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<document id="61DD5B47AFFA0FA1894981B16393087D" ID-CLB-Dataset="22901" ID-DOI="10.11646/zootaxa.4763.3.4" ID-GBIF-Dataset="68dd0ab1-ecc3-4ac5-a00d-84d38ed0e5d4" ID-ISSN="1175-5334" ID-Zenodo-Dep="3759527" ID-ZooBank="urn:lsid:zoobank.org:pub:721665BA-2B50-407E-9561-FBAD1CA161F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587482723970" checkinUser="carolina" docAuthor="Daniel, Gimo M., Deschodt, Christian M., Davis, Adrian L. V. &amp; Sole, Catherine L." docDate="2020" docId="03EABB021577A14910D88E0CFDC642E3" docLanguage="en" docName="zootaxa.4763.3.4.pdf.imf" docOrigin="Zootaxa 4763 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stiptopodius savuti Daniel &amp; Deschodt &amp; Davis &amp; Sole 2020, new species" docType="treatment" docVersion="8" lastPageNumber="400" masterDocId="FFD3C37A1572A14F104F8F7DFFD84314" masterDocTitle="The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa" masterLastPageNumber="404" masterPageNumber="394" pageNumber="399" updateTime="1698834361793" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B5DAB922A99E6DD011F3963BBA5374E0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E4FA79D737E562699E33AA5FC416E6C6">
<mods:title id="365D3AAD498EF5E10122CAE9D8C8E444">The enigmatic dung beetle genus Stiptopodius Harold, 1871 (Coleoptera Scarabaeidae: Scarabaeinae) revisited: new species descriptions from southern Africa</mods:title>
</mods:titleInfo>
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<mods:role id="B52F6E662B826F2548A778846D08A9EB">
<mods:roleTerm id="1BE79C7FC5BF610C23D222E31C1109D6">Author</mods:roleTerm>
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<mods:namePart id="C9D48568B248506F36D05CA1A80200D7">Daniel, Gimo M.</mods:namePart>
<mods:nameIdentifier id="BFCF896156888B11ECEBFD551F635595" type="ORCID">0000-0003-3602-5034</mods:nameIdentifier>
<mods:affiliation id="495FB708EF2AF07B37C7376B1C8A7478">Department of Terrestria Invertebrates, The National Museum South Africal</mods:affiliation>
<mods:nameIdentifier id="77A752EC664934E4344CCD5E814D503E" type="URL">https://orcid.org/0000-0003-3602-5034</mods:nameIdentifier>
<mods:nameIdentifier id="FB2424E30477B048917D994A9E475981" type="email">gimo.daniel@nasmus.co.za</mods:nameIdentifier>
</mods:name>
<mods:name id="CBF5201F8F4F8A819E9B004FA09BD450" type="personal">
<mods:role id="291A9FBC8327DA60C7C4375F39FB6979">
<mods:roleTerm id="A1166C698AC9B637A185202F0198DD24">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="66F1464C4126576E34877BA5D80D3B74">Deschodt, Christian M.</mods:namePart>
<mods:affiliation id="6942594BCDCD9C50E929CE85F580ECCA">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="B504AB420BE03644190A4DEA981DBA6C" type="email">cdeschodt@zoology.up.ac.za</mods:nameIdentifier>
</mods:name>
<mods:name id="822E8B5C3E969B802804DDFA6F3E9DCA" type="personal">
<mods:role id="666C9F62F2396CED9EC1D1BE54AD1A38">
<mods:roleTerm id="F5A5CE260D8062D48DDD32A60110F356">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86C6FF038BDFB881559C1746C5F673E8">Davis, Adrian L. V.</mods:namePart>
<mods:affiliation id="69EFC53B4848ACA790525E84BF9C86FC">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="BE3DC6F897648C6B0AB71536502FD868" type="email">adrian.davis04@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AC63CBDB6BED5B3EE96A252D7201553A" type="personal">
<mods:role id="4EBBD668C7EC2338A8C38B55F292509C">
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</mods:role>
<mods:namePart id="56B12FF2B332256DE1FB9DAF02C56887">Sole, Catherine L.</mods:namePart>
