849]" captionTargetId-2="figure-350@40.[186,1401,1296,1889]" captionTargetId-3="figure-204@41.[183,1404,918,1784]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionTargetPageId-3="41" captionText-0="Fig. 29. Priscula pastaza Huber sp. nov.; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101; left male palp, prolateral, dorsal, and retrolateral views. Scale line: 0.5 mm." captionText-1="Fig. 30. Priscula pastaza Huber sp. nov.; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views; asterisks: whitish protruding area. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Scale lines: 0.5 mm." captionText-2="Fig. 31. Priscula pastaza Huber sp. nov.; male and female paratypes from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." captionText-3="Fig. 32. Priscula pastaza Huber sp. nov.; female paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–B. Abdomen, ventral and lateral views. C–D. Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10462932" figureDoi-1="http://doi.org/10.5281/zenodo.10462934" figureDoi-2="http://doi.org/10.5281/zenodo.10462936" figureDoi-3="http://doi.org/10.5281/zenodo.10462938" httpUri-0="https://zenodo.org/record/10462932/files/figure.png" httpUri-1="https://zenodo.org/record/10462934/files/figure.png" httpUri-2="https://zenodo.org/record/10462936/files/figure.png" httpUri-3="https://zenodo.org/record/10462938/files/figure.png" pageId="37" pageNumber="38">29–32</figureCitation>
, 
<figureCitation id="9AFD81C0FFC4A649FD41FB77616AFA8F" box="[649,772,1279,1305]" captionStart="Fig" captionStartId="51.[189,233,1773,1799]" captionTargetBox="[183,1404,265,1703]" captionTargetId="figure-86@51.[183,1404,265,1703]" captionTargetPageId="51" captionText="Fig. 41. Priscula spp., SEM photographs of female ocular areas and spinnerets. A–B. Ocular areas, frontal views; Priscula pastaza Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C–E. ALS, P. pastaza (C), P. esmeraldas Huber sp. nov. (D), and P. lumbaqui sp. nov. (E). F–H. Distinctive structures in front of spinnerets (in place of colulus); P. chapintza Huber sp. nov. (F), P. esmeraldas (G), and P. pastaza (H). Scale lines: 100 µm (A–B); 10 µm (C–G); 2 µm (H)." figureDoi="http://doi.org/10.5281/zenodo.10462956" httpUri="https://zenodo.org/record/10462956/files/figure.png" pageId="37" pageNumber="38">41A, C, H</figureCitation>
, 
<figureCitation id="9AFD81C0FFC4A649FCD8FB7761E4FA8F" box="[784,906,1279,1305]" captionStart="Fig" captionStartId="52.[189,232,1775,1801]" captionTargetBox="[181,1402,265,1699]" captionTargetId="figure-113@52.[181,1402,265,1699]" captionTargetPageId="52" captionText="Fig. 42. Priscula spp., SEM photographs of male pedipalps. A–C. Tip of left procursus in dorsal view (A) and hair-like processes on its prolateral (B) and retrolateral (C) sides; P. esmeraldas Huber sp. nov. D–E. Tip of left procursus in retrolateral view (D) and hair-like processes in more detail (E); P. pastaza Huber sp. nov. F–G. Left procursus and bulb of P. bonita Huber sp. nov., showing position of tarsal organ (arrow in F) and position of sperm duct opening (arrow in G). H. Sperm duct opening (arrow) on bulbal process in P. pastaza. Scale lines: 100 µm (A, F–G); 10 µm (B–C, E); 20 µm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.10462958" httpUri="https://zenodo.org/record/10462958/files/figure.png" pageId="37" pageNumber="38">42D–E, H</figureCitation>
, 
<figureCitation id="9AFD81C0FFC4A649FC5EFB77667FFA8F" box="[918,1041,1279,1305]" captionStart="Fig" captionStartId="53.[189,233,1771,1797]" captionTargetBox="[183,1404,265,1697]" captionTargetId="figure-98@53.[183,1404,265,1697]" captionTargetPageId="53" captionText="Fig. 43. Priscula spp., SEM photographs of female palps and of male and female tarsal organs. A–B. Tips of female palps in dorsal views; Priscula esmeraldas Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C. Male palpal tarsal organ, P. pastaza Huber sp. nov. D. Female palpal tarsal organ, P. lumbaqui. E–H. Female tarsal organs on leg 1 (E, P. lumbaqui), leg 2 (F, P. esmeraldas), leg 3 (G, P. pastaza), and leg 4 (H, P. pastaza). Scale lines: 10 µm (A, C–F); 20 µm (B); 2 µm (G–H)." figureDoi="http://doi.org/10.5281/zenodo.10462960" httpUri="https://zenodo.org/record/10462960/files/figure.png" pageId="37" pageNumber="38">43C, G–H</figureCitation>
, 
<figureCitation id="9AFD81C0FFC4A649FBD5FB7766FFFA8F" box="[1053,1169,1279,1305]" captionStart="Fig" captionStartId="54.[189,231,1774,1800]" captionTargetBox="[183,1404,265,1696]" captionTargetId="figure-96@54.[183,1404,265,1696]" captionTargetPageId="54" captionText="Fig. 44. Priscula spp., SEM photographs of female legs. A–B. Oval plates of unknown function, in Priscula lumbaqui Huber sp. nov. (A, metatarsus 4) and P. pastaza Huber sp. nov. (B, metatarsus 4). C–F. Tarsal claws of leg 1 (C, P. pastaza), leg 2 (D, P. pastaza), leg 3 (E, P. bonita Huber sp. nov.), and leg 4 (F, P. pastaza). G. Microstructure of claw tines (P. chapintza Huber sp. nov.). H. Basis of mechanoreceptive hair (P. chapintza; metatarsus 4), showing distal bulge (on right side). Scale lines: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10462964" httpUri="https://zenodo.org/record/10462964/files/figure.png" pageId="37" pageNumber="38">44B–D, F</figureCitation>
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<emphasis id="30B24157FFC4A649FF75FAC86358FACC" bold="true" box="[189,310,1344,1370]" pageId="37" pageNumber="38">Diagnosis</emphasis>
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<paragraph id="02799D45FFC4A649FF75FAE763D2F9AF" blockId="37.[189,1399,1391,1593]" pageId="37" pageNumber="38">
Distinguished from known congeners by details of procursus (
<figureCitation id="9AFD81C0FFC4A649FC29FAE76614FA1F" box="[993,1146,1391,1417]" captionStart="Fig" captionStartId="39.[189,232,1928,1954]" captionTargetBox="[199,1395,472,1826]" captionTargetId="figure-98@39.[186,1401,472,1849]" captionTargetPageId="39" captionText="Fig. 30. Priscula pastaza Huber sp. nov.; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views; asterisks: whitish protruding area. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462934" httpUri="https://zenodo.org/record/10462934/files/figure.png" pageId="37" pageNumber="38">Fig. 30A–C</figureCitation>
; tip with two large membranous elements: wide dorsal flap and long ventral flap with pointed tip), genital bulb (
<figureCitation id="9AFD81C0FFC4A649FB36FA1A62BFFA46" captionStart="Fig" captionStartId="39.[189,232,1928,1954]" captionTargetBox="[199,1395,472,1826]" captionTargetId="figure-98@39.[186,1401,472,1849]" captionTargetPageId="39" captionText="Fig. 30. Priscula pastaza Huber sp. nov.; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views; asterisks: whitish protruding area. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462934" httpUri="https://zenodo.org/record/10462934/files/figure.png" pageId="37" pageNumber="38">Fig. 30D– F</figureCitation>
