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<document ENCODING="UTF8" ID-DOI="http://dx.doi.org/10.3897/phytokeys.22.4041" ID-PMC="PMC3689140" ID-Pensoft-Pub="1314-2003-22-1" ID-Pensoft-UUID="FFC8FFF7FF8EFFFE9F6EFFEEFFFF3860" ID-PubMed="23794937" ID-Zenodo-Dep="576167" ModsDocID="1314-2003-22-1" checkinTime="1451251931319" checkinUser="pensoft" docAuthor="Knapp, Sandra" docDate="2013" docId="603BB09E469D5CB2BE8AA3045C457A3B" docLanguage="en" docName="PhytoKeys 22: 1-428" docOrigin="PhytoKeys 22" docPubDate="2013-05-10" docSource="http://dx.doi.org/10.3897/phytokeys.22.4041" docTitle="Solanum ruizii S. Knapp 1989" docType="treatment" docVersion="5" id="FFC8FFF7FF8EFFFE9F6EFFEEFFFF3860" lastPageNumber="214" masterDocId="FFC8FFF7FF8EFFFE9F6EFFEEFFFF3860" masterDocTitle="A revision of the Dulcamaroid Clade of Solanum L. (Solanaceae)" masterLastPageNumber="432" masterPageNumber="1" pageNumber="211" updateTime="1668140799926" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Dulcamaroid Clade of Solanum L. (Solanaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knapp, Sandra</mods:namePart>
<mods:affiliation>Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom</mods:affiliation>
<mods:nameIdentifier type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-05-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>22</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>432</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.22.4041</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.22.4041</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-22-1</mods:identifier>
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<treatment ID-GBIF-Taxon="182229100" LSID="urn:lsid:plazi:treatment:603BB09E469D5CB2BE8AA3045C457A3B" httpUri="http://treatment.plazi.org/id/603BB09E469D5CB2BE8AA3045C457A3B" lastPageId="213" lastPageNumber="214" pageId="210" pageNumber="211">
<subSubSection pageId="210" pageNumber="211" type="nomenclature">
<paragraph pageId="210" pageNumber="211">
32.
<taxonomicName LSID="603BB09E-469D-5CB2-BE8A-A3045C457A3B" authority="S. Knapp" authorityName="S. Knapp" authorityYear="1989" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum ruizii" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="ruizii">Solanum ruizii S. Knapp</taxonomicName>
,
<bibCitation journal="Bull. Brit. Mus. Nat. Hist. (Bot.)" page="91" score="37.32816182331285" volume="19" year="1989">Bull. Brit. Mus. Nat. Hist. (Bot.) 19: 91. 1989</bibCitation>
<figureCitation captionStart="Figure 80" captionStartId="F80" captionText="Figure 80. Solanum ruizii S. Knapp. (A-B drawn from Macbride 4351). Reproduced from Knapp (1989) with permission of the Natural History Museum Botany Library. Illustration by Margaret Tebbs." httpUri="https://binary.pensoft.net/fig/10337" pageId="210" pageNumber="211">Figure 80</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="210" pageNumber="211" type="reference_group">
<paragraph pageId="210" pageNumber="211">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="lanceolatum">Solanum lanceolatum</taxonomicName>
Ruiz &amp; Pav., Fl. Peruv. 2: 33, tab. 164a. 1799, non
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis italics="true" pageId="210" pageNumber="211">Solanum lanceolatum</emphasis>
</taxonomicName>
Cav. 1795. Type: Peru.
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
:
<normalizedToken originalValue="Muña">Muna</normalizedToken>
,
<emphasis italics="true" pageId="210" pageNumber="211">
H. Ruiz &amp; J.
<normalizedToken originalValue="Pavón">Pavon</normalizedToken>
</emphasis>
s.n. (lectotype, designated by Knapp, 2008c, pg. 320: MA [MA-747163]; isolectotypes: G [G00357895], MA [MA-747164, MA-747165]).
</paragraph>
<paragraph pageId="210" pageNumber="211">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="patulum">Solanum patulum</taxonomicName>
Pers., Syn. 1: 223. 1805, non
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="patulum">
<emphasis italics="true" pageId="210" pageNumber="211">Solanum patulum</emphasis>
</taxonomicName>
(L.) Roth, 1800. Type: Based on
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis italics="true" pageId="210" pageNumber="211">Solanum lanceolatum</emphasis>
</taxonomicName>
Ruiz &amp; Pav., non
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis italics="true" pageId="210" pageNumber="211">Solanum lanceolatum</emphasis>
</taxonomicName>
Cav., 1795.