<mods:affiliation id="66449FEF75C99D7CC20D695240530BF8">Department of Zoology &amp; Entomology, University of Pretoria</mods:affiliation>
<mods:nameIdentifier id="6D1732EB7EBBFBC496D634D5CF69B2A1" type="email">catherine.sole@up.ac.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B02ADA44FF5C6A96FFA8130C4C425EE7">text</mods:typeOfResource>
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<mods:title id="13D528307D5772CAB66D9D718FD8DA00">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EEED2FD2451E43382289DE00A8C36EA8">
<mods:date id="EF51C4C10298F3AA995ED059D302ACC6">2020</mods:date>
<mods:detail id="BE9289F904CB14D6A9501A6735A5B232" type="pubDate">
<mods:number id="DDB557904FCAF2DA938A2EC27C7489C0">2020-04-14</mods:number>
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<mods:number id="927F53F25B85CAFA39F2D68230DC2E57">4763</mods:number>
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<mods:classification id="794307611BC113E7704D1A8C5EB9BFD7">journal article</mods:classification>
<mods:identifier id="503F8DC8FE02F6739E3AC4CCA6F3A68A" type="CLB-Dataset">22901</mods:identifier>
<mods:identifier id="948179E967608241EA261AA7513030EB" type="DOI">10.11646/zootaxa.4763.3.4</mods:identifier>
<mods:identifier id="5AB472C509E8B87788EECEC9E4FEA870" type="GBIF-Dataset">68dd0ab1-ecc3-4ac5-a00d-84d38ed0e5d4</mods:identifier>
<mods:identifier id="CC8A18677226DAAAF7273A40C169E1A4" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03EABB021577A14910D88E0CFDC642E3" ID-DOI="http://doi.org/10.5281/zenodo.3805329" ID-GBIF-Taxon="163726935" ID-Zenodo-Dep="3805329" LSID="urn:lsid:plazi:treatment:03EABB021577A14910D88E0CFDC642E3" httpUri="http://treatment.plazi.org/id/03EABB021577A14910D88E0CFDC642E3" lastPageId="6" lastPageNumber="400" pageId="5" pageNumber="399">
<subSubSection id="C359599F1577A14A10D88E0CFCD6429F" box="[151,782,369,396]" pageId="5" pageNumber="399" type="nomenclature">
<paragraph id="8BFC0A141577A14A10D88E0CFCD6429F" blockId="5.[151,782,369,431]" box="[151,782,369,396]" pageId="5" pageNumber="399">
<heading id="D0B4BD781577A14A10D88E0CFCD6429F" bold="true" box="[151,782,369,396]" fontSize="10" level="2" pageId="5" pageNumber="399" reason="2">
<emphasis id="B937D6061577A14A10D88E0CFCD6429F" bold="true" box="[151,782,369,396]" pageId="5" pageNumber="399">
<taxonomicName id="4C4371971577A14A10D88E0CFDAA4298" ID-CoL="6ZW2K" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[151,626,369,396]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="savuti" status="sp. nov.">
<emphasis id="B937D6061577A14A10D88E0CFEA4429F" bold="true" box="[151,380,369,396]" italics="true" pageId="5" pageNumber="399">Stiptopodius savuti</emphasis>
Daniel &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A2046B7D1577A14A12318E0FFCD6429F" box="[638,782,369,396]" pageId="5" pageNumber="399" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14A10D88EE8FEFC42BB" box="[151,292,405,431]" pageId="5" pageNumber="399" type="description">
<paragraph id="8BFC0A141577A14A10D88EE8FEFC42BB" blockId="5.[151,782,369,431]" box="[151,292,405,431]" pageId="5" pageNumber="399">
(
<figureCitation id="137816911577A14A10D08EE8FED942BB" box="[159,257,405,431]" captionStart="FIGURES 1" captionStartId="3.[151,264,1877,1902]" captionTargetBox="[163,1426,206,1839]" captionTargetId="figure@3.[151,1436,198,1849]" captionTargetPageId="3" captionText="FIGURES 1A–D. Habitus of Stiptopodius Harold, 1871 species, holotypes. A, Stiptopodius peringueyi Daniel &amp; Deschodt, new species; B, Stiptopodius savuti Daniel &amp; Deschodt, new species; C, Stiptopodius muellerae Daniel &amp; Deschodt, new species; D, Stiptopodius singularis (Péringuey, 1901). Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3759529" httpUri="https://zenodo.org/record/3759529/files/figure.png" pageId="5" pageNumber="399">Figs. 1B</figureCitation>