; main bulbal process weakly curved with obtuse tip), epigynum (
<figureCitation id="9AFD81C0FFC4A649FC2FFA3D6614FA59" box="[999,1146,1461,1487]" captionStart="Fig" captionStartId="41.[189,232,1860,1886]" captionTargetBox="[183,1404,918,1784]" captionTargetId="figure-204@41.[183,1404,918,1784]" captionTargetPageId="41" captionText="Fig. 32. Priscula pastaza Huber sp. nov.; female paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–B. Abdomen, ventral and lateral views. C–D. Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462938" httpUri="https://zenodo.org/record/10462938/files/figure.png" pageId="37" pageNumber="38">Fig. 32A–B</figureCitation>
; semicircular, longer than in putatively closest known relatives, 
<taxonomicName id="C5C6E6C6FFC4A649FD1DFA516111FA64" authority="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera, 2023" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[725,895,1496,1522]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="esmeraldas" status="sp. nov.">
<emphasis id="30B24157FFC4A649FD1DFA516111FA64" box="[725,895,1496,1522]" italics="true" pageId="37" pageNumber="38">P. esmeraldas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFC4A649FC43FA516185FA65" box="[907,1003,1497,1523]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="C5C6E6C6FFC4A649FBE5FA5166ACFA64" authority="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera, 2023" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[1069,1218,1496,1522]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="chapintza" status="sp. nov.">
<emphasis id="30B24157FFC4A649FBE5FA5166ACFA64" box="[1069,1218,1496,1522]" italics="true" pageId="37" pageNumber="38">P. chapintza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFC4A649FB06FA51675CFA64" box="[1230,1330,1496,1523]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
), and female internal genitalia (
<figureCitation id="9AFD81C0FFC4A649FE33FA736004F980" box="[507,618,1531,1558]" captionStart="Fig" captionStartId="40.[190,233,1928,1954]" captionTargetBox="[191,1393,1304,1888]" captionTargetId="figure-350@40.[186,1401,1296,1889]" captionTargetPageId="40" captionText="Fig. 31. Priscula pastaza Huber sp. nov.; male and female paratypes from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462936" httpUri="https://zenodo.org/record/10462936/files/figure.png" pageId="37" pageNumber="38">Fig. 31C</figureCitation>
; round pore plates similar to 
<taxonomicName id="C5C6E6C6FFC4A649FC13FA746601F983" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[987,1135,1531,1557]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="chapintza">
<emphasis id="30B24157FFC4A649FC13FA746601F983" box="[987,1135,1531,1557]" italics="true" pageId="37" pageNumber="38">P. chapintza</emphasis>
</taxonomicName>
but not connected to sclerites posteriorly).
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFC4A64AFF75F9E86070FDB2" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="02799D45FFC4A649FF75F9E86302F9EC" blockId="37.[189,364,1632,1658]" box="[189,364,1632,1658]" pageId="37" pageNumber="38">
<heading id="59312A29FFC4A649FF75F9E86302F9EC" bold="true" box="[189,364,1632,1658]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="30B24157FFC4A649FF75F9E86302F9EC" bold="true" box="[189,364,1632,1658]" pageId="37" pageNumber="38">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFC4A649FF22F907636EF884" blockId="37.[189,1399,1679,1810]" pageId="37" pageNumber="38">
<materialsCitation id="B2AE9718FFC4A649FF22F907636EF884" collectedFrom="in cave" collectingDate="2021-09-25" collectionCode="MECN" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" county="Cavernas del Anzu Forest Reserve" elevation="1160" latitude="-1.4067" location="Caverna de los Continentes" longLatPrecision="7" longitude="-78.0449" municipality="Caverna" pageId="37" pageNumber="38" specimenCode="MECN-ARAC- 37-T" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="holotype">
<emphasis id="30B24157FFC4A649FF22F9076332F93F" bold="true" box="[234,348,1679,1705]" pageId="37" pageNumber="38">
<typeStatus id="DD7D23E7FFC4A649FF22F9076332F93F" box="[234,348,1679,1705]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7AD1DDD5FFC4A649FF75F93A6325F95A" box="[189,331,1714,1740]" name="Ecuador" pageId="37" pageNumber="38">ECUADOR</collectingCountry>
– 
<emphasis id="30B24157FFC4A649FEA5F93A63A3F95A" bold="true" box="[365,461,1714,1740]" pageId="37" pageNumber="38">
<collectingRegion id="C00253A7FFC4A649FEA5F93A63A3F95A" box="[365,461,1714,1740]" country="Ecuador" name="Pastaza" pageId="37" pageNumber="38">Pastaza</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFC4A649FE22F93A606DF95A" box="[490,515,1714,1740]" count="1" pageId="37" pageNumber="38" type="male">♂</specimenCount>
; 
<collectingCounty id="EB18E5C9FFC4A649FDDAF93A61DFF95A" box="[530,945,1714,1740]" pageId="37" pageNumber="38">Cavernas del Anzu Forest Reserve</collectingCounty>
, 
<location id="0719CB9EFFC4A649FC08F93A6762F95A" LSID="urn:lsid:plazi:treatment:8A6F2C53FFC4A645FD96FB3061A1FCD6:0719CB9EFFC4A649FC08F93A6762F95A" box="[960,1292,1714,1740]" country="Ecuador" county="Cavernas del Anzu Forest Reserve" latitude="-1.4067" longLatPrecision="7" longitude="-78.0449" municipality="Caverna" name="Caverna de los Continentes" pageId="37" pageNumber="38" stateProvince="Pastaza">Caverna de los Continentes</location>
; 
<geoCoordinate id="67F2FB82FFC4A649FAD2F93A62A1F979" degrees="1.4067" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="-1.4067">1.4067° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFC4A649FF14F95D6302F966" box="[220,364,1749,1776]" degrees="78.0449" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="5" value="-78.0449">78.0449° W</geoCoordinate>
; 
<elevation id="89EB7A76FFC4A649FEB1F95D607FF979" box="[377,529,1749,1776]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="37" pageNumber="38" unit="m" value="1160.0">
<quantity id="C53E30A0FFC4A649FEB1F95D63BFF966" box="[377,465,1749,1776]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="37" pageNumber="38" unit="m" value="1160.0">1160 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFC4A649FDD6F95D60D2F979" box="[542,700,1749,1776]" pageId="37" pageNumber="38" value="2021-09-25">