</paragraph>
<paragraph pageId="210" pageNumber="211">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="patulum">Solanum patulum</taxonomicName>
Pers. var.
<emphasis italics="true" pageId="210" pageNumber="211">pilosistylum</emphasis>
Bitter, Bot. Jahrb. Syst. 54, Beibl. 119: 9. 1916. Type: Peru. Huancavelica: Tayacaja, cerros al lado derecho del
<normalizedToken originalValue="Río">Rio</normalizedToken>
Mantaro al sur de Surcubamba, 3800 m, 12 Mar 1913,
<emphasis italics="true" pageId="210" pageNumber="211">A. Weberbauer 6477</emphasis>
(holotype: B, destroyed; lectotype, designated by
<bibRefCitation author="Knapp, S" journalOrPublisher="Bulletin of the British Museum of Natural History (Botany)" pageId="320" pageNumber="321" pagination="63 - 112" refId="B68" refString="Knapp, S, 1989. A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae). Bulletin of the British Museum of Natural History (Botany) 19: 63 - 112" title="A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae)." volume="19" year="1989">Knapp 1989</bibRefCitation>
, pg. 91: MOL; isolectotypes: G [G00070195], MOL [2 sheets]).
</paragraph>
<paragraph pageId="210" pageNumber="211">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="patulum">Solanum patulum</taxonomicName>
Pers. forma
<emphasis italics="true" pageId="210" pageNumber="211">album</emphasis>
<bibCitation author="J. F. Macbr." journal="Publ. Field Columbian Mus., Bot. Ser." page="111" score="0.28006613869424335" volume="8" year="1930">
J.F.Macbr.,
<bibCitation journal="Publ. Field Columbian Mus., Bot. Ser." page="111" score="0.28006613869424335" volume="8" year="1930">Publ. Field Columbian Mus., Bot. Ser. 8: 111. 1930</bibCitation>
</bibCitation>
. Type: Peru.
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
: Tambo de Vaca, 3900 m, 10 Jun 1923,
<emphasis italics="true" pageId="210" pageNumber="211">J.F. Macbride 4441</emphasis>
(holotype: F [F-535527]; isotypes: G [G00070196], MA [MA-205962], US [US-1592826], W [W-1936-3811]).
</paragraph>
</subSubSection>
<subSubSection pageId="210" pageNumber="211" type="type">
<paragraph pageId="210" pageNumber="211">Type.</paragraph>
<paragraph pageId="210" pageNumber="211">
Based on
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis italics="true" pageId="210" pageNumber="211">Solanum lanceolatum</emphasis>
</taxonomicName>
Ruiz &amp; Pav., non
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis italics="true" pageId="210" pageNumber="211">Solanum lanceolatum</emphasis>
</taxonomicName>
Cav. 1795.
</paragraph>
</subSubSection>
<subSubSection lastPageId="211" lastPageNumber="212" pageId="210" pageNumber="211" type="description">
<paragraph pageId="210" pageNumber="211">Description.</paragraph>
<paragraph lastPageId="211" lastPageNumber="212" pageId="210" pageNumber="211">
Shrubs or small trees, 2-6 m tall. Stems densely pubescent with golden-yellow echinoid trichomes, these often elongate and tree-like; leaf scars somewhat raised; new growth densely pubescent with yellow echinoid and tree-like trichomes both above and below. Bark of older stems pale yellowish-white, glabrate. Sympodial units plurifoliate. Leaves narrowly elliptic, 7-13.5 cm long, 3-4 cm wide, the adaxial surfaces drying black with scattered golden echinoid and tree-like trichomes, these denser along the veins, the abaxial surfaces pubescent with the same echinoid trichomes, not drying as dark as the upper surfaces; primary veins 7-8 pairs, pubescent; base acute, not decurrent on to the petiole; margins entire; apex acuminate; petiole 1-1.5 cm long. Inflorescences terminal, becoming lateral with overtopping of the shoot, very large, 