,
<figureCitation id="137816911577A14A11428EE8FEC442BB" box="[269,284,405,431]" captionStart="FIGURE 4" captionStartId="8.[151,250,1093,1118]" captionTargetBox="[151,1412,151,1074]" captionTargetId="figure@8.[151,1412,151,1074]" captionTargetPageId="8" captionText="FIGURE 4. Map showing the type localities of Stiptopodius muellerae Daniel &amp; Deschodt, new species (.), Stiptopodius savuti Daniel &amp; Deschodt, new species (A) and Stiptopodius peringueyi Daniel &amp; Deschodt, new species (■)." figureDoi="http://doi.org/10.5281/zenodo.3759535" httpUri="https://zenodo.org/record/3759535/files/figure.png" pageId="5" pageNumber="399">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14A10D88EA3FD4F42EC" box="[151,663,477,504]" pageId="5" pageNumber="399" type="distribution">
<paragraph id="8BFC0A141577A14A10D88EA3FD4F42EC" blockId="5.[151,1437,477,2016]" box="[151,663,477,504]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10D88EA3FEE842EC" bold="true" box="[151,304,477,504]" pageId="5" pageNumber="399">
<typeStatus id="54F8B4B61577A14A10D88EA3FF0A42EC" box="[151,210,478,504]" pageId="5" pageNumber="399">Type</typeStatus>
locality
</emphasis>
: Savuti, North West,
<collectingCountry id="F3544A841577A14A126E8EA3FD4B42EC" box="[545,659,478,504]" name="Botswana" pageId="5" pageNumber="399">Botswana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14A10888D7FFDCE4177" pageId="5" pageNumber="399" type="materials_examined">
<paragraph id="8BFC0A141577A14A10888D7FFDCE4177" blockId="5.[151,1437,477,2016]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888D7FFEAD4108" bold="true" box="[199,373,513,540]" pageId="5" pageNumber="399">Type material.</emphasis>
<materialsCitation id="3B2B00491577A14A11348D7CFDCE4177" ID-GBIF-Occurrence="2603207388" collectingDate="2005-12-15" collectionCode="TMSA" country="Botswana" latitude="-18.53092" location="Site" longLatPrecision="1" longitude="24.08378" municipality="North West" pageId="5" pageNumber="399" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54F8B4B61577A14A11348D7CFE3E410F" box="[379,486,513,539]" pageId="5" pageNumber="399" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D45C19D1577A14A11BE8D7CFD9A410F" box="[497,578,513,539]" pageId="5" pageNumber="399" type="female">female</specimenCount>
: Savuti [
<collectingCountry id="F3544A841577A14A12EF8D7FFCCC4108" box="[672,788,514,540]" name="Botswana" pageId="5" pageNumber="399">Botswana</collectingCountry>
,
<collectingMunicipality id="6B98906E1577A14A13508D7CFC7C4108" box="[799,932,513,540]" pageId="5" pageNumber="399">North West</collectingMunicipality>
],
<location id="8E9C5CCF1577A14A13FA8D7CFC39410F" LSID="urn:lsid:plazi:treatment:03EABB021577A14910D88E0CFDC642E3:8E9C5CCF1577A14A13FA8D7CFC39410F" box="[949,993,513,539]" country="Botswana" latitude="-18.53092" longLatPrecision="1" longitude="24.08378" municipality="North West" name="Site" pageId="5" pageNumber="399">Site</location>
4 |
<geoCoordinate id="EE776CD31577A14A14468D7CFB544108" box="[1033,1164,513,540]" degrees="18.53092" direction="south" orientation="latitude" pageId="5" pageNumber="399" precision="1" value="-18.53092">S18.53092</geoCoordinate>
[°];
<geoCoordinate id="EE776CD31577A14A14E38D7CFAF74108" box="[1196,1327,513,540]" degrees="24.08378" direction="east" orientation="longitude" pageId="5" pageNumber="399" precision="1" value="24.08378">E24.08378</geoCoordinate>