<collectingDate id="663C426DFFC4A649FDD6F95D60D2F979" box="[542,700,1749,1776]" pageId="37" pageNumber="38" value="2021-09-25">25 Sep. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFC4A649FD00F95D613EF979" box="[712,848,1749,1775]" pageId="37" pageNumber="38">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFC4A649FC42F95E6663F966" box="[906,1037,1750,1776]" pageId="37" pageNumber="38">M. Herrera</collectorName>
leg.; 
<collectedFrom id="B1DADEB8FFC4A649FB87F95D66C8F966" box="[1103,1190,1749,1776]" pageId="37" pageNumber="38">in cave</collectedFrom>
; 
<specimenCode id="5260353EFFC4A649FB7BF95D6292F884" pageId="37" pageNumber="38">MECN–ARAC– 37–T</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="02799D45FFC4A649FF22F8B260D6F80C" blockId="37.[189,1398,1850,1947]" pageId="37" pageNumber="38">
<materialsCitation id="B2AE9718FFC4A649FF22F8B260DAF80C" collectedFrom="in cave" collectingDate="2021-09-25" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" county="Cavernas del Anzu Forest Reserve" elevation="1160" latitude="-1.4067" location="Pastaza" longLatPrecision="7" longitude="-78.0449" municipality="Caverna" pageId="37" pageNumber="38" specimenCode="MECN-ARAC-38-T" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<emphasis id="30B24157FFC4A649FF22F8B26309F8C2" bold="true" box="[234,359,1850,1876]" pageId="37" pageNumber="38">
<typeStatus id="DD7D23E7FFC4A649FF22F8B26309F8C2" box="[234,359,1850,1876]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="7AD1DDD5FFC4A649FF75F8D56325F8E1" box="[189,331,1885,1911]" name="Ecuador" pageId="37" pageNumber="38">ECUADOR</collectingCountry>
– 
<emphasis id="30B24157FFC4A649FEA2F8D563A4F8E1" bold="true" box="[362,458,1885,1911]" pageId="37" pageNumber="38">
<collectingRegion id="C00253A7FFC4A649FEA2F8D563A4F8E1" box="[362,458,1885,1911]" country="Ecuador" name="Pastaza" pageId="37" pageNumber="38">Pastaza</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFC4A649FE2EF8D5607BF8E1" box="[486,533,1885,1911]" count="1" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
, 
<specimenCount id="14C056CCFFC4A649FDEAF8D5600BF8E1" box="[546,613,1885,1911]" count="2" pageId="37" pageNumber="38" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="14C056CCFFC4A649FDBEF8D56094F8E1" box="[630,762,1885,1911]" count="1" pageId="37" pageNumber="38" type="female">one female</specimenCount>
used for SEM); same collection data as for holotype; 
<specimenCode id="5260353EFFC4A649FF75F80863D1F80C" box="[189,447,1920,1946]" pageId="37" pageNumber="38">MECN–ARAC–38–T</specimenCode>
, in 
<collectionCode id="64D70580FFC4A649FE21F8096054F80D" box="[489,570,1921,1947]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="37" pageNumber="38" type="Museum">ZFMK</collectionCode>
Ar 24101
</materialsCitation>
.
</paragraph>
<paragraph id="02799D45FFC7A64AFF75FE806397FEB4" blockId="38.[189,505,264,290]" box="[189,505,264,290]" pageId="38" pageNumber="39">
<heading id="59312A29FFC7A64AFF75FE806397FEB4" bold="true" box="[189,505,264,290]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="30B24157FFC7A64AFF75FE806397FEB4" bold="true" box="[189,505,264,290]" pageId="38" pageNumber="39">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFC7A64AFF75FEBF6070FDB2" blockId="38.[189,1399,311,549]" pageId="38" pageNumber="39">
<materialsCitation id="B2AE9718FFC7A64AFF75FEBF6376FEE2" collectedFrom="in cave" collectingDate="2021-09-25" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" county="Cavernas del Anzu Forest Reserve" elevation="1160" latitude="-1.4067" location="Pastaza" longLatPrecision="7" longitude="-78.0449" municipality="Caverna" pageId="38" pageNumber="39" specimenCount="5" specimenCount-female="3" stateProvince="Pastaza">
<collectingCountry id="7AD1DDD5FFC7A64AFF75FEBF6323FEC7" box="[189,333,311,337]" name="Ecuador" pageId="38" pageNumber="39">ECUADOR</collectingCountry>
– 
<emphasis id="30B24157FFC7A64AFEA3FEBF63A3FEC7" bold="true" box="[363,461,311,337]" pageId="38" pageNumber="39">
<collectingRegion id="C00253A7FFC7A64AFEA3FEBF63A3FEC7" box="[363,461,311,337]" country="Ecuador" name="Pastaza" pageId="38" pageNumber="39">Pastaza</collectingRegion>
</emphasis>
• 
<specimenCount id="14C056CCFFC7A64AFE2FFEBF6045FEC4" box="[487,555,311,338]" count="3" pageId="38" pageNumber="39" type="female">3 ♀♀</specimenCount>
, 
<specimenCount id="14C056CCFFC7A64AFDF2FEBF60ECFEC7" box="[570,642,311,337]" pageId="38" pageNumber="39">2 juvs</specimenCount>
(in pure ethanol); same collection data as for holotype; 
<collectionCode id="64D70580FFC7A64AFAECFEB06718FEC4" box="[1316,1398,312,338]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="38" pageNumber="39" type="Museum">ZFMK</collectionCode>
Ecu174
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFC7A64AFEF0FED2610FFE2D" collectingDate="2021-09-26" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1140" latitude="-1.4054" location="Cueva Copa del Mundo" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" pageId="38" pageNumber="39" specimenCode="MECN-ARAC-39-T" specimenCount="5" specimenCount-female="1" specimenCount-male="2" stateProvince="Pastaza">
<specimenCount id="14C056CCFFC7A64AFEF0FED26313FEE3" box="[312,381,346,373]" count="2" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
(one palp used for SEM), 
<specimenCount id="14C056CCFFC7A64AFD07FED26090FEE3" box="[719,766,346,373]" count="1" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
, 
<specimenCount id="14C056CCFFC7A64AFCD9FED2610EFEE2" box="[785,864,346,372]" pageId="38" pageNumber="39">2 juvs</specimenCount>
; 
<collectingMunicipality id="E21D073FFFC7A64AFCA7FED26775FEE3" box="[879,1307,346,373]" pageId="38" pageNumber="39">Cavernas del Anzu Forest Reserve</collectingMunicipality>
, 
<location id="0719CB9EFFC7A64AFAE2FED263E3FE01" LSID="urn:lsid:plazi:treatment:8A6F2C53FFC4A645FD96FB3061A1FCD6:0719CB9EFFC7A64AFAE2FED263E3FE01" country="Ecuador" latitude="-1.4054" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" name="Cueva Copa del Mundo" pageId="38" pageNumber="39" stateProvince="Pastaza">Cueva Copa del Mundo</location>
; 
<geoCoordinate id="67F2FB82FFC7A64AFE53FEF5607BFE0E" box="[411,533,381,408]" degrees="1.4054" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-1.4054">1.4054° S</geoCoordinate>
, 
<geoCoordinate id="67F2FB82FFC7A64AFDECFEF660D6FE0E" box="[548,696,382,408]" degrees="78.0433" direction="west" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="-78.0433">78.0433° W</geoCoordinate>
; 
<elevation id="89EB7A76FFC7A64AFD0EFEF6610AFE01" box="[710,868,381,408]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="38" pageNumber="39" unit="m" value="1140.0">
<quantity id="C53E30A0FFC7A64AFD0EFEF6614FFE0E" box="[710,801,382,408]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="38" pageNumber="39" unit="m" value="1140.0">1140 m</quantity>
a.s.l.