12-15 cm long, narrowly elliptic in outline, branching 8-10 times, with 10-25 flowers, each branch 0.5-3 cm long, densely pubescent with echinoid and tree-like trichomes; peduncle 1-2 cm long; pedicels 1.8-2.2 cm long, tapering from a basal diameter of 0.5-1 mm to an apical diameter of 1-1.5 mm, sparsely to densely pubescent with golden echinoid and tree-like trichomes, deflexed or horizontal at anthesis, articulated at the base and inserted in a sleeve ca. 0.5 mm long; pedicel scars closely spaced and clustered at the inflorescence branch tips. Buds ellipsoid, the corolla strongly exserted from the calyx tube. Flowers all perfect, 5-merous. Calyx tube 4-6 mm long, conical, the lobes 4-5 mm long, narrowly triangular, pubescent abaxially with the same trichomes as those of the pedicels, densely pubescent with golden dendritic trichomes adaxially. Corolla 4-4.8 cm in diameter, rotate-stellate, violet, very large and showy, lobed 1/2 of the way to the base, the lobes 23-25 mm long, 12-15 mm wide, planar or slightly cupulate at anthesis, densely pubescent abaxially with golden dendritic trichomes and with a few dendritic trichomes along the main veins adaxially. Filament tube minute or absent; free portion of the filaments ca. 1 mm long, glabrous; anthers ca. 5 mm long, 1-1.5 mm wide, loosely connivent, poricidal at the tips, the pores becoming slit-like with age. Ovary glabrous or with a few golden dendritic trichomes at the apex; style 1-1.2 cm long, densely pubescent at the base with golden dendritic trichomes; stigma bilobed, the surface minutely papillose. Fruit a globose berry ca. 1.1 cm in diameter (immature?), purplish-black with thin pericarp; fruiting pedicels 1.5-2 cm long, somewhat woody, erect or nodding,. Seeds ca. 30 per
<pageBreakToken pageId="211" pageNumber="212" start="start">berry</pageBreakToken>
, c. 3.5 mm long, 3 mm wide, reddish-brown, flattened lenticular, the surfaces minutely pitted. Chromosome number: not known.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10337" pageId="211" pageNumber="212" start="Figure 80" startId="F80">
<paragraph pageId="211" pageNumber="212">
<emphasis bold="true" pageId="211" pageNumber="212">Figure 80.</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="211" pageNumber="212" phylum="Tracheophyta" rank="species" species="ruizii">
<emphasis italics="true" pageId="211" pageNumber="212">Solanum ruizii</emphasis>
</taxonomicName>
S. Knapp. (
<emphasis bold="true" pageId="211" pageNumber="212">A-B</emphasis>
drawn from
<emphasis italics="true" pageId="211" pageNumber="212">Macbride 4351</emphasis>
). Reproduced from
<bibRefCitation author="Knapp, S" journalOrPublisher="Bulletin of the British Museum of Natural History (Botany)" pageId="320" pageNumber="321" pagination="63 - 112" refId="B68" refString="Knapp, S, 1989. A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae). Bulletin of the British Museum of Natural History (Botany) 19: 63 - 112" title="A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae)." volume="19" year="1989">Knapp (1989)</bibRefCitation>
with permission of the Natural History Museum Botany Library. Illustration by Margaret Tebbs.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="211" pageNumber="212" type="distribution">
<paragraph pageId="211" pageNumber="212">Distribution</paragraph>
<paragraph pageId="211" pageNumber="212">
(
<figureCitation captionStart="Figure 81" captionStartId="F81" captionText="Figure 81. Distribution of Solanum ruizii S. Knapp." httpUri="https://binary.pensoft.net/fig/10338" pageId="211" pageNumber="212">Figure 81</figureCitation>
). Endemic to central Peru from Depts.