[°] |
<date id="FFFD2CD41577A14A151A8D7CFF174154" pageId="5" pageNumber="399" value="2005-12-15">
<collectingDate id="EFB9D53C1577A14A151A8D7CFF174154" pageId="5" pageNumber="399" value="2005-12-15">15.xii. 2005</collectingDate>
</date>
|
<location id="8E9C5CCF1577A14A10A98D58FE14412B" LSID="urn:lsid:plazi:treatment:03EABB021577A14910D88E0CFDC642E3:8E9C5CCF1577A14A10A98D58FE14412B" box="[230,460,549,575]" country="Botswana" latitude="-18.53092" longLatPrecision="1" longitude="24.08378" municipality="North West" name="Deschodt &amp; Tshikae" pageId="5" pageNumber="399">Deschodt &amp; Tshikae</location>
|
<location id="8E9C5CCF1577A14A11AD8D58FDE0412B" LSID="urn:lsid:plazi:treatment:03EABB021577A14910D88E0CFDC642E3:8E9C5CCF1577A14A11AD8D58FDE0412B" box="[482,568,549,575]" country="Botswana" latitude="-18.53092" longLatPrecision="1" longitude="24.08378" municipality="North West" name="Carrion" pageId="5" pageNumber="399">Carrion</location>
pitfall trap ||
<emphasis id="B937D6061577A14A129C8D58FCA34154" bold="true" box="[723,891,549,576]" pageId="5" pageNumber="399">HOLOTYPE:</emphasis>
<emphasis id="B937D6061577A14A13CC8D58FBF84154" box="[899,1056,549,576]" italics="true" pageId="5" pageNumber="399">
<taxonomicName id="4C4371971577A14A13CC8D58FBD7412B" authorityName="Harold" authorityYear="1871" box="[899,1039,549,575]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="genus">
<location id="8E9C5CCF1577A14A13CC8D58FBD7412B" LSID="urn:lsid:plazi:treatment:03EABB021577A14910D88E0CFDC642E3:8E9C5CCF1577A14A13CC8D58FBD7412B" box="[899,1039,549,575]" country="Botswana" latitude="-18.53092" longLatPrecision="1" longitude="24.08378" municipality="North West" name="Stiptopodius" pageId="5" pageNumber="399">Stiptopodius</location>
</taxonomicName>
|
</emphasis>
<taxonomicName id="4C4371971577A14A14678D5BFA96412B" authority="Daniel &amp; Deschodt" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[1064,1358,549,575]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="savuti">
<emphasis id="B937D6061577A14A14678D5BFBB3412B" box="[1064,1131,550,575]" italics="true" pageId="5" pageNumber="399">savuti</emphasis>
<location id="8E9C5CCF1577A14A143C8D58FA96412B" LSID="urn:lsid:plazi:treatment:03EABB021577A14910D88E0CFDC642E3:8E9C5CCF1577A14A143C8D58FA96412B" box="[1139,1358,549,575]" country="Botswana" latitude="-18.53092" longLatPrecision="1" longitude="24.08378" municipality="North West" name="Daniel &amp; Deschodt" pageId="5" pageNumber="399">Daniel &amp; Deschodt</location>
</taxonomicName>
| 2020 [printed on the red card] (
<collectionCode id="ED5292D11577A14A11F98D34FDD24177" box="[438,522,585,611]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="399">TMSA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14A10888D10FE504563" pageId="5" pageNumber="399" type="description">
<paragraph id="8BFC0A141577A14A10888D10FC45419C" blockId="5.[151,1437,477,2016]" box="[199,925,621,648]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888D10FE81419C" bold="true" box="[199,345,621,648]" pageId="5" pageNumber="399">Description.</emphasis>
<typeStatus id="54F8B4B61577A14A112F8D10FE134193" box="[352,459,621,647]" pageId="5" pageNumber="399" type="holotype">Holotype</typeStatus>
, female. Length:
<quantity id="4CBBA7F11577A14A12C18D10FD3E419C" box="[654,742,621,648]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="399" unit="mm" value="4.3">4.3 mm</quantity>
, width:
<quantity id="4CBBA7F11577A14A130F8D10FC41419C" box="[832,921,621,648]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="399" unit="mm" value="1.9">1.9 mm</quantity>
.