</elevation>
; 
<date id="7678BB85FFC7A64AFCBCFEF56676FE0E" box="[884,1048,381,408]" pageId="38" pageNumber="39" value="2021-09-26">
<collectingDate id="663C426DFFC7A64AFCBCFEF56676FE0E" box="[884,1048,381,408]" pageId="38" pageNumber="39" value="2021-09-26">26 Sep. 2021</collectingDate>
</date>
; 
<collectorName id="AF33F893FFC7A64AFBEEFEF666DCFE01" box="[1062,1202,381,408]" pageId="38" pageNumber="39">B.A. Huber</collectorName>
and 
<collectorName id="AF33F893FFC7A64AFB38FEF66718FE0E" box="[1264,1398,382,408]" pageId="38" pageNumber="39">M. Herrera</collectorName>
leg.; in cave; 
<specimenCode id="5260353EFFC7A64AFEABFE296007FE2D" box="[355,617,417,443]" pageId="38" pageNumber="39">MECN–ARAC–39–T</specimenCode>
, in 
<collectionCode id="64D70580FFC7A64AFD5DFE296088FE2D" box="[661,742,417,443]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="38" pageNumber="39" type="Museum">ZFMK</collectionCode>
Ar 24102
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFC7A64AFCB5FE29611CFE48" collectingDate="2021-09-26" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1140" latitude="-1.4054" location="Cueva Copa del Mundo" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" pageId="38" pageNumber="39" specimenCode="MECN-ARAC-40-T" specimenCount="1" specimenCount-female="1" stateProvince="Pastaza">
<specimenCount id="14C056CCFFC7A64AFCB5FE2961C4FE2D" box="[893,938,417,443]" count="1" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
with egg-sac; same collection data as for preceding; 
<specimenCode id="5260353EFFC7A64AFEBAFE4C6019FE48" box="[370,631,452,478]" pageId="38" pageNumber="39">MECN–ARAC–40–T</specimenCode>
, in 
<collectionCode id="64D70580FFC7A64AFD6DFE4C6098FE48" box="[677,758,452,478]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="38" pageNumber="39" type="Museum">ZFMK</collectionCode>
Ar 24103
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFC7A64AFC47FE4C60E2FD97" collectingDate="2021-09-26" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. Herrera" country="Ecuador" elevation="1140" latitude="-1.4054" location="Cueva Copa del Mundo" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Pastaza">
<specimenCount id="14C056CCFFC7A64AFC47FE4C61BDFE48" box="[911,979,452,478]" count="1" pageId="38" pageNumber="39" type="generic">1 juv.</specimenCount>
(in pure ethanol); same collection data as for preceding; 
<collectionCode id="64D70580FFC7A64AFE1CFE606048FD94" box="[468,550,488,514]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="38" pageNumber="39" type="Museum">ZFMK</collectionCode>
Ecu183
</materialsCitation>
• 
<materialsCitation id="B2AE9718FFC7A64AFD65FE6F6074FDB2" collectingDate="2013-07-17" collectionCode="QCAZ" collectorName="M. Archambault" country="Ecuador" elevation="1140" latitude="-1.4054" location="Cueva Copa del Mundo" longLatPrecision="7" longitude="-78.0433" municipality="Cavernas del Anzu Forest Reserve" pageId="38" pageNumber="39" specimenCount="3" specimenCount-female="1" stateProvince="Pastaza">
<specimenCount id="14C056CCFFC7A64AFD65FE6F60B2FD94" box="[685,732,487,514]" count="1" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
, 
<specimenCount id="14C056CCFFC7A64AFD27FE6F6150FD97" box="[751,830,487,513]" pageId="38" pageNumber="39">2 juvs</specimenCount>
; same locality as for preceding; 
<date id="7678BB85FFC7A64AFB1DFE6F671FFD97" box="[1237,1393,487,513]" pageId="38" pageNumber="39" value="2013-07-17">
<collectingDate id="663C426DFFC7A64AFB1DFE6F671FFD97" box="[1237,1393,487,513]" pageId="38" pageNumber="39" value="2013-07-17">17 Jul. 2013</collectingDate>
</date>
; 
<collectorName id="AF33F893FFC7A64AFF75FD8363EFFDB2" box="[189,385,522,549]" pageId="38" pageNumber="39">M. Archambault</collectorName>
leg.; 
<collectionCode id="64D70580FFC7A64AFE0DFD826074FDB2" box="[453,538,522,548]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="38" pageNumber="39" type="Museum">QCAZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFC7A64AFF75FDC46195FD02" box="[189,1019,588,661]" pageId="38" pageNumber="39" type="etymology">
<paragraph id="02799D45FFC7A64AFF75FDC4632AFDF0" blockId="38.[189,324,588,614]" box="[189,324,588,614]" pageId="38" pageNumber="39">
<heading id="59312A29FFC7A64AFF75FDC4632AFDF0" bold="true" box="[189,324,588,614]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="30B24157FFC7A64AFF75FDC4632AFDF0" bold="true" box="[189,324,588,614]" pageId="38" pageNumber="39">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFC7A64AFF75FDF26195FD02" blockId="38.[189,1019,634,661]" box="[189,1019,634,661]" pageId="38" pageNumber="39">
The species name is derived from the 
<typeStatus id="DD7D23E7FFC7A64AFDB3FDF360C0FD03" box="[635,686,635,661]" pageId="38" pageNumber="39">type</typeStatus>
locality, noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFC7A645FF75FD33613BFEFF" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39" type="description">
<paragraph id="02799D45FFC7A64AFF75FD336320FD43" blockId="38.[189,334,699,725]" box="[189,334,699,725]" pageId="38" pageNumber="39">
<heading id="59312A29FFC7A64AFF75FD336320FD43" bold="true" box="[189,334,699,725]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="30B24157FFC7A64AFF75FD336320FD43" bold="true" box="[189,334,699,725]" pageId="38" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFC7A64AFF22FD6263DEFC92" blockId="38.[189,1399,746,914]" box="[234,432,746,772]" pageId="38" pageNumber="39">
<emphasis id="30B24157FFC7A64AFF22FD6263DEFC92" bold="true" box="[234,432,746,772]" pageId="38" pageNumber="39">
Male (
<typeStatus id="DD7D23E7FFC7A64AFEF1FD6263C6FC92" box="[313,424,746,772]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="02799D45FFC7A64AFF75FC8660E8FC07" blockId="38.[189,1399,746,914]" pageId="38" pageNumber="39">
<smallCapsWord id="049F0B99FFC7A64AFF75FC866302FCB0" baselines="801,801" box="[189,364,782,808]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="38" pageNumber="39">MEASUREMENTS</smallCapsWord>
. Total body length 4.2, carapace width 1.7. Distance PME–PME 200 µm; diameter PME 130 ×150 µm; distance PME–ALE 50 µm; AME absent (cf. female: 
<figureCitation id="9AFD81C0FFC7A64AFC32FCB86607FCDD" box="[1018,1129,816,843]" captionStart="Fig" captionStartId="51.[189,233,1773,1799]" captionTargetBox="[183,1404,265,1703]" captionTargetId="figure-86@51.[183,1404,265,1703]" captionTargetPageId="51" captionText="Fig. 41. Priscula spp., SEM photographs of female ocular areas and spinnerets. A–B. Ocular areas, frontal views; Priscula pastaza Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C–E. ALS, P. pastaza (C), P. esmeraldas Huber sp. nov. (D), and P. lumbaqui sp. nov. (E). F–H. Distinctive structures in front of spinnerets (in place of colulus); P. chapintza Huber sp. nov. (F), P. esmeraldas (G), and P. pastaza (H). Scale lines: 100 µm (A–B); 10 µm (C–G); 2 µm (H)." figureDoi="http://doi.org/10.5281/zenodo.10462956" httpUri="https://zenodo.org/record/10462956/files/figure.png" pageId="38" pageNumber="39">Fig. 41A</figureCitation>
). ALE and PLE larger than PME (diameter ALE 180 µm, PLE 160 µm). Leg 1: 41.4 (10.3+0.8+ 10.4 +17.5 +2.4), tibia 2: 7.7, tibia 3: 5.7, tibia 4: 7.3; tibia 1 L/d: 69.