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
to Huancavelica; from 3000-4000 m.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10338" pageId="211" pageNumber="212" start="Figure 81" startId="F81">
<paragraph pageId="211" pageNumber="212">
<emphasis bold="true" pageId="211" pageNumber="212">Figure 81.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="211" pageNumber="212" phylum="Tracheophyta" rank="species" species="ruizii">
<emphasis italics="true" pageId="211" pageNumber="212">Solanum ruizii</emphasis>
</taxonomicName>
S.Knapp.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="211" pageNumber="212" type="ecology">
<paragraph pageId="211" pageNumber="212">Ecology.</paragraph>
<paragraph pageId="211" pageNumber="212">In moist situations in cloud forests and forest margins; often growing in open grassy areas.</paragraph>
</subSubSection>
<subSubSection pageId="211" pageNumber="212" type="common names">
<paragraph pageId="211" pageNumber="212">Common names.</paragraph>
<paragraph pageId="211" pageNumber="212">
Peru. uruhuacta (
<emphasis italics="true" pageId="211" pageNumber="212">Macbride 4351</emphasis>
)
</paragraph>
<paragraph pageId="211" pageNumber="212">
Conservation status. Vulnerable (VU); EOO &lt;10,000 km2 (VU) and AOO &lt;4,500 km2 (NT). See
<bibRefCitation author="Moat, J" journalOrPublisher="Pesquisas, Botanica" pageId="322" pageNumber="323" refId="B116" refString="Moat, J, 2007. Conservation assessment tools extension for ArcView 3.x, version 1.2. GIS Unit, Royal Botanic Gardens, Kew. Available at http://www.rbgkew.org.uk/cats." title="Conservation assessment tools extension for ArcView 3. x, version 1.2. GIS Unit, Royal Botanic Gardens, Kew. Available at http: // www. rbgkew. org. uk / cats" year="2007">Moat (2007)</bibRefCitation>
for explanation of measurements.
</paragraph>
</subSubSection>
<subSubSection lastPageId="213" lastPageNumber="214" pageId="212" pageNumber="213" type="discussion">
<paragraph pageId="212" pageNumber="213">
<pageBreakToken pageId="212" pageNumber="213" start="start">Discussion</pageBreakToken>
.
</paragraph>
<paragraph lastPageId="213" lastPageNumber="214" pageId="212" pageNumber="213">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="212" pageNumber="213" phylum="Tracheophyta" rank="species" species="ruizii">
<emphasis italics="true" pageId="212" pageNumber="213">Solanum ruizii</emphasis>
</taxonomicName>
is certainly one of the most spectacular members of the
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="212" pageNumber="213" phylum="Tracheophyta" rank="species" species="nitidum">
<emphasis italics="true" pageId="212" pageNumber="213">Solanum nitidum</emphasis>
</taxonomicName>
species group with its large, deep purple flowers. All of the specimens of this species characterize it as being very showy. Label data on
<emphasis italics="true" pageId="212" pageNumber="213">Woytkowski 147</emphasis>
(F) state that
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="212" pageNumber="213" phylum="Tracheophyta" rank="species" species="ruizii">
<emphasis italics="true" pageId="212" pageNumber="213">Solanum ruizii</emphasis>
</taxonomicName>
is &quot;very common up to 3600 m.a.s level. Sometimes, growing in open places, attains considerable height and forms trees&quot;.
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="212" pageNumber="213" phylum="Tracheophyta" rank="species" species="ruizii">
<emphasis italics="true" pageId="212" pageNumber="213">Solanum ruizii</emphasis>
</taxonomicName>
has not been collected since the early part of the 20th century, but large-scale habitat destruction probably accounts for this. Collections of this species come from slightly drier areas than typical 'ceja de la
<normalizedToken originalValue="montaña">montana'</normalizedToken>
vegetation in which other members of the
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="212" pageNumber="213" phylum="Tracheophyta" rank="species" species="nitidum">
<emphasis italics="true" pageId="212" pageNumber="213">Solanum nitidum</emphasis>
</taxonomicName>
<pageBreakToken pageId="213" pageNumber="214" start="start">group</pageBreakToken>
(sensu
<bibRefCitation author="Knapp, S" journalOrPublisher="Bulletin of the British Museum of Natural History (Botany)" pageId="320" pageNumber="321" pagination="63 - 112" refId="B68" refString="Knapp, S, 1989. A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae). Bulletin of the British Museum of Natural History (Botany) 19: 63 - 112" title="A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae)." volume="19" year="1989">Knapp 1989</bibRefCitation>
) occur. These drier areas are characterized by extensive expanses of high elevation grassland or
<normalizedToken originalValue="pajonal">'pajonal'</normalizedToken>
.