</paragraph>
<paragraph id="8BFC0A141577A14A10888DECFAB841BF" blockId="5.[151,1437,477,2016]" box="[199,1376,657,684]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888DECFEF841B8" bold="true" box="[199,288,657,684]" pageId="5" pageNumber="399">Colour.</emphasis>
Dorsally black; lateral edge of clypeus, legs, and last abdominal ventrite brown; setae yellowish.
</paragraph>
<paragraph id="8BFC0A141577A14A10888DCBFE514090" blockId="5.[151,1437,477,2016]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888DCBFED641C4" bold="true" box="[199,270,694,720]" pageId="5" pageNumber="399">Head.</emphasis>
Incision between medial clypeal teeth deeply concave; latero-clypeal edges not sinuate; genae with external protrusion sharply triangular; surface of the head smooth and punctate; ocellate punctures sparse on the middle of vertex; some large punctures with outer edges moderately raised on clypeus, otherwise, finely punctate; clypeo-frontal carina not visible; clypeal-genal suture present; vertex with carina occupying half the space between eyes, carina interrupted in the middle clearly forming two small cephalic tubercles. Antennae with eight antennomeres, labium setose.
</paragraph>
<paragraph id="8BFC0A141577A14A10888CF3FBC840E4" blockId="5.[151,1437,477,2016]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888CF3FE9F40BC" bold="true" box="[199,327,910,936]" pageId="5" pageNumber="399">Pronotum.</emphasis>
Convex, wider than long; base of pronotum marginate; anterior margin wider than posterior edge; in lateral view, lateral edge strongly sinuate before postero-lateral angles, central disc glabrous; punctation ocellate with sparse, short setae antero-laterally that emanate from centres of punctures.
</paragraph>
<paragraph id="8BFC0A141577A14A10888C87FE8F4748" blockId="5.[151,1437,477,2016]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888C87FEC24700" bold="true" box="[199,282,1018,1044]" pageId="5" pageNumber="399">Elytra.</emphasis>
Laterally straight, somewhat convex distally and concave proximally; sparsely and finely punctate in external-most four striae, the rest apunctate; interstriae with sparse, short setae visible basally, only interstriae V–VIII punctate.
</paragraph>
<paragraph id="8BFC0A141577A14A10888B18FAE54794" blockId="5.[151,1437,477,2016]" box="[199,1341,1125,1152]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888B18FE9A4794" bold="true" box="[199,322,1125,1152]" pageId="5" pageNumber="399">Pygidium.</emphasis>
Convex with dense ocellate punctation bearing setae emanating from centres of punctures.
</paragraph>
<paragraph id="8BFC0A141577A14A10888BF4FC1D4604" blockId="5.[151,1437,477,2016]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888BF4FE1C47B0" bold="true" box="[199,452,1161,1188]" pageId="5" pageNumber="399">Abdominal ventrites.</emphasis>
Prosternum with anterior angles slanted forward and inwards, surface somewhat excavated. Mesoventrite dull, setose and punctate. Metaventrite shiny with long setae anteriorly, glabrous mid-posteriorly. Abdominal ventrites glabrous medially, shiny laterally, setigerous with ocellate punctures, setae arranged in lateral rows; last ventrites wider medially than others (female character).