</paragraph>
<paragraph id="02799D45FFC7A64AFF75FC3160CAFBAB" blockId="38.[189,1399,952,1085]" pageId="38" pageNumber="39">
<smallCapsWord id="049F0B99FFC7A64AFF75FC316367FC47" baselines="972,972" box="[189,265,953,979]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="38" pageNumber="39">COLOR</smallCapsWord>
(in ethanol). Carapace light-brown, with whitish marks beside ocular area, clypeus with large light brown band narrowing towards chelicerae; sternum light brown with thin darker brown margins; legs monochromous light brown, without darker rings; abdomen monochromous pale gray, only ventrally with darker ochre mark in gonopore area.
</paragraph>
<caption id="56B9CDCDFFC7A64AFF77F82366DAF87F" ID-DOI="http://doi.org/10.5281/zenodo.10462932" ID-Zenodo-Dep="10462932" httpUri="https://zenodo.org/record/10462932/files/figure.png" pageId="38" pageNumber="39" startId="38.[191,234,1963,1989]" targetBox="[183,1404,1160,1885]" targetPageId="38">
<paragraph id="02799D45FFC7A64AFF77F82366DAF87F" blockId="38.[191,1400,1963,2025]" pageId="38" pageNumber="39">
<emphasis id="30B24157FFC7A64AFF77F8236375F853" bold="true" box="[191,283,1963,1989]" pageId="38" pageNumber="39">Fig. 29.</emphasis>
<taxonomicName id="C5C6E6C6FFC7A64AFEE9F823605CF853" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[289,562,1963,1989]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pastaza" status="sp. nov.">
<emphasis id="30B24157FFC7A64AFEE9F823638BF853" box="[289,485,1963,1989]" italics="true" pageId="38" pageNumber="39">Priscula pastaza</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFC7A64AFDF1F82460F8F850" box="[569,662,1964,1990]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101; left male palp, prolateral, dorsal, and retrolateral views. Scale line: 0.5 mm.
</paragraph>
</caption>
<paragraph id="02799D45FFC6A64BFF75FE816398FE1A" blockId="39.[189,1399,265,396]" pageId="39" pageNumber="40">
<smallCapsWord id="049F0B99FFC6A64BFF75FE816293FEB7" baselines="284,284" box="[189,253,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="39" pageNumber="40">BODY</smallCapsWord>
. Habitus as in 
<figureCitation id="9AFD81C0FFC6A64BFE63FE816066FEB5" box="[427,520,265,291]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[183,1404,633,1854]" captionTargetId="figure-130@12.[183,1404,633,1854]" captionTargetPageId="12" captionText="Fig. 6. Priscula spp., binghamae and andinensis groups, live specimens. A–B. P. pastaza Huber sp. nov.; male and female with egg-sac from Cavernas del Anzu Forest Reserve. C–D. P. bonita Huber sp. nov.; male from La Bonita-Santa Barbara and female with egg-sac from Gruta de los Tayos. E–F. P. lumbaqui Huber sp. nov.; male and female from near Lumbaqui." figureDoi="http://doi.org/10.5281/zenodo.10462882" httpUri="https://zenodo.org/record/10462882/files/figure.png" pageId="39" pageNumber="40">Fig. 6A</figureCitation>
. Ocular area raised, without hump on posterior side, with comb of stronger hairs at median side of each ocular triad. Deep thoracic groove. Clypeus unmodified except sclerotized rim. Sternum wider than long (1.15/0.80), unmodified. Abdomen globular to slightly higher than long, dorso-posteriorly rounded.
</paragraph>
<caption id="56B9CDCDFFC6A64BFF75F8006728F87F" ID-DOI="http://doi.org/10.5281/zenodo.10462934" ID-Zenodo-Dep="10462934" httpUri="https://zenodo.org/record/10462934/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1928,1954]" targetBox="[199,1395,472,1826]" targetPageId="39">
<paragraph id="02799D45FFC6A64BFF75F8006728F87F" blockId="39.[189,1398,1928,2025]" pageId="39" pageNumber="40">
<emphasis id="30B24157FFC6A64BFF75F8006377F834" bold="true" box="[189,281,1928,1954]" pageId="39" pageNumber="40">Fig. 30.</emphasis>
<taxonomicName id="C5C6E6C6FFC6A64BFED7F800605EF834" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[287,560,1928,1954]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pastaza" status="sp. nov.">
<emphasis id="30B24157FFC6A64BFED7F800638DF834" box="[287,483,1928,1954]" italics="true" pageId="39" pageNumber="40">Priscula pastaza</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFC6A64BFDFFF80060FAF834" box="[567,660,1928,1954]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. 
<emphasis id="30B24157FFC6A64BFEF7F8236311F853" bold="true" box="[319,383,1963,1989]" pageId="39" pageNumber="40">A–C.</emphasis>
Left tarsus and procursus, prolateral, dorsal, and retrolateral views; asterisks: whitish protruding area. 
<emphasis id="30B24157FFC6A64BFE48F84663D4F87E" bold="true" box="[384,442,1998,2024]" pageId="39" pageNumber="40">D–F.</emphasis>
Left genital bulb, dorsal, retrolateral, and ventral views. Scale lines: 0.5 mm.
</paragraph>
</caption>
<paragraph id="02799D45FFC9A644FF75FE816020FED0" blockId="40.[189,1398,265,326]" pageId="40" pageNumber="41">
<smallCapsWord id="049F0B99FFC9A644FF75FE81632DFEB7" baselines="284,284" box="[189,323,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Chelicerae" pageId="40" pageNumber="41">CHELICERAE</smallCapsWord>
. As in 
<figureCitation id="9AFD81C0FFC9A644FE53FE816044FEB5" box="[411,554,265,291]" captionStart="Fig" captionStartId="40.[190,233,1928,1954]" captionTargetBox="[191,1393,1304,1888]" captionTargetId="figure-350@40.[186,1401,1296,1889]" captionTargetPageId="40" captionText="Fig. 31. Priscula pastaza Huber sp. nov.; male and female paratypes from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462936" httpUri="https://zenodo.org/record/10462936/files/figure.png" pageId="40" pageNumber="41">Fig. 31A–B</figureCitation>
, with short entapophyses, pair of short frontal apophyses close to fang joints; without stridulatory ridges.