</paragraph>
<paragraph pageId="213" pageNumber="214">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="213" pageNumber="214" phylum="Tracheophyta" rank="species" species="ruizii">
<emphasis italics="true" pageId="213" pageNumber="214">Solanum ruizii</emphasis>
</taxonomicName>
can be distinguished from its closest relative
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="213" pageNumber="214" phylum="Tracheophyta" rank="species" species="cutervanum">
<emphasis italics="true" pageId="213" pageNumber="214">Solanum cutervanum</emphasis>
</taxonomicName>
by flower size and by the number of seeds in the berries (ca. 30 versus only 7-8 in
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="213" pageNumber="214" phylum="Tracheophyta" rank="species" species="cutervanum">
<emphasis italics="true" pageId="213" pageNumber="214">Solanum cutervanum</emphasis>
</taxonomicName>
). Mature fruits of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="213" pageNumber="214" phylum="Tracheophyta" rank="species" species="ruizii">
<emphasis italics="true" pageId="213" pageNumber="214">Solanum ruizii</emphasis>
</taxonomicName>
are not known, but the label on
<emphasis italics="true" pageId="213" pageNumber="214">Pearce</emphasis>
s.n. from
<normalizedToken originalValue="Muña">Muna</normalizedToken>
states they are
<normalizedToken originalValue="“large”">&quot;large&quot;</normalizedToken>
, suggesting that the fruits I have observed are immature.
</paragraph>
<paragraph pageId="213" pageNumber="214">
Holotypes or lectotypes for Ruiz and
<normalizedToken originalValue="Pavón">Pavon</normalizedToken>
names in the Madrid herbarium (MA) not specifying a particular sheet are not sufficiently precise.
<bibRefCitation author="Knapp, S" journalOrPublisher="Bulletin of the British Museum of Natural History (Botany)" pageId="320" pageNumber="321" pagination="63 - 112" refId="B68" refString="Knapp, S, 1989. A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae). Bulletin of the British Museum of Natural History (Botany) 19: 63 - 112" title="A revision of the Solanum nitidum species group (section Holophylla pro parte: Solanaceae)." volume="19" year="1989">Knapp (1989)</bibRefCitation>
lectotypified
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Solanales" pageId="213" pageNumber="214" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis italics="true" pageId="213" pageNumber="214">Solanum lanceolatum</emphasis>
</taxonomicName>
citing only a sheet in MA; this was rectified in 2008 by citation of the particular sheet (
<bibRefCitation author="Knapp, S" journalOrPublisher="Anales del Jardin Botanico de Madrid" pageId="320" pageNumber="321" pagination="307 - 329" publicationUrl="10.3989/ajbm.2008.v65.i2.295" refId="B78" refString="Knapp, S, 2008c. Lectotypification of Ruiz and Pavon's names in Solanum (Solanaceae). Anales del Jardin Botanico de Madrid 65: 307 - 329, 10.3989/ajbm.2008.v65.i2.295" title="Lectotypification of Ruiz and Pavon's names in Solanum (Solanaceae)." url="10.3989/ajbm.2008.v65.i2.295" volume="65" year="2008 c">Knapp 2008c</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="213" pageNumber="214" type="specimens examined">
<paragraph pageId="213" pageNumber="214">Specimens examined.</paragraph>
<paragraph pageId="213" pageNumber="214">
<emphasis bold="true" pageId="213" pageNumber="214">Peru</emphasis>
.
<emphasis bold="true" pageId="213" pageNumber="214">Huancavelica</emphasis>
: Tayacaja, Huaribamba, 1 km before Huari, 3170 m, 28 Jul 1968,
<emphasis italics="true" pageId="213" pageNumber="214">Saunders 1173</emphasis>
(F); Tayacaja, Tayacaja, 3700 m, 10 Aug 1949,
<emphasis italics="true" pageId="213" pageNumber="214">
Velarde
<normalizedToken originalValue="Nuñez">Nunez</normalizedToken>
2008
</emphasis>
(US);
<emphasis bold="true" pageId="213" pageNumber="214">
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
</emphasis>
: Tambo de Vaca, 3963 m, 10 Jun 1923,
<emphasis italics="true" pageId="213" pageNumber="214">Macbride 4351</emphasis>
(G, US);
<normalizedToken originalValue="Muña">Muna</normalizedToken>
, May 1863,
<emphasis italics="true" pageId="213" pageNumber="214">Pearce 156</emphasis>
(BM);
<emphasis bold="true" pageId="213" pageNumber="214">
<normalizedToken originalValue="Junín">Junin</normalizedToken>
</emphasis>
: between Acopalca and Huari in steep canyon, NE of Huancayo, 4024 m, 19 Aug 1977,
<emphasis italics="true" pageId="213" pageNumber="214">Duncan et al. 2737</emphasis>
(NY).
</paragraph>
</subSubSection>
</treatment>
</document>