</paragraph>
<paragraph id="8BFC0A141577A14A10888A67FC914547" blockId="5.[151,1437,477,2016]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888A67FEDE4620" bold="true" box="[199,262,1306,1332]" pageId="5" pageNumber="399">Legs.</emphasis>
External and internal margins of profemora punctate, with well-developed setae, central area apunctate and glabrous; external and internal edges strongly and slightly marginate, respectively. Meso- and metafemora with external and internal edges sparsely punctate, all punctures bearing setae (shorter than those in profemora), surface between external and internal edges glabrous, strongly marginate internally. Protibiae crenulate with four teeth on external margin, a line of setae around teeth in dorsal view; in ventral view, protibiae with two carinae, internally punctate and pilose. Protibial spur long and inward curving. Meso- and metatibiae triangular and strongly widened apically. Protarsi broken off in both legs. Meso- and metatarsomeres I–IV triangular, widened apically, wider than long, strongly concave medially, external and internal edges with distal lobe; meso- and metatarsomere V rectangular and longer than others, with pair of very tiny apical claws.
</paragraph>
<paragraph id="8BFC0A141577A14A10888923FE504563" blockId="5.[151,1437,477,2016]" box="[199,392,1629,1656]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888923FED3456C" bold="true" box="[199,267,1630,1656]" pageId="5" pageNumber="399">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14A108889FCFBA04588" box="[199,1144,1665,1692]" pageId="5" pageNumber="399" type="distribution">
<paragraph id="8BFC0A141577A14A108889FCFBA04588" blockId="5.[151,1437,477,2016]" box="[199,1144,1665,1692]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A108889FCFEB94588" bold="true" box="[199,353,1665,1692]" pageId="5" pageNumber="399">Distribution.</emphasis>
This species is only known from the
<typeStatus id="54F8B4B61577A14A134B89FFFCEC4588" box="[772,820,1666,1692]" pageId="5" pageNumber="399">type</typeStatus>
locality Savuti in
<collectingCountry id="F3544A841577A14A144E89FFFBAB4588" box="[1025,1139,1666,1692]" name="Botswana" pageId="5" pageNumber="399">Botswana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14A108889DBFB1245AB" box="[199,1226,1701,1728]" pageId="5" pageNumber="399" type="etymology">
<paragraph id="8BFC0A141577A14A108889DBFB1245AB" blockId="5.[151,1437,477,2016]" box="[199,1226,1701,1728]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A108889DBFE9645D4" bold="true" box="[199,334,1702,1728]" pageId="5" pageNumber="399">Etymology.</emphasis>
The specific name, a noun in apposition, refers to the name of the
<typeStatus id="54F8B4B61577A14A147689DBFBB145D4" box="[1081,1129,1702,1728]" pageId="5" pageNumber="399">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14A108889B4FC464467" pageId="5" pageNumber="399" type="diagnosis">
<paragraph id="8BFC0A141577A14A108889B4FC464467" blockId="5.[151,1437,477,2016]" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A108889B4FE9A45F0" bold="true" box="[199,322,1737,1764]" pageId="5" pageNumber="399">Diagnosis.</emphasis>
<taxonomicName id="4C4371971577A14A110489B4FDFC45F7" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[331,548,1737,1763]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="savuti" status="sp. nov.">
<emphasis id="B937D6061577A14A110489B4FDFC45F7" box="[331,548,1737,1763]" italics="true" pageId="5" pageNumber="399">Stiptopodius savuti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D1577A14A126189B7FD6F45F7" box="[558,695,1737,1764]" pageId="5" pageNumber="399" rank="species">new species</taxonomicNameLabel>
is morphologically similar to