</paragraph>
<paragraph id="02799D45FFC9A644FF75FEE660D2FD37" blockId="40.[189,1399,366,673]" pageId="40" pageNumber="41">
<smallCapsWord id="049F0B99FFC9A644FF75FEE66291FE10" baselines="385,385" box="[189,255,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Palps" pageId="40" pageNumber="41">PALPS</smallCapsWord>
. As in 
<figureCitation id="9AFD81C0FFC9A644FE90FEE66386FE1E" box="[344,488,366,392]" captionStart="Fig" captionStartId="38.[191,234,1963,1989]" captionTargetBox="[183,1404,1160,1885]" captionTargetId="figure-280@38.[183,1404,1160,1885]" captionTargetPageId="38" captionText="Fig. 29. Priscula pastaza Huber sp. nov.; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101; left male palp, prolateral, dorsal, and retrolateral views. Scale line: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462932" httpUri="https://zenodo.org/record/10462932/files/figure.png" pageId="40" pageNumber="41">Fig. 29A–C</figureCitation>
; coxa unmodified, trochanter slightly protruding ventrally, femur large, with unsclerotized retrolateral process proximally followed distally by sclerotized indentation, distal ventral rim not protruding; patella ventrally reduced to strongly sclerotized narrow rim; tibia small relative to femur; palpal tarsal organ exposed, weakly raised (
<figureCitation id="9AFD81C0FFC9A644FCF4FE5F61C4FE67" box="[828,938,471,497]" captionStart="Fig" captionStartId="53.[189,233,1771,1797]" captionTargetBox="[183,1404,265,1697]" captionTargetId="figure-98@53.[183,1404,265,1697]" captionTargetPageId="53" captionText="Fig. 43. Priscula spp., SEM photographs of female palps and of male and female tarsal organs. A–B. Tips of female palps in dorsal views; Priscula esmeraldas Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C. Male palpal tarsal organ, P. pastaza Huber sp. nov. D. Female palpal tarsal organ, P. lumbaqui. E–H. Female tarsal organs on leg 1 (E, P. lumbaqui), leg 2 (F, P. esmeraldas), leg 3 (G, P. pastaza), and leg 4 (H, P. pastaza). Scale lines: 10 µm (A, C–F); 20 µm (B); 2 µm (G–H)." figureDoi="http://doi.org/10.5281/zenodo.10462960" httpUri="https://zenodo.org/record/10462960/files/figure.png" pageId="40" pageNumber="41">Fig. 43C</figureCitation>
); procursus (
<figureCitation id="9AFD81C0FFC9A644FB8CFE5F66BCFE67" box="[1092,1234,471,497]" captionStart="Fig" captionStartId="39.[189,232,1928,1954]" captionTargetBox="[199,1395,472,1826]" captionTargetId="figure-98@39.[186,1401,472,1849]" captionTargetPageId="39" captionText="Fig. 30. Priscula pastaza Huber sp. nov.; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views; asterisks: whitish protruding area. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462934" httpUri="https://zenodo.org/record/10462934/files/figure.png" pageId="40" pageNumber="41">Fig. 30A–C</figureCitation>
) with distinct whitish protruding area dorsally, distinctive prolateral and dorsal membranous elements composed of hair-like processes (dorsal part shown in 
<figureCitation id="9AFD81C0FFC9A644FD56FD966147FDAE" box="[670,809,542,568]" captionStart="Fig" captionStartId="52.[189,232,1775,1801]" captionTargetBox="[181,1402,265,1699]" captionTargetId="figure-113@52.[181,1402,265,1699]" captionTargetPageId="52" captionText="Fig. 42. Priscula spp., SEM photographs of male pedipalps. A–C. Tip of left procursus in dorsal view (A) and hair-like processes on its prolateral (B) and retrolateral (C) sides; P. esmeraldas Huber sp. nov. D–E. Tip of left procursus in retrolateral view (D) and hair-like processes in more detail (E); P. pastaza Huber sp. nov. F–G. Left procursus and bulb of P. bonita Huber sp. nov., showing position of tarsal organ (arrow in F) and position of sperm duct opening (arrow in G). H. Sperm duct opening (arrow) on bulbal process in P. pastaza. Scale lines: 100 µm (A, F–G); 10 µm (B–C, E); 20 µm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.10462958" httpUri="https://zenodo.org/record/10462958/files/figure.png" pageId="40" pageNumber="41">Fig. 42D–E</figureCitation>
), distal ventral sclerite flat and weakly sclerotized; genital bulb (
<figureCitation id="9AFD81C0FFC9A644FE92FDC9638BFDCD" box="[346,485,577,603]" captionStart="Fig" captionStartId="39.[189,232,1928,1954]" captionTargetBox="[199,1395,472,1826]" captionTargetId="figure-98@39.[186,1401,472,1849]" captionTargetPageId="39" captionText="Fig. 30. Priscula pastaza Huber sp. nov.; male paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–C. Left tarsus and procursus, prolateral, dorsal, and retrolateral views; asterisks: whitish protruding area. D–F. Left genital bulb, dorsal, retrolateral, and ventral views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462934" httpUri="https://zenodo.org/record/10462934/files/figure.png" pageId="40" pageNumber="41">Fig. 30D–F</figureCitation>
) with small proximal sclerite connecting to tarsus, with large whitish area on retrolateral-ventral side, strong and slightly spiraling main bulbal process with subdistal sperm duct opening (arrow in 
<figureCitation id="9AFD81C0FFC9A644FE5EFD0F606FFD37" box="[406,513,647,673]" captionStart="Fig" captionStartId="52.[189,232,1775,1801]" captionTargetBox="[181,1402,265,1699]" captionTargetId="figure-113@52.[181,1402,265,1699]" captionTargetPageId="52" captionText="Fig. 42. Priscula spp., SEM photographs of male pedipalps. A–C. Tip of left procursus in dorsal view (A) and hair-like processes on its prolateral (B) and retrolateral (C) sides; P. esmeraldas Huber sp. nov. D–E. Tip of left procursus in retrolateral view (D) and hair-like processes in more detail (E); P. pastaza Huber sp. nov. F–G. Left procursus and bulb of P. bonita Huber sp. nov., showing position of tarsal organ (arrow in F) and position of sperm duct opening (arrow in G). H. Sperm duct opening (arrow) on bulbal process in P. pastaza. Scale lines: 100 µm (A, F–G); 10 µm (B–C, E); 20 µm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.10462958" httpUri="https://zenodo.org/record/10462958/files/figure.png" pageId="40" pageNumber="41">Fig. 42H</figureCitation>
) and obtuse tip.
</paragraph>
<paragraph id="02799D45FFC9A644FF75FD4166BAFCBF" blockId="40.[189,1399,713,810]" pageId="40" pageNumber="41">
<smallCapsWord id="049F0B99FFC9A644FF75FD416298FD77" baselines="732,732" box="[189,246,713,739]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="40" pageNumber="41">LEGS</smallCapsWord>
. Without spines; with curved hairs on tibiae and metatarsi (mainly legs 1 and 2); with few short vertical hairs; retrolateral trichobothrium of tibia 1 at 6%; prolateral trichobothrium present on all leg tibiae; tarsi without regular pseudosegmentation but rather with many indistinct platelets.
</paragraph>
<paragraph id="02799D45FFC9A644FF22FCD963D9FCFC" blockId="40.[189,1399,848,945]" box="[234,439,848,875]" pageId="40" pageNumber="41">
<emphasis id="30B24157FFC9A644FF22FCD963D9FCFC" bold="true" box="[234,439,848,875]" pageId="40" pageNumber="41">Male (variation)</emphasis>
</paragraph>
<paragraph id="02799D45FFC9A644FF75FCFC63C9FC27" blockId="40.[189,1399,848,945]" pageId="40" pageNumber="41">
Tibia 
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three other males: 9.3, 11.0, 11.3. AME pigment always present but lenses tiny (~20 µm diameter) or absent.
</paragraph>
<paragraph id="02799D45FFC9A644FF22FC50632BFC64" blockId="40.[189,1398,984,1222]" box="[234,325,984,1010]" pageId="40" pageNumber="41">
<emphasis id="30B24157FFC9A644FF22FC50632BFC64" bold="true" box="[234,325,984,1010]" pageId="40" pageNumber="41">Female</emphasis>
</paragraph>
<paragraph id="02799D45FFC9A645FF75FC74613BFEFF" blockId="40.[189,1398,984,1222]" lastBlockId="41.[189,1398,265,362]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
In general similar to male (
<figureCitation id="9AFD81C0FFC9A644FDEDFC7360E6FB83" box="[549,648,1019,1045]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[183,1404,633,1854]" captionTargetId="figure-130@12.[183,1404,633,1854]" captionTargetPageId="12" captionText="Fig. 6. Priscula spp., binghamae and andinensis groups, live specimens. A–B. P. pastaza Huber sp. nov.; male and female with egg-sac from Cavernas del Anzu Forest Reserve. C–D. P. bonita Huber sp. nov.; male from La Bonita-Santa Barbara and female with egg-sac from Gruta de los Tayos. E–F. P. lumbaqui Huber sp. nov.; male and female from near Lumbaqui." figureDoi="http://doi.org/10.5281/zenodo.10462882" httpUri="https://zenodo.org/record/10462882/files/figure.png" pageId="40" pageNumber="41">Fig. 6B</figureCitation>
) but clypeus rim not sclerotized and hairs on ocular area unmodified. AME variable as in males. Tibia 
<quantity id="C53E30A0FFC9A644FD3CFB97614AFBAF" box="[756,804,1055,1081]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="40" pageNumber="41" unit="in" value="1.0">1 in</quantity>
<specimenCount id="14C056CCFFC9A644FCE6FB9761A6FBAF" box="[814,968,1055,1081]" count="5" pageId="40" pageNumber="41" type="female">five females</specimenCount>
: 7.7–8.8 (mean 8.2). Tarsal organs exposed (
<figureCitation id="9AFD81C0FFC9A644FEFDFBCA63A1FBCA" box="[309,463,1090,1116]" captionStart="Fig" captionStartId="53.[189,233,1771,1797]" captionTargetBox="[183,1404,265,1697]" captionTargetId="figure-98@53.[183,1404,265,1697]" captionTargetPageId="53" captionText="Fig. 43. Priscula spp., SEM photographs of female palps and of male and female tarsal organs. A–B. Tips of female palps in dorsal views; Priscula esmeraldas Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C. Male palpal tarsal organ, P. pastaza Huber sp. nov. D. Female palpal tarsal organ, P. lumbaqui. E–H. Female tarsal organs on leg 1 (E, P. lumbaqui), leg 2 (F, P. esmeraldas), leg 3 (G, P. pastaza), and leg 4 (H, P. pastaza). Scale lines: 10 µm (A, C–F); 20 µm (B); 2 µm (G–H)." figureDoi="http://doi.org/10.5281/zenodo.10462960" httpUri="https://zenodo.org/record/10462960/files/figure.png" pageId="40" pageNumber="41">Fig. 43G–H</figureCitation>
); main tarsal claws with ~14–17 tines, tarsus 4 claws more evenly curved and with shorter tines than tarsi 1–3 claws (
<figureCitation id="9AFD81C0FFC9A644FD18FBED61EAFBE9" box="[720,900,1125,1151]" captionStart="Fig" captionStartId="54.[189,231,1774,1800]" captionTargetBox="[183,1404,265,1696]" captionTargetId="figure-96@54.[183,1404,265,1696]" captionTargetPageId="54" captionText="Fig. 44. Priscula spp., SEM photographs of female legs. A–B. Oval plates of unknown function, in Priscula lumbaqui Huber sp. nov. (A, metatarsus 4) and P. pastaza Huber sp. nov. (B, metatarsus 4). C–F. Tarsal claws of leg 1 (C, P. pastaza), leg 2 (D, P. pastaza), leg 3 (E, P. bonita Huber sp. nov.), and leg 4 (F, P. pastaza). G. Microstructure of claw tines (P. chapintza Huber sp. nov.). H. Basis of mechanoreceptive hair (P. chapintza; metatarsus 4), showing distal bulge (on right side). Scale lines: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10462964" httpUri="https://zenodo.org/record/10462964/files/figure.png" pageId="40" pageNumber="41">Fig. 44C–D, F</figureCitation>
). ALS with one strongly widened spigot, one pointed spigot, and one large and four small cylindrical spigots (
<figureCitation id="9AFD81C0FFC9A644FBDEFB0066E9FB34" box="[1046,1159,1160,1186]" captionStart="Fig" captionStartId="51.[189,233,1773,1799]" captionTargetBox="[183,1404,265,1703]" captionTargetId="figure-86@51.[183,1404,265,1703]" captionTargetPageId="51" captionText="Fig. 41. Priscula spp., SEM photographs of female ocular areas and spinnerets. A–B. Ocular areas, frontal views; Priscula pastaza Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C–E. ALS, P. pastaza (C), P. esmeraldas Huber sp. nov. (D), and P. lumbaqui sp. nov. (E). F–H. Distinctive structures in front of spinnerets (in place of colulus); P. chapintza Huber sp. nov. (F), P. esmeraldas (G), and P. pastaza (H). Scale lines: 100 µm (A–B); 10 µm (C–G); 2 µm (H)." figureDoi="http://doi.org/10.5281/zenodo.10462956" httpUri="https://zenodo.org/record/10462956/files/figure.png" pageId="40" pageNumber="41">Fig. 41C</figureCitation>
); with distinctively sculptured area medially in front of ALS (
<figureCitation id="9AFD81C0FFC9A644FD02FB236150FB50" box="[714,830,1195,1222]" captionStart="Fig" captionStartId="51.[189,233,1773,1799]" captionTargetBox="[183,1404,265,1703]" captionTargetId="figure-86@51.[183,1404,265,1703]" captionTargetPageId="51" captionText="Fig. 41. Priscula spp., SEM photographs of female ocular areas and spinnerets. A–B. Ocular areas, frontal views; Priscula pastaza Huber sp. nov. (A) and P. lumbaqui Huber sp. nov. (B). C–E. ALS, P. pastaza (C), P. esmeraldas Huber sp. nov. (D), and P. lumbaqui sp. nov. (E). F–H. Distinctive structures in front of spinnerets (in place of colulus); P. chapintza Huber sp. nov. (F), P. esmeraldas (G), and P. pastaza (H). Scale lines: 100 µm (A–B); 10 µm (C–G); 2 µm (H)." figureDoi="http://doi.org/10.5281/zenodo.10462956" httpUri="https://zenodo.org/record/10462956/files/figure.png" pageId="40" pageNumber="41">Fig. 41H</figureCitation>
). Epigynum (
<figureCitation id="9AFD81C0FFC9A644FC2EFB236614FB50" box="[998,1146,1195,1222]" captionStart="Fig" captionStartId="41.[189,232,1860,1886]" captionTargetBox="[183,1404,918,1784]" captionTargetId="figure-204@41.[183,1404,918,1784]" captionTargetPageId="41" captionText="Fig. 32. Priscula pastaza Huber sp. nov.; female paratype from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–B. Abdomen, ventral and lateral views. C–D. Cleared genitalia, ventral and dorsal views. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462938" httpUri="https://zenodo.org/record/10462938/files/figure.png" pageId="40" pageNumber="41">Fig. 32A–B</figureCitation>
) main anterior plate semicircular, slightly protruding, posteriorly with lighter semicircular area; posterior epigynal plate medially divided by whitish area. Internal genitalia (
<figureCitation id="9AFD81C0FFC8A645FC52FEA46667FED0" box="[922,1033,300,326]" captionStart="Fig" captionStartId="40.[190,233,1928,1954]" captionTargetBox="[191,1393,1304,1888]" captionTargetId="figure-350@40.[186,1401,1296,1889]" captionTargetPageId="40" captionText="Fig. 31. Priscula pastaza Huber sp. nov.; male and female paratypes from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. A–B. Male chelicerae, frontal and lateral views. C. Cleared female genitalia, dorsal view. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10462936" httpUri="https://zenodo.org/record/10462936/files/figure.png" pageId="41" pageNumber="42">Fig. 31C</figureCitation>
) with simple sclerotized arc, membranous ‘valve’, and pair or roundish pore plates.