<taxonomicName id="4C4371971577A14A145789B7FB6B45F7" authorityName="Branco" authorityYear="2009" box="[1048,1203,1737,1764]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B937D6061577A14A145789B7FB6B45F7" box="[1048,1203,1737,1764]" italics="true" pageId="5" pageNumber="399">S. granulosus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C4371971577A14A157589B7FA4445F7" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[1338,1436,1738,1764]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="savuti" status="sp. nov.">
<emphasis id="B937D6061577A14A157589B7FA4445F7" box="[1338,1436,1738,1764]" italics="true" pageId="5" pageNumber="399">S. savuti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D1577A14A10D88993FEC64413" box="[151,286,1773,1800]" pageId="5" pageNumber="399" rank="species">new species</taxonomicNameLabel>
can be distinguished by having the dorsal cephalic and pronotal surface smooth while in
<taxonomicName id="4C4371971577A14A154B8993FA444413" authorityName="Branco" authorityYear="2009" box="[1284,1436,1773,1800]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B937D6061577A14A154B8993FA444413" box="[1284,1436,1773,1800]" italics="true" pageId="5" pageNumber="399">S. granulosus</emphasis>
</taxonomicName>
they are densely rugose. The new species bears a carina which is interrupted medially and clearly forms two small cephalic tubercles, unlike
<taxonomicName id="4C4371971577A14A11F0884BFD80445B" authorityName="Branco" authorityYear="2009" box="[447,600,1845,1872]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B937D6061577A14A11F0884BFD80445B" box="[447,600,1845,1872]" italics="true" pageId="5" pageNumber="399">S. granulosus</emphasis>
</taxonomicName>
, whose cephalic carina does not form tubercles. Furthermore, the head of the new species is glabrous, whereas in
<taxonomicName id="4C4371971577A14A121A8827FD354467" authorityName="Branco" authorityYear="2009" box="[597,749,1881,1908]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B937D6061577A14A121A8827FD354467" box="[597,749,1881,1908]" italics="true" pageId="5" pageNumber="399">S. granulosus</emphasis>
</taxonomicName>
it is setigerous.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1577A14910888803FAC14257" lastPageId="6" lastPageNumber="400" pageId="5" pageNumber="399" type="discussion">
<paragraph id="8BFC0A141577A14910888803FAC14257" blockId="5.[151,1437,477,2016]" lastBlockId="6.[151,1437,153,503]" lastPageId="6" lastPageNumber="400" pageId="5" pageNumber="399">
<emphasis id="B937D6061577A14A10888803FEEC448C" bold="true" box="[199,308,1918,1944]" pageId="5" pageNumber="399">Remarks</emphasis>
: The Afrotropical
<taxonomicName id="4C4371971577A14A124C8800FD014483" authority="Daniel &amp; Deschodt &amp; Davis &amp; Sole, 2020" authorityName="Daniel &amp; Deschodt &amp; Davis &amp; Sole" authorityYear="2020" box="[515,729,1917,1943]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="savuti" status="sp. nov.">
<emphasis id="B937D6061577A14A124C8800FD014483" box="[515,729,1917,1943]" italics="true" pageId="5" pageNumber="399">Stiptopodius savuti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2046B7D1577A14A12908803FCBD4483" box="[735,869,1917,1944]" pageId="5" pageNumber="399" rank="species">new species</taxonomicNameLabel>
seems to be similar to the brown Somalian species of
<taxonomicName id="4C4371971577A14A10FA88DCFDBD44AF" authority="Muller, 1942" authorityName="Muller" authorityYear="1942" box="[181,613,1953,1979]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="glabricollis">
<emphasis id="B937D6061577A14A10FA88DCFE1144AF" box="[181,457,1953,1979]" italics="true" pageId="5" pageNumber="399">Stiptopodius glabricollis</emphasis>
Müller, 1942
</taxonomicName>
. Our knowledge on Afrotropical dung beetle biogeographical distribution patterns suggests that it is very unlikely that the inland species from
<collectingCountry id="F3544A841577A14A13FF88BBFBC644F4" box="[944,1054,1990,2016]" name="Botswana" pageId="5" pageNumber="399">Botswana</collectingCountry>
could be a sub-population of the coastal eastern horn of Africa species,
<taxonomicName id="4C4371971574A149120A8FE4FD3943A7" authorityName="Muller" authorityYear="1942" box="[581,737,153,179]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="400" phylum="Arthropoda" rank="species" species="glabricollis">
<emphasis id="B937D6061574A149120A8FE4FD3943A7" box="[581,737,153,179]" italics="true" pageId="6" pageNumber="400">S. glabricollis</emphasis>
</taxonomicName>
(see also a similar case in
<bibRefCitation id="EFD277E51574A14914448FE4FAE643A7" author="Genier, F. &amp; Josso, J. F." box="[1035,1342,153,179]" firstAuthor="Genier" pageId="6" pageNumber="400" pagination="333 - 342" refId="ref7273" refString="Genier, F. &amp; Josso, J. F. (2016) Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae). Zootaxa, 4072 (3), 333 - 342. https: // doi. org / 10.11646 / zootaxa. 4072.3.3" type="journal article" year="2016">Génier &amp; Josso 2017: 338).</bibRefCitation>
As such a wide and disjunct distribution is very rare among Afrotropical Scarabaeinae, we therefore consider the specimen from Savuti (
<collectingCountry id="F3544A841574A14911658F9CFE4443EF" box="[298,412,225,251]" name="Botswana" pageId="6" pageNumber="400">Botswana</collectingCountry>
) as a new species.
<taxonomicName id="4C4371971574A14912208F9CFCA543EF" authorityName="Branco" authorityYear="2009" box="[623,893,225,251]" class="Insecta" family="Scarabaeidae" genus="Stiptopodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="400" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B937D6061574A14912208F9CFCA543EF" box="[623,893,225,251]" italics="true" pageId="6" pageNumber="400">Stiptopodius granulosus</emphasis>
</taxonomicName>
(from
<collectingCountry id="F3544A841574A14913848F9CFBFA43E8" box="[971,1058,225,252]" name="Zambia" pageId="6" pageNumber="400">Zambia</collectingCountry>
) is morphologically similar to the new species. Since both species occur in neighbour countries, there is a huge potential of misidentification between them, that’s why the diagnosis of the new species is compared to the closest species in southern Africa.
</paragraph>
</subSubSection>
<subSubSection id="C359599F1574A14910888E31FDC642E3" pageId="6" pageNumber="400" type="conservation">
<paragraph id="8BFC0A141574A14910888E31FDC642E3" blockId="6.[151,1437,153,503]" pageId="6" pageNumber="400">
<emphasis id="B937D6061574A14910888E31FD644273" bold="true" box="[199,700,332,359]" pageId="6" pageNumber="400">Conservation status and potential threats.</emphasis>
This species was recorded from the well-protected Savuti area of Chobe National Park in northern
<collectingCountry id="F3544A841574A14912478E0CFDA2429F" box="[520,634,369,395]" name="Botswana" pageId="6" pageNumber="400">Botswana</collectingCountry>
. The vegetation consists of somewhat open to dense stands of
<taxonomicName id="4C4371971574A14915618E0CFE2242BB" authority="(Schinz) Schrire. The" authorityName="Schrire. The" baseAuthorityName="Schinz" class="Magnoliopsida" family="Fabaceae" genus="Philenoptera" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="400" phylum="Tracheophyta" rank="species" species="nelsii">
<emphasis id="B937D6061574A14915618E0CFEDF42BB" italics="true" pageId="6" pageNumber="400">Philenoptera nelsii</emphasis>
(Schinz) Schrire. The
</taxonomicName>
trees are about one to two meters tall with about 30% canopy cover (CMD, personal observation). Although it is protected at Savuti, the extent of occurrence of this species is not known and it should thus be assessed as Data Deficient.
</paragraph>
</subSubSection>
</treatment>
</document>
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