</paragraph>
</subSubSection>
<caption id="56B9CDCDFFC9A644FF76F800602EF87F" ID-DOI="http://doi.org/10.5281/zenodo.10462936" ID-Zenodo-Dep="10462936" httpUri="https://zenodo.org/record/10462936/files/figure.png" pageId="40" pageNumber="41" startId="40.[190,233,1928,1954]" targetBox="[191,1393,1304,1888]" targetPageId="40">
<paragraph id="02799D45FFC9A644FF76F800602EF87F" blockId="40.[188,1398,1928,2025]" pageId="40" pageNumber="41">
<emphasis id="30B24157FFC9A644FF76F8006373F834" bold="true" box="[190,285,1928,1954]" pageId="40" pageNumber="41">Fig. 31.</emphasis>
<taxonomicName id="C5C6E6C6FFC9A644FEEEF800602FF834" authority="Huber" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[294,577,1928,1954]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pastaza" status="sp. nov.">
<emphasis id="30B24157FFC9A644FEEEF8006383F834" box="[294,493,1928,1954]" italics="true" pageId="40" pageNumber="41">Priscula pastaza</emphasis>
Huber
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFC9A644FD82F80060C6F834" box="[586,680,1928,1954]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
; male and female paratypes from Cavernas del Anzu Forest Reserve, ZFMK Ar 24101. 
<emphasis id="30B24157FFC9A644FE33F8236057F853" bold="true" box="[507,569,1963,1989]" pageId="40" pageNumber="41">A–B.</emphasis>
Male chelicerae, frontal and lateral views. 
<emphasis id="30B24157FFC9A644FBE6F8236625F853" bold="true" box="[1070,1099,1963,1989]" pageId="40" pageNumber="41">C.</emphasis>
Cleared female genitalia, dorsal view. Scale lines: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="4ADCCECEFFC8A645FF75FE1863F0FE6B" pageId="41" pageNumber="42" type="distribution">
<paragraph id="02799D45FFC8A645FF75FE186338FE3C" blockId="41.[189,342,400,426]" box="[189,342,400,426]" pageId="41" pageNumber="42">
<heading id="59312A29FFC8A645FF75FE186338FE3C" bold="true" box="[189,342,400,426]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="30B24157FFC8A645FF75FE186338FE3C" bold="true" box="[189,342,400,426]" pageId="41" pageNumber="42">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFC8A645FF75FE4863F0FE6B" blockId="41.[189,1398,447,509]" pageId="41" pageNumber="42">
Known from two neighboring caves in the Cavernas del Anzu Forest Reserve, 
<collectingRegion id="C00253A7FFC8A645FB57FE48671CFE4F" box="[1183,1394,447,474]" country="Ecuador" name="Pastaza" pageId="41" pageNumber="42">Pastaza Province</collectingRegion>
, 
<collectingCountry id="7AD1DDD5FFC8A645FF75FE6A634EFE6A" box="[189,288,482,508]" name="Ecuador" pageId="41" pageNumber="42">Ecuador</collectingCountry>
(
<figureCitation id="9AFD81C0FFC8A645FEF9FE6A63FEFE6B" box="[305,400,482,509]" captionStart="Fig" captionStartId="9.[189,232,1821,1847]" captionTargetBox="[183,1405,1129,1775]" captionTargetId="figure-307@9.[184,1405,1129,1774]" captionTargetPageId="9" captionText="Fig. 4. Known distribution of Priscula in Ecuador. A. Representatives of the gularis group. Circles: P. gularis (TL=type locality) and specimens treated herein as “P. gularis?” and “P. cf. gularis”. Squares: other Ecuadorian representatives of the gularis group. B. Representatives of the andinensis group (red) and binghamae group (blue) in Ecuador. The background colors indicate biogeographic regions of Ecuador (as far as relevant for Priscula, i.e., Andean regions from western foothill forests to eastern foothill forests), taken from Torres-Carvajal et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10462874" httpUri="https://zenodo.org/record/10462874/files/figure.png" pageId="41" pageNumber="42">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4ADCCECEFFC8A645FF75FDAC61A1FCD6" pageId="41" pageNumber="42" type="biology_ecology">
<paragraph id="02799D45FFC8A645FF75FDAC6310FDA8" blockId="41.[189,382,548,574]" box="[189,382,548,574]" pageId="41" pageNumber="42">
<heading id="59312A29FFC8A645FF75FDAC6310FDA8" bold="true" box="[189,382,548,574]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="30B24157FFC8A645FF75FDAC6310FDA8" bold="true" box="[189,382,548,574]" pageId="41" pageNumber="42">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="02799D45FFC8A645FF75FDDA61A1FCD6" blockId="41.[188,1400,594,832]" pageId="41" pageNumber="42">
All spiders were collected in the interior of two caves; adults were only found in the aphotic zones, but juveniles were also found in the twilight area (the first meters of the entrance areas of both caves were occupied by a different species that also occurs in the neighboring forest; see 
<taxonomicName id="C5C6E6C6FFC8A645FBA0FD1266BCFD25" authority="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera, 2023" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[1128,1234,665,691]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="bonita" status="sp. nov.">
<emphasis id="30B24157FFC8A645FBA0FD1266BCFD25" box="[1128,1234,665,691]" italics="true" pageId="41" pageNumber="42">P. bonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFC8A645FB16FD116752FD25" box="[1246,1340,665,691]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
). No specimens of 
<taxonomicName id="C5C6E6C6FFC8A645FE92FD3563BEFD40" authority="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera, 2023" authorityName="Huber &amp; Meng &amp; Dupérré &amp; Astrin &amp; Herrera" authorityYear="2023" box="[346,464,701,726]" class="Arachnida" family="Pholcidae" genus="Priscula" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="pastaza" status="sp. nov.">
<emphasis id="30B24157FFC8A645FE92FD3563BEFD40" box="[346,464,701,726]" italics="true" pageId="41" pageNumber="42">P. pastaza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B81FC2CFFC8A645FE1FFD35605EFD41" box="[471,560,701,727]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
were found in the well-preserved neighboring forest, suggesting that the species might be a troglobite, i.e. strictly bound to underground habitats. The spiders were hanging in very fine and barely visible webs, freely exposed among rocks or in wall niches. Two egg-sacs contained six and 
<specimenCount id="14C056CCFFC8A645FED0FCAE63F4FCD6" box="[280,410,806,832]" count="7" pageId="41" pageNumber="42" type="egg">seven eggs</specimenCount>
, respectively, with an egg diameter of 
<quantity id="C53E30A0FFC8A645FC97FCAD61A5FCD6" box="[863,971,805,832]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="41" pageNumber="42" unit="mm" value="0.95">0.95